EVOLUTION (Two Lectures) I. INTRODUCTION A. Observations 1. All life forms share certain common characteristics (unity) 2. There are many forms of life on earth today (diversity) B. Explanation: Principle of Evolution 1. Unity: common descent 2. Diversity: adapt to environment (Natural Selection) 3. Define: Hypothesis Theory Principle: Scientific Method observations hypothesis scientific (controlled) observations/experiments interpretation of new data conclusion reject/support hypothesis Many scientific observations/experiments over long time period = THEORY Theories that are generally accepted as valid by an overwhelming # of scientists = PRINCIPLE / LAW II. III. CHARLES DARWIN A. 1831 - MHS Beagle naturalist (see p 241 map) B. 1858 - Darwin & Wallace both proposed natural selection. Data to back theory: Origin of Species C. Other hypotheses for agents of evolutionary change rejected by physical evidence: Ex: Lamarck: Inheritance of acquired characteristics 1. If you don't use it, you loose it 2. No scientific evidence to support Ex: muscles atrophy in individual; offspring have potential muscle development POPULATION GENETICS (MICROEVOLUTION) Changes in a population's allele frequencies over time. A. Definitions: 1. Population = all members of single species occupying a particular area at the same time. 2. Species = interbreed & produce viable offspring 3. Allele = version of a gene. It shows one particular manifestation of a trait. 4. Gene = section of DNA with enough code to make one protein. Each protein contributes to a trait that is inherited. 5. Gene pool = all the various alleles of all the genes in all the members of a population 1 6. 7. B. Gene flow = movement of alleles between populations Mutation = Change in alleles a. Translocation b. Deletion c. Inversion d. Duplication Population variations members of population vary in function, physical appearance & behavior C. Reproductive potential vs reality 1. Mathematical calc shows an animal population unchecked would increase faster than food supply (Thomas Malthus, 1798) 2. Available resources not sufficient to support all members of population 3. Only certain members of population survive & reproduce D. Survival of the Fittest (Differential reproduction) 1. Fitness = ability of individual to survive & reproduce in its local environment 2. Ex: Convergent evolution a. Dissimilar species living in similar environments have similar physical appearances b. Ex.: i. Sharks, seals, whales, penguins & sea turtles have flippers ii. Bats, birds & butterflies have wings E. Natural selection 1. Stabilization selection - intermediate phenotype favored 2. Disruptive selection – both extreme phenotypes favored 3. Directional selection - one extreme phenotype favored F. Agents of evolutionary change 1. Mutation = Raw material for evolutionary change 2. Natural Selection 3. Adaptive radiation = rapid development from 1 sp. many Occurs with lack of competition a) Galapagos: tortoises; finches b) Australia: marsupials 4. Interruption of gene flow (Gene flow keeps gene pools similar & large Prevents speciation) a. Reproductive isolation = increased chances of speciation 1) Allopatric speciation Ex: geographic isolation 2 4. 5. 2) Sympatric speciation a) Rare in animals b) Polyploidy in some plants Nonrandom mating = mating preference for particular phenotype a. Decreases # of heterozygotes b. Increases # of homozygotes Genetic drift = changes in allelic frequencies due to chance Most noticeable in small populations a. Founder Effect When few individuals found new colony, bring only fraction of original gene pool, which alleles are carried determined by chance Ex: 1) Smith Is.: blonde hair & blue eyes 2) Amish of Lancaster, PA i. 1 in 14 carries recessive allele that causes unusual form of dwarfism (affects lower arms/legs only) and polydactylism (extra fingers) ii. 1 in 1000 carries allele in pop. at large b. Bottleneck effect = natural or manmade disaster drives population to near extinction 1) Few individuals surviving = small gene pool 2) Surviving alleles selected by chance, not better adapted (endangered species) G. VI. Population NOT evolving Frequency of alleles remains constant Hardy - Weinberg rule SUMMARY Principle of Evolution - explanation of unity & diversity of life MACROEVOLUTION Reading fossils Comparative embryology and early development Comparative anatomy MICROEVOLUTION Comparative biochemistry Proteins: yeasts & humans have 90% of the same proteins DNA fingerprinting 3 1. 99% shared with chimpanzees (Jane Goodall) 2. 18 of 23 human chromosomes nearly identical to chimpanzees & gorillas Other 5 differ at inverted & translocated regions VII. HOMEWORK DUE AT START OF NEXT LECTURE Required: After reading these chapters, collect one living organism (See outline of homework for details) (Optional Extra Credit): INDEPENDENT STUDY FOR EXTRA CREDIT: KINGDOM PROTISTA (See outline of homework for details) INDEPENDENT STUDY FOR EXTRA CREDIT: THE FUNGI KINGDOM (See outline of homework for details) 4