37.1.6
B
S
T
Elymus kronokensis (Kom.) Tzvelev in Grubov (1968), Pl. As. Centr. 4: 216.
Agropyron kronokense Kom. (1915), Feddes Repert. 13: 87.
Roegneria kronokensis (Kom.) Tzvelev in Tolm. (1964), Fl. Arct. URSS 2: 246.
Russian Far East: Kamtchatka, Kronotskii pass, dry tundra in the alpine zone,
10.08.1909, leg. V. Komarov 3180 (LE), holotype?
Comments:
(1)
Tzvelev in Fl. Arct. URSS (1964, 1995) writes that virtually only one character
separates the three species Roegneria borealis, R. kronokensis and R. scandicus: leaf
surface emergences. The characters given by Melderis in Tutin et al. (1980) for
separation of the subspecies borealis, scandicus and subalpinus have proved
inapplicable for separation of entities in N Europe. As the three subspecies now
proposed in Tzvelev's draft ('subalpinus' is by me included in 'scandicus') also are
reported as mainly sympatric, their value as separate taxa should be very critically
evaluated. We don't even recognize varities on such meagre criteria in other families.
(Elven)
FIRST WARNING! Elymus kronokensis will be merged with E. alaskanus, under the
former name, if very good arguments for the current solution are not proposed.
SECOND WARNING! Subspecies will be merged if not much better justified.
37.1.6.1 Elymus kronokensis (Kom.) Tzvelev subsp. kronokensis
S
2n=
28 (4x).
2nD
?
G
RFE ALA
Comments:
(1)
What is the source for the report of this 'subspecies' from Alaska? (Elven)
37.1.6.2 Elymus kronokensis (Kom.) Tzvelev subsp. borealis (Turcz.) Tzvelev (1973), Nov.
Sist. Vyssh. Rast. 10: 24.
B
Triticum boreale Turcz. (1856), Fl. Baic.-Dahur. 2,1: 345.
S
Agropyron boreale (Turcz.) Drobov (1916), Tr. Bot. Muz. Imp. Akad. Nauk 16: 84;
Roegneria borealis (Turcz.) Nevski in Kom. (1934), Fl. SSSR 2: 624 [or 264?];
Elymus alaskanus (Scribn. & Merr.) Á. Löve subsp. borealis (Turcz.) Á. Löve & D.
Löve (1976), Bot. Not. 128: 502.
T
Siberia: Yakutsk ASSR, Ad viam Ochotensem ad fl. Aldan, 1835, leg. Kuznetzov
(LE) holotype?
2n=
28 (4x).
2nD
?
G
RUS SIB RFE ALA CAN
Comments:
(1)
The chromosome counts listed by Löve & Löve (1975) under E. trachycaulus subsp.
borealis probably all stem from E. kronokensis in Tzvelev's concept but from several
subspecies. (Elven)
37.1.6.3 Elymus kronokensis (Kom.) Tzvelev subsp. scandicus (Nevski) Tzvelev (2000),
Nov. Sist. Vyssh. Rast. 32: 182.
B
Roegneria scandica Nevski in Kom. (1934), Fl. URSS 2: 624.
S
E. alaskanus (Scribn. & Merr.) Á. Löve subsp. scandicus (Nevski) Melderis (1978),
Bot. J. Linn. Soc. 76: 375; E. kronokensis (Kom.) Tzvelev subsp. subalpinus (Neum.)
Tzvelev (1973), Nov. Sist. Vyssh. Rast. 10: 24; E. alaskanus (Scribn. & Merr.) Á.
Löve subsp. subalpinus (Neum.) Á. Löve & D. Löve (1976), Bot. Not. 128: 502.
T
Lapponia ponojensis, Orlov, ad rupes litorales, 31.07.1889, leg. Kihlman (LE) ***
2n=
28 (4x).
2nD
Knaben & Engelskjøn (1971 S & N Nor).
G
RUS SIB RFE ALA
Comments:
(1)
The 'subalpinus' synonyms should probably be included as they are names accepted
by Löve & Löve (1975), Tzvelev (1976: 120) and Lid & Lid (1994). Löve & Löve
list two Russian counts, both as arctic, under 'subalpinus'. (Elven)
37.1.7 Elymus sajanensis (Nevski) Tzvelev (1972), Nov. Sist. Vyssh. Rast. 9: 61.
B
Roegneria sajanensis Nevski in Kom. (1934), Fl. SSSR 2: 624.
S
T
Siberia: E Sayan, Tuniskii region, Garganskii pass, 31.07.1902, leg. Komorov (LE),
holotype?
Comments:
(1)
See the discussion by Tzvelev in Fl. Arct. URSSS (1964, 1995) concerning the (very
small) difference between subsp. hyperarcticus and subsp. villosus; one character is
reported. As both entities now are recognized in his draft from both Asia and
America, his last argument for keeping them apart, as geographical races, is
weakened. Which means that 'villosus' should be considered synonymous with
'hyperarcticus'.
Subsp. caeruleus also needs good morphological and/or genetical
justification before acceptance.
A third question is whether there is good correspondence between the S
Siberian plants ('sajanensis' s. str.) and the major arctic ones ('hyperarcticus').
Another solution could then be E. sajanensis with one (or more?) southern subspecies
and one major northern one, for which the name subsp. hyperarcticus probably would
have priority. If they are considered as species, Roegneria villosa (1954) has priority
before Roegneria hyperarctica (1964). It is also a question whether the 'hyperarctica'
entity could not be merged into a larger E. alaskanus. (Elven)
WARNING! Subspecies might be omitted if not (much) better justified.
37.1.7.1 Elymus sajanensis (Nevski) Tzvelev subsp. hyperarcticus (Polunin) Tzvelev
(1973), Nov. Sist. Vyssh. Rast. 10: 24.
B
Agropyron violaceum (Hornem.) Lange var. hyperarcticum Polunin (1940), Bull.
Natl. Mus. Can. 92, Biol. Ser., 24: 95.
S
Roegneria hyperarctica (Polunin) Tzvelev in Tolm. (1964), Fl. Arct. URSS 2: 244;
Agropyron boreale (Turcz.) Drobov subsp. hyperarcticum (Polunin) Melderis in
Hultén (1968), Ark. Bot., ser. 2, 7, 1: 19; Elymus hyperarcticus (Polunin) Tzvelev
(1972), Nov. Sist. Vyssh. Rast. 9: 61; Elymus alaskanus (Scribn. & Merr.) Á. Löve
subsp. hyperarcticus (Polunin) Á. Löve & D. Löve (1976), Bot. Not. 128: 502.
T
Canada: Baffin Isl., Arctic Bay, 09.09.1936, leg. N. Polunin 2531 (BM) holotype.
2n=
28 (4x).
2nD
Böcher & Larsen (1950 Grl); Sokolovskaya & Probatova in Tzvelev (1978).
G
SIB RFE ALA CAN GRL
Comments:
37.1.7.2 Elymus sajanensis (Nevski) Tzvelev subsp. villosus (V. Vassil.) Tzvelev (1973),
Nov. Sist. Vyssh. Rast. 10: 24.
B
Roegneria villosa V. Vassil. (1954), Bot. Mater. Gerb. Bot. Inst. AN SSSR 16: 57.
S
Elymus alaskanus (Scribn. & Merr.) Á. Löve subsp. villosus (V. Vassil.) Á. Löve &
D. Löve (1976), Bot. Not. 128: 502.
T
Russian Far East: Chukotsk region, Chegituni valley, sand-pebble deposits,
08.08.1938, leg. Trushkovskii (LE), holotype?
2n=
28 (4x).
2nD
Zhukova (1965b); Zhukova & Petrovsky (1976).
G
SIB RFE ALA
Comments:
37.1.7.3 Elymus sajanensis (Nevski) Tzvelev subsp. caeruleus (Jurtz.) Tzvelev (2000), Nov.
Sist. Vyssh. Rast. 32: 182.
B
Roegneria villosa V. Vassil. subsp. caerulea Jurtz. (1989), Bot. Zhurn. 74: 113.
S
2n=
2nD
G
RFE
Comments:
The Elymus macrourus aggregate (E. jacutensis, E. macrourus, ?E. sericeus)
Comments:
(1)
The aggregate might naturally disappear if E. jacutensis is merged with E.
macrourus, a solution indicated below, and if Agropyron sericeum only is accepted in
synonymy. (Elven)
(2)
Agropyron sericeum Hitchc. (1915), Amer. J. Bot. 1915, 2: 309, should be included
somewhere - either as an accepted taxon or in synonymy - as it is accepted as species
by Porsild & Cody (1980). Hultén (1968) only mentions the name as a synonym of
Agropyron macrourum, but Björn Salomon (pers. comm.) doubts whether the
American plants belong within E. macrourus s. str. (Elven)
37.1.8 Elymus macrourus (Turcz.) Tzvelev (1970), Sched. Herb. Fl. URSS 18: 30.
B
Triticum macrourum Turcz. in Steud. (1855), Syn. Pl. Glum. 1: 343.
S
Agropyron macrourum (Turcz.) Drobov (1916), Trav. Mus. Bot. Acad. Petersb. 16:
86; Roegneria macroura (Turcz.) Nevski in Kom. (1934), Fl. SSSR 2: 627.
T
Siberia: In sabulosis ad fl. Angaram superiorem, 1834, leg. Kuznetzov (LE) ***.
Comments:
(1)
According to Tzvelev in Fl Arct. URSS (1964, 1995), Roegneria macroura, R.
nepliana (at that time only known from a lost type specimen), R. turuchanensis, and
R. jacutensis are separated only by single characters. They are also virtually
sympatric, the more narrowly distributed entities all occur within the wider range of
E. macrourus s. str. and subsp. turuchanensis is found 'on both sides of' subsp.
neplinus. In other families, such character variation would only justify varieties, at
best. The lack of intermediacy in such characters is more often a result of the
characters being under a simple genetical control (e.g. awned and awnless, and
smooth, scabrous and hairy) and not a sign of high age and taxonomic significance.
We should consider reduction of all the subspecies, and E. jacutensis, to synonymy.
(Elven)
WARNING! Subspecies will disappear if not more well-founded than currently.
37.1.8.1 Elymus macrourus (Turcz.) Tzvelev subsp. macrourus
S
2n=
28 (4x).
2nD
Zhukova & Petrovsky (1975).
G
RUS SIB RFE ALA CAN
Comments:
37.1.8.2 Elymus macrourus (Turcz.) Tzvelev subsp. neplianus (V. Vassil.) Tzvelev (1973),
Nov. Sist. Vyssh. Rast. 10: 25.
B
Roegneria nepliana V. Vassil. (1954), Bot. Mater. Gerb. Bot. Inst. AN SSSR 16: 56.
T
Russian Far East: Northern slope of the Anyuiskii range, valley of Umkaveem River
upstream, 04.09.1950, leg. Nepli (LE), holotype.
S
2n=
28 (4x).
2nD
Zhukova (1967a); Zhukova & Petrovsky (1976 Chukotka).
G
SIB RFE
Comments:
37.1.8.3 Elymus macrourus (Turcz.) Tzvelev subsp. turuchanensis (Reverd.) Tzvelev
(1971), Nov. Sist. Vyssh. Rast. 8: 63.
B
Agropyron turuchanense Reverd. (1932), Sist. Zam. Mater. Gerb. Tomsk Univ. 4: 2.
S
Roegneria turuchanensis (Reverd.) Nevski in Kom. (1934), Fl. SSSR 2: 626.
T
Siberia: lower reaches of Enisei, Lapkaikha valley, meadows along bank, 30.07.1914,
leg. Reverdatto (TK), lectotype selected by ***.
2n=
28 (4x).
2nD
Sokolovskaya (1970 Komi area).
G
RUS SIB ALA
Comments:
37.1.9
B
S
T
Elymus jacutensis (Drobov) Tzvelev (1972), Nov. Sist. Vyssh. Rast. 9: 61.
Agropyron jacutense Drobov (1916), Tr. Bot. Muz. Peterb. Akad. Nauk 16: 94.
Roegneria jacutensis (Drobov) Nevski in Kom. (1934), Fl. SSSR 2: 625.
Siberia: Yakutia, Vilyuskii area, Bilyuchai River (right tributary of Vilyui), 40 versts
from the mouth, pebbled bank, 15.08.1914, leg. Drobov 625 (LE), lectotype selected
by ***.
2n=
28 (4x).
2nD
Zhukova (1967a, 1969).
G
SIB RFE
Comments:
(1)
The comment made by Tzvelev in Fl. Arct. URSS (1964, 1995, p. 294 in English
translation) throws serious doubt on this as a taxon separate from E. macrourus.
Similar awn variation, even with signs of sterility in the few intermediates known, is
documented in N European E. caninus without discernible genetical difference and
without high rank being given. There might be a very simple genetic background.
Perhaps a variety, at best? (Elven)
37.1.10 Elymus fibrosus (Schrenk) Tzvelev (1970), Sched. Herb. Fl. URSS 18: 29.
B
Triticum fibrosum Schrenk (1845), Bull. Cl. Phys.-Math. Acad. Imp. Sci. SaintPétersbourg 3: 209.
S
Roegneria fibrosa (Schrenk) Nevski in Kom. (1934), Fl. SSSR 2: 625.
T
Russia?: Songaria, in montibus Karkaraly, 1843, leg. Schrenk 1541 (LE), holotype.
Comments:
37.1.10.1 Elymus fibrosus (Schrenk) Tzvelev subsp. fibrosus
S
2n=
28 (4x).
2nD
Melderis (1955); Löve & Löve (1961); Sokolovskaya (1970).
G
RUS
Comments:
37.1.10.2 Elymus fibrosus (Schrenk) Tzvelev subsp. subfibrosus (Tzvelev) Tzvelev (1973),
Nov. Sist. Vyssh. Rast. 10: 25.
B
Roegneria subfibrosa Tzvelev in Tolm. (1964), Fl. Arct. URSS 2: 238.
S
T
Siberia: obl. Jakutsk, Balaganach, in arenosis ad fluminem, 15.07.1898, leg. Nilsson
(LE) holotype.
2n=
28 (4x).
2nD
Sokolovskaya & Probatova in Tzvelev (1976).
G
SIB RFE
Comments:
37.1.11 Elymus caninus (L.) L. (1755), Fl. Suec., ed. 2, 39.
B
Triticum caninum L. (1753), Sp. Pl. 86.
S
Agropyron caninum (L.) Beauv. (1812), Ess. Agrostogr. 101, 146; Roegneria canina
(L.) Nevski in Kom. (1934), Fl. SSSR 2: 617.
2n=
28 (4x).
2nD
Löve & Löve (1975) list numerous counts, one Icelandic.
G
ICE NOR RUS
Comments:
(1)
Similar awn-less plants are found here as given rank by Tzvelev in, e.g., E.
kronokensis and E. macrourus - jacutensis. They have sometimes been named (e.g.
Roegneria behmii) but have been found not to differ significantly genetically.
However, they may have their own distribution and ecology and even be partly sterile
in crosses with normal awned plants. Varietal rank might be appropriate. (Elven)
37.1.12 Elymus confusus (Roshev.) Tzvelev in Grubov (1968), Pl. As. Centr. 4: 221.
B
Agropyron confusum Roshev. (1924), Bot. Mater. Glavn. Bot. Sada RSFSR 5: 150.
S
Roegneria confusa (Roshev.) Nevski in Kom. (1934), Fl. SSSR 2: 605.
T
Siberia: Ircutia, 1828, leg. Turczaninow (LE), lectotype selected by ***.
2n=
28 (4x).
2nD
Zhukova (1967a); Sokolovskaya & Probatova in Tzvelev (1976).
G
SIB RFE
Comments:
37.1.13 Elymus asiaticus Á. Löve (1984), Feddes Repert. 95: 465.
S
2n=
28 (4x).
2nD
?
G
SIB
Comments: