Marjorie Angelica Balaguer Baccalaureate Biology HL 11-1/2 Option D: Evolution Phylogeny and Systematics I-II (4.22 – 4.30 2010) Phylogeny and Systematics I Phylogeny – the study of evolutionary relatedness among various groups of organisms.1 (Common ancestry or convergent, DNA & genetic similarities) Systematics – the study of the diversification of life on the planet Earth, both past and present, and the relationships among living things through time.2 + Outline the value of classifying organisms Classifying organisms allows us to; - Identify unknown organisms - Determine evolutionary relationships - Make predictions about members of a group, based on data from another + Define clade and cladistics Clade – (monophyletic group) A common ancestor and all its descendants Cladistics – A system of natural classification in which organisms are grouped based on characteristics that they share amongst themselves and a recent common ancestor + Distinguish, with examples, between homologous characteristics and analogous characteristics Homologous characteristics – are characteristics shared by two or more species that are derived from (were present in) a common ancestor. The structure of homologous characteristics is the same or similar among organisms, though the function may vary. Examples include the forelimbs of land vertebrates and the vertebral column of all vertebrates. Analogous characteristics – are characteristics shared by two or more species due to convergent evolution. These structures have the same function, but may be structurally different. Examples include insect and fly wings and the shark and cetacean dorsal fins. + Explain the biochemical evidence provided by the university of DNA and protein structures for the common ancestry of living organisms Early ideas about evolutionary relationships were based on anatomical evidence. It is now known that DNA is the molecule of inheritance used by all organisms and the genetic code is universal. Species hypothesized to be closely related to one another have more similar DNA sequences to each other. Protein sequences and structures are also similar among closely related organisms. Taken together, these biochemical data support the concept of common ancestry. 1. 2. http://en.wikipedia.org/wiki/Phylogenetics http://en.wikipedia.org/wiki/Systematics Marjorie Angelica Balaguer Baccalaureate Biology HL 11-1/2 + Explain how variations in specific molecules can indicate phylogeny (relatedness) Shared sequences of amino acids (proteins) and nucleotides (DNA) as well as the number of differences among sequences allow determination of relationships among organisms. In order to compare these data, alignments (comparisons) of sequences must be performed. For example: (DNA sequences are taken from both nuclear DNA and mitochondrial DNA) Species #1 #2 #3 #4 1 A A A A 2 A A A A 3 A A A C 4 A A A A 5 T C C T 6 T T T C 7 T T T T 8 T A A T 9 C C C C 10 C C C C 11 C C C A 12 C G G C 13 G G C G 14 G G G C 15 G G G T 16 G G G G Bold & Boxed – differences in sequence + Discuss how biochemical variations can be used as an “evolutionary clock” In addition to determining relationships, differences in DNA sequences can be used to estimate the amount of time that has passed since the divergence of two species from a common ancestor. However, because rates in mutation and evolution are not constant, there is some uncertainty (+/- x years) involved in determining divergence times. 1. 2. http://en.wikipedia.org/wiki/Phylogenetics http://en.wikipedia.org/wiki/Systematics Marjorie Angelica Balaguer Baccalaureate Biology HL 11-1/2 Phylogeny and Systematics II + Outline the methods used to construct cladograms and the conclusions that can be drawn from them. 1. Make a list of all of the organisms included in the study 2. List the characteristics included in the study 3. Make a table with the characteristics in the heading columns and the taxa (Kingdom, Phylum, Class, Order, Family, Genus or Species) in the rows. 4. Order characteristics left to right from the most widely shared characteristic to the lease widely shared. 5. Order the taxa top to bottom so the organism with the fewest traits is at the top, most at the bottom. *** The cladogram graphically illustrates the taxa organized in terms of shared characteristics derived from recent ancestors. + Sample cladogram: (2 step process) Step 1: Table and arrangement of taxa and characteristics *Table below is already ordered from least to greatest – it may not come out this way when observing real data or other information. Taxa Lamprey Sunfish Newt Lizard Bear Chimpanzee Human Backbone Jaw Forelimbs 1 1 1 1 1 1 1 0 1 1 1 1 1 1 0 0 1 1 1 1 1 KEY 1 – Yes, organism has characteristic 0 – No, organism does not have characteristic present 1. 2. http://en.wikipedia.org/wiki/Phylogenetics http://en.wikipedia.org/wiki/Systematics Amniotic Egg 0 0 0 1 1 1 1 Mamary Glands 0 0 0 0 1 1 1 Opposable Thumbs 0 0 0 0 0 1 1 Upright Posture 0 0 0 0 0 0 1 Marjorie Angelica Balaguer Baccalaureate Biology HL 11-1/2 Step 2: Cladogram with arrangement of taxa and characteristics Lamprey Sunfish Newt Lizard Bear Chimp Human Upright Posture Opposable Thumbs Mammary Glands Amniotic Egg Forelimbs Jaw + Discuss the relationship between cladograms and the classification of living organisms Cladograms are organized such that taxa are grouped based on evidence of relationships. Each derived characteristic that is added to a data set helps define a particular group (ex: hair defines mamalia) The Linnaean system of classification that provided names for many groups does not operate based on evidence for relationships, thus cladistics is now widely used for classification. 1. 2. http://en.wikipedia.org/wiki/Phylogenetics http://en.wikipedia.org/wiki/Systematics