Chapter 24 Obtaining Food in Animals Foraging: Animals show distinct behaviour patterns, organised to gain maximum energy when they go searching for food ( foraging). Problems they Face: availability of food competition predation physical barriers economy Examples of Foraging Behaviour Planaria: complete 1 Chapter 24 Search Patterns: 1) Bees: 2) Ants: 3) Higher animals: 2 Chapter 24 Economics of Foraging Behaviour Foraging itself is very costly. The costs will include energy losses due to handling, digestion and foraging for food as well as fighting to overcome prey, avoiding conflict with competitors and so on. Predators have to weigh up these losses against potential gains and select their options. Optimal foraging To be economic the animal must consume those food items which will give it the best return for the time and energy spent. If the energy gained from the food that it finds is less than the energy expended on the search, then the animal suffers a net loss of energy. Factors Affecting Foraging Behaviour and Subsequent Choice of Food Items 1. Time Search time - amount of time spent locating the food. Pursuit time - time spent obtaining the food once located. Where search time is short and pursuit time long e.g. lion and gazelles, then it is economic to be selective i.e. the lion picks off an old or weak gazelle. Where search time is long and pursuit time short e.g. a bird searching in dense foliage and finding a rich supply of easily captured insects, then it is economic to be non selective and eat as many prey items as possible since it may take a long time to find a similar haul. 3 Chapter 24 2. Unproductive versus Productive Ecosystem (a) In a poor ecosystem, the forager cannot afford to be choosy. If it ate only desirable prey, too much time and energy would be spent on the search. If it ate only poor prey, the predator would not gain enough energy. To be economical, it has to settle for a mixture of items with those of intermediate quality giving the best net energy return for the time spent on the search. (b) In a rich ecosystem, search time is reduced and the forager can afford to be more selective since it can obtain all of its energy requirements from a few types of choice food items. However, in some cases, it is economical for a herbivore to settle for the food available in a poorer ecosystem if foraging in a food-rich area exposes it unduly to the risk of attack by predators. Copy Fig 24.4 p 191 Effect of poor ecosystem on economics of foraging 4 Chapter 24 3. Size of Prey At first, net energy gain increases with increasing size of prey since larger items contain more energy than smaller ones. However, an optimum prey size is eventually reached. Beyond this point, net energy gain decreases because the very largest prey is the scarcest (hence involves a longer search time) and it tends to put up the best fight. It is not economical if the predator has to expend much energy subduing the prey. Copy Fig 24.5 p 191 Effect of food size on economics of foraging 5 Chapter 24 Competition 1) Interspecific Competition Arises when 2 organisms of different species occupy the same ecological niche i.e. the same position in a food web. Interspecific competition or a limited resource may become so fierce that one species ousts the other. Example 1. Paramecium: complete including graphs p192 6 Chapter 24 Example 2. Squirrels : complete Interspecific competition can be reduced when different species of animals in a community eat different food, nest in slightly different habitats or seek food at different times of the day etc. Example 3. Cormorants : complete 7 Chapter 24 2) Intraspecific Competition Arises when competition exists between members of the same resources in areas of overcrowding. Intraspecific competition is therefore even more intense than intraspecific competition when there is a scarcity of some resource e.g. food This will result in behavioural responses such as migration and aggression e.g. bird colonies, lemmings. Territorial Behaviour - Intraspecific competition often takes the form of behaviour which involves competition between members of the same species for territory. The size of this area defended (territory) will depend on a number of factors, one of the most important of which is the availability of food. Territorial behaviour can be considered in terms of costs and benefits. The major factors are: a) If a territory is too small, not enough food will be obtained. b) If a territory is too big, it costs too much energy too defend it. c) As food quality goes up, the size of the territory which needs to be defended goes down. d) Territories which are settled, result in reduced aggression, with consequent energy saving. Territorial Behaviour in the Robin complete 8 Chapter 24 Social Groups Dominance Hierarchy Complete: This is a system where ….. The linear form of social organisation in birds is called a …………. …………. Make note on dominance hierarchy in mammals List the advantages of dominance hierarchy 9 Chapter 24 Co-operative Hunting List 4 examples of animals which rely on co-operation between members of the social group to hunt their prey. Describe what is meant by: Ambush strategy Running down The dominant members of the social group are likely to get the biggest share of the kill. Explain the advantages of co-operative hunting to the social group as a whole which result in subordinate members of the group being willing to participate in co-operative hunting. 10