Week 5 Study Guide Big-picture topic — Explain the sequential adaptations to life on dry land illustrated in the rise of plants, and culminating in the most successful group, the angiosperms. (A) Discuss traits important to survival of the adult stage, including transport tissues and organs, adaptations that allow gas exchange while preventing water loss, and the switch to a dominant and independent sporophyte stage in the life cycle. (B) Discuss traits involved in reproduction, including (a) the feature that freed plants from their early dependency on environmental water for fertilization; (b) heterospory; (c) seeds; (d) double fertilization and nourishment of the developing embryo; (e) flowers and fruit, and how they allow symbiosis with animals that improves pollen transfer and seed dispersal Gymnosperms — Understand the following themes in plant evolution (be able to provide examples): - progressive reduction of gametophyte and dominance of sporophyte - improved vascular tissues allowing growth to greater heights above ground - greater protection of reproductive parts - pollen (immature male gametophyte) replaces swimming sperm Understand how the seed develops from the ovule, and be able to define these terms Know the basic life cycle of the pine tree. Where does meiosis occur? What is the multicellular haploid stage(s)? Understand adaptations that that enable conifers to survive in drought (= dry conditions), including tough needle-like leaves with dense tissues, thick cuticles, recessed stomata What is inefficient about the way most gymnosperms transfer their pollen to the female cones of other plants? What is a sporophyll? What part of a gymnosperm is the sporophyll? What part on an angiosperm is the sporophyll? Angiosperms — Know that angiosperms replaced gymnosperms as the dominant land plant group around 65 million years ago at the end of the Mesozoic Era, following a mass extinction event. Explain the key differences between angiosperms and gymnosperms, including: - flowers vs. cones - naked seeds vs. seeds enclosed in fruit - improved transport tissues (e.g. xylem with vessel elements) - role of animals in pollination and seed dispersal Be able to define flower and fruit and understand their basic parts, and where they fit in the life cycle. Understand the angiosperm life cycle; be able to explain double fertilization Be able to provide examples of heterotrophic (parasitic or carnivorous) angiosperms Understand the concepts of symbiosis and coevolution as they apply to angiosperms, animals, and fungi. In particular, understand that many angiosperms have mutualistic relationships with (1) mycorrhizal fungi, and (2) animal pollinators and fruit dispersers. Be able to give examples of pollination syndromes that illustrate (1) mutualism between plant and animal, (2) exploitation of the animal partner by the plant. How do angiosperms protect themselves from competitors and from herbivores? How does coevolution contribute to biodiversity -- meaning why does it tend to increase the overall number of species present in an ecosystem over time? Be able to explain the economic and ecological importance of angiosperms, and the human uses for plant defensive chemicals Terms to know -Angiosperm Flower Fruit Pericarp Ovule Seed Tracheid Vessel elements Pollination syndromes Double fertilization Pollen Sporophyll