N.G. Prilepsky, A.N. Demidova
Collection of Vietnamese plants in the Herbarium of Moscow University (MW)
E-mail: nprilepsky@mail.ru, DEMIDOVA_A@rambler.ru
Abstract. The article presents data on vascular plant collection from Vietnam stored at Foreign Asia
section of the Moscow State University (MSU) Herbarium named after D.P. Syreishchikov (MW) by
January 1, 2010. Collection includes 1607 specimens from 135 families and 382 genera. The article
contains data on collectors, places of collection and information on number of genera and families
presented in the collection. The essential part of the collection makes specimens collected by Nickolaj
G. Prilepsky at Ma Da forestland (Dong Nai province, South Vietnam) and Bien Lac – Nui Ong
forestry (Binh Thuan province).
Keywords: Vietnam, flora, Herbarium of Moscow State University (MSU), collection, collectors.
The purpose of the present article is generalization of data on collection of specimens from
Vietnam stored at Foreign Asia section of the Moscow State University (MSU) Herbarium named after
D.P. Syreishchikov by January 1, 2010. The Herbarium is known as «Moscow University Herbarium»1
and has its own international acronym – MW. Moscow University Herbarium is the oldest of now
existing Herbariums in Russia (was founded in 1765). It occupies the second place (after the
Herbarium of Botanical Institute of Russian Academy of Sciences named after V.L. Komarov in Saint
Petersburg), as far as size and scientific significance are concerned, among main Herbariums of Russia
(Balandin, 2006). Collections of the MSU Herbarium include more than 900 thousand specimens
(mosses and liverworts included), many of which have grate scientific, historical and cultural value. In
the resolution of the international conference «Conservation of botanical collections» that took place
under the auspices of UNESCO and International Council of Scientific Unions (Saint Petersburg, 8-10
December 1993) the MSU Herbarium is recognized as an Institution with collections of world
significance (Balandin, 2004). One of the most interesting collections of the MSU Herbarium is the
collection of Vietnamese flora2 being the part of Foreign Asia section.
By January 1, 2010 the Foreign Asia (without Mongolia, Central Asia and Kazakhstan)
collection included 17458 specimens from 6743 species, 2323 genera and 328 families. Vietnamese
collection (with exchange pool) by January 1, 2010 enumerated 1607 specimens. As a result of
inventory work at Moscow University Herbarium (Foreign Asia section) as well as on the basis of
literary sources, the authors of the present article have composed a database on all Vietnamese
specimens. It includes the following data: sheet number according to Herbarium Index, family name,
specific name (or generic name in case of incomplete identification), all possible synonyms according
to literary sources listed in this article, life form of plants, geographical names of collection points with
indication of provinces, districts etc., geographical coordinates, plant community names, collector
names, date of collection and different comments.
The following list includes collectors of Vietnamese flora whose collections are located in the
MSU Herbarium with comments on place and date of collection and number of sheets:
Averyanov L., N.Q. Binh, Phan Ke Loc – Lam Dong province, 31 km to the north-east of Da
Lat city, Lac Duong district, Da Chay municipality, in the neighborhood of Klong Lanh village – 1
sheet (1997);
Averyanov L., N.T. Ban, N.Q. Binh, А. Budantsev, L. Budantsev, N.T. Hiep, D.D. Huyen, P.К.
Loc, N.X. Tam, G. Yakovlev – Kon Tum province, north-west slopes of Ngoc Linh mountain system –
1 sheet (1995);
Arnautov N.N., Phan Ke Loc, Fam Van Nguyen, Maj Ngi, determined by Grushvitsky I.V.,
Skvortsova N.T. – Habak province, Tuan Mau settlement – 1 sheet (1970);
1
Formally Herbarium is the subdivision of the Geobotany Department of Biology Faculty of the MSU, but in all guides of
the world and among the specialists it is known as Moscow University Herbarium.
2
At present time the most significant herbarium from Vietnam (as well as from Laos and Cambodia) is located in France in
National Museum of Natural History in Paris. Part of the herbarium is located in Vietnam in the Institute of Tropical
Biology and Biological Resources (Ho Chi Minh city) and in the analogous Institute in Hanoi city (Kuznetsov, 2003b).
1
Belyaeva N.V.3 – Dong Nai province, to the north of Ho Tri An lake, Rang Rang neighborhood
– 2 sheets (1995);
Belyaeva N.V. – Lam Dong province, Da Lat plateau – 6 sheets (1995);
Belyaeva N.V. – Lam Dong province, Da Lat plateau, Cong Troi notch – 8 sheets (1995-1996);
Gordeeva М.М., Lazareva N.S., Kuleshova L.V.; determined by Nguen Tien Ban, Serebryanyj
М.М., Fam Dinh Tam, Bung Bang, Vu Nguen Ty – Hoa Binh province, Kuk Phyong national park –
138 sheets (1989);
Gordeeva М.М., Lazareva N.S., Kuleshova L.V.; determined by Serebryanyj М.М., Vu Nguen
Ty – Vinh Phu province, Tam Dao mountain massif – 10 sheets (1989);
Gorovoj P.G.; determined by P.G. Gorovoj and V.N. Tichomirov – Lao Cai province, Sa Pa
district – 7 sheets (1963).
Gruschwitski I.V. et al. – Hoa Binh province, Cai mountain – 1 sheet (1969);
Gruschwitski I.V., Arnautov N.N., La Dinh, Mai Ni et al., determined by Gruschwitski I.V.,
Skvortsova N.T. – Cao Bang province, Lea notch – 1 sheet (1972);
Gruschwitski I.V., Arnautov N.N.; determined by Gruschwitski I.V., Skvortsova N.T. –Hoa
Binh province, Pa Co – 1 sheet (1969);
Gruschwitski I.V., Arnautov N.N., Pham Van Nguyen, Phan Ke Loc; determined by
Gruschwitski I.V., Skvortsova N.T. – Nam Ha province – 1 sheet (1970);
Gruschwitski I.V., Arnautov N.N., Pham Van Nguyen, Phan Ke Loc, Nguyen Guang Hao,
determined by Gruschwitski I.V., Skvortsova N.T. – Hoa Binh province – 1 sheet (1970);
Gruschwitski I.V., Arnautov N.N., Pham Van Nguyen, determined by Ha Thu Zung, Mai Ni –
Ninh Binh province, Kuk Phyn nature reserve – 1 sheet (1969);
Gruschwitski I.V., Arnautov N.N., Pham Van Nguyen, Phan Ke Loc, Mai Ni, determined by
Gruschwitski I.V., Skvortsova N.T. – Ha Bac province, Hung Son settlement – 1 sheet (1969);
Gruschwitski I.V., Arnautov N.N., Pham Van Nguyen, Phan Ke Loc, Ha Thu Zung, Mai Ni, So
Va Ni; determined by Gruschwitski I.V., Skvortsova N.T. – Hoa Binh province, Pa Co settlement – 1
sheet (1969);
Gruschwitski I.V., Arnautov N.N., Pham Van Nguyen, Phan Ke Loc et al., determined by
Gruschwitski I.V., Skvortsova N.T. – Hoa Binh province – 1 sheet (1969);
Gruschwitski I.V.; determined by Gruschwitski I.V., Skvortsova N.T. – Hoa Binh province,
about 7 km to the south-west of Hoa Binh city, near the river, within Nha Bo settlement – 1 sheet
(1969);
Gruschwitski I.V., Pham Van Nguen et al., determined by Gruschwitski I.V., Skvortsova N.T.
– Hoa Binh province, Lac Son district, to the south of Vu Ban district center. – 1 sheet (1969);
Gruschwitski I.V., Phan Ke Loc, Pham Van Nguen; determined by Ha Thu Zung – Hoa Binh
province – 1 sheet (1969);
Dao Tan; determined by Phan Ke Loc –province Vinh Phu, Tam Dao – 3 sheets (1959);
Kuznetsov G.V.; determined by V.N. Pavlov et al. – interfluve of upper reaches of Ba and Kon
rivers within Gia Lai and Kon Tum provinces – 68 sheets (1990);
Lichnova O.P.4 – Lam Dong province, Da Lat plateau – 1 sheet (1995);
Lichnova O.P. – Lam Dong province, Da Lat plateau, Cong Troi notch – 101 sheet (1995);
Lichnova O.P. – Lam Dong province, Da Lat plateau, Phi Lieng – 61 sheet (1995);
Mikeshin G.V. – Quang Tri province, 5 km to the north of Vinh Linh – 2 sheets (1963);
Mikeshin G.V. –Nghia Lo province, in Phuyen neighborhood – 1 sheet (1963);
Mikeshin G.V.; determined by V.D. Bochkin – Quang Tri province, 5 km to the north of Vinh
Linh – 1 sheet (1963);
Mikeshin G.V.; determined by O.V. Daeva – Lai Chau province, Thuan Ciao district, Ma Lu
Thang mountains, Thanh Hoh neighborhood – 3 sheets (1963);
Mikeshin G.V. – Iter 5 – 8 sheets (1963);
Mikeshin G.V. – Iter 6 – 2 sheets (1963);
Mikeshin G.V. – Quang Binh province, near Dong Hoi city – 1 sheet (1963);
Mikeshin G.V. – place is unknown – 2 sheets (1963);
3
4
Specimens collected by N.V. Belyaeva were determined by N.G. Prilepsky.
Specimens collected by O.P. Lichnova were determined by N.G. Prilepsky.
2
Mokievsky V. – Siam bay, Phu Quok island – 12 sheets (2009); Khanh Hoa province, Nha
Trang city and Nha Trang bay – 41 sheet (2007-2009); Khanh Hoa province, Hon Heo peninsula – 3
sheets (2008);
Nuraliev M.S.; determined by Nuraliev M.S. – Lao Cai province, Sa Pa district, Hoang Lien
national park – 74 sheets (2009);
Prilepsky N.G.5 – Ho Chi Minh city – 7 sheets (1995-1996);
Prilepsky N.G. – Ba Ria – Vung Tau province, Vung Tau city neighborhood – 46 sheets (19951996);
Prilepsky N.G. –Binh Thuan province, Bien Lac – Nui Ong forestry – 122 sheets (1995);
Prilepsky N.G. – Khanh Hoa province, Nha Trang city neighborhood – 6 sheets (1996);
Prilepsky N.G. – Lam Dong province, Cat Tien national park – 2 sheets (1996);
Prilepsky N.G. – Dong Nai province, to the north of Ho Tri An lake, Ma Da forest – 808 sheets
(1995-1996);
Sokolov D.D. – Ho Chi Minh city – 2 sheets (1995).
Phan Ke Loc, Le Minh Tuan; determined by Phan Ke Loc – Gia Lai province, Chu Prong town
– 2 sheets (1980);
Phan Ke Loc, Le Minh Tuan; determined by Phan Ke Loc – Dac Lac province, Dak Mil town,
Nam Da – 2 sheets (1980);
Phan Ke Loc, Le Munh Tunn; determined by Phan Ke Loc – Dac Lac province, Dak Mil town,
Nam Nung – 1 sheet (1980);
Phan Ke Loc, Nguen Hai Ha; determined by Phan Ke Loc – Gia Lai province, Chu Pah town –
1 sheet (1980);
Phan Ke Loc; determined by Nguyen Tu –Nam Ha province, Kim Bang Ba Sao – 1 sheet
(1976);
Phan Ke Loc, determined by Phan Ke Loc – Lang Son province, Bao Son district, Vu Le
settlement – 1 sheet (1970);
Phan Ke Loc, Phung Dang Bach; determined by Phan Ke Loc – Lang Son province, Bao Son
district, Vu Le settlement – 1 sheet (1970);
Khokhryakov A.P. – Ho Chi Minh city – 19 sheets (1979).
Tsyurupa P.; determined by A. Filonenko – place unknown – 13 sheets (1958).
Collectors unknown:
Collectors unknown, determined by Phan Ke Loc – Lao Cai province, Sa Pa district – 1 sheet (1957);
Collectors unknown – Gia Lai province, Quang Tram Lap – 1 sheet (1989);
Collectors unknown, determined by Tyavkov /?/ – Vinh Phu province, Tam Dao – 1 sheet (1981).
The most essential part of Vientamese flora herbarium is composed by specimens collected by
N.G. Prilepsky. This part of herbarium was collected during the period when he worked as a researcher
within South department of Russian-Vietnamese Tropical Center. The research was conducted since
July 1995 till May 1996 mainly at Ma Da forest (Dong Nai province) in the neighborhood of «Ma Da»
Field ecological station. The purpose of his work was to study diversity of plants and plant
communities of South Vietnam. Comparative study of vegetation of forest edges, forest gaps, forest
tracks, clear cuts and processes of their regeneration was conducted (Prilepsky, 1997).
Ma Da forest is closed humid flat (not higher than 50 m above sea level) forest with the
predominance of Dipterocarpus dyeri Pierre. The remaining fragments of dipterocarpus forest are
characterized by multilevel structure of forest stand and high floristic diversity. There are 4-5
underlevels in forest stand of such forests. More than 400 tree species (among them 140 being
background ones) and 130 species of lianas are present in the composition of dipterocarpus forest. The
second and third underlevels are characterized by maximal diversity. Numerous epiphytes and
hemiepiphytes occur in crowns and on trunks of trees. Herb layer in forest is fragmentary because of
high shade at soil surface (Kuznetsov, 2003а).
Part of specimens were collected by N.G. Prilepsky during short-term expedition to Bien Lac –
Nui Ong forestry (Binh Thuan province) in September 1995 as well as in Vung Tau neigborhood (Ba
Ria – Vung Tau province) in 1995-1996. He determined the majority of collected specimens. Plus to
5
Specimens collected by N.G. Prilepsky were determined by him.
3
this he determined specimens collected by O.P. Lichnova in December 1995 and N.V. Belyaeva in
April 1995 and March 1996 at Da Lat mountain plateau in Lam Dong province (mainly specimens
from Cong Troi notch). The specimens were determined with the help of “Illustrated flora of Vietnam”
(Pham- Hoàng Hô, 1991-1993); some plant names were specified according to “Illustrated flora of
South Vietnam” (Pham- Hoàng Hô, 1970-1972). Specification of some plant names was made with the
help of herbarium specimens stored at herbarium of Institute of Ecology and Biological Resources of
Ho Chi Minh city. Further specification and determination of some collected specimens is possible
only with the help of specialists on concrete taxonomic groups.
Of special interest is a collection of fruits of tropical plants collected by N.G. Prilepsky in
1995-1996 in Bien Lac – Nui Ong forestry (Binh Thuan province), Cat Tien national park (Lam Dong
province), Nha Trang neighborhood (Khanh Hoa province), Ma Da forest (Dong Nai province), Vung
Tau neighborhood (Ba Ria – Vung Tau province) and Ho Chi Minh city. The collection contains fruits
of: Acacia auriculaeformis A.Cunn. ex Benth., A. megaladena Desv., Afzelia xylocarpa (Kurz) Craib,
Ancistrocladus sp., Bombax sp., Buchanania sp., Ceiba pentandra (L.) Gaertn., Delonix regia (Hook.)
Raf., Dioscorea sp., Dipterocarpus sp., D. dyeri Pierre, D. turbinatus Gaertn. f., Garcinia sp., Gnetum
sp., Irvingia sp., Knema sp., Lagerstroemia duperreana Pierre ex Gagnep. (L. thorelii Gagnep.),
Lasiobema cardinale (Pierre ex Gagnep.) de Wit, Leea indica (Burm.f.) Merr. (L. sambucina Willd.),
Mimosa pigra L. (M. aspera M.E. Jones), Mussaenda sp., Pandanus sp., Parkia sumatrana Miq.,
Pecteilis susannae (L.) Raf. (Habenaria susannae (L.) R.Br.), Pteris decrescens H. Christ,
Pterospermum sp., Randia dasycarpa (Kurz) Bakh. f. (R. tomentosa Bl. in DC.), Sonneratia sp.,
Syzygium sp., Tacca sp., Ternstroemia sp., Uvaria micrantha Hook. f.&Thoms., Ventilago sp., Viscum
articulatum Burm. f., Vitex sp., Zingiber sp. (?).
The list of families and genera of plants represented in the collection of Vietnamese flora is
given below. Number of specimens for each family and genus is given in parentheses.
The list of plant genera according to families with indication of number of sheets in the
collection:
Acanthaceae (21) – Acanthus (1), Codonacanthus (1) Cyclacanthus (5), Peristrophe (2),
Phlogacanthus (6), Pseuderanthemum (2), Thunbergia (4).
Aceraceae (1) – Acer (1).
Actinidiaceae (1) – Saurauja (1).
Adiantaceae (18) – Acrostichum (4), Adianthum (3), Pteris (6), Taenitis (5).
Agavaceae (12) – Dracaena (12).
Aizoaceae (1) – Sesuvium (1).
Amaranthaceae (1) – Amaranthus (1).
Amarillidaceae (6) – Curculigo (6).
Anacardiaceae (5) – Buchanania (3), Mangifera (2).
Ancistrocladaceae (5) – Ancistrocladus (5).
Annonaceae (47) – Dasymaschalon (5), Desmos (2), Friesodielsa (4), Goniothalamus (7), Polyalthia
(14), Uvaria (15).
Apiaceae (14) – Bupleurum (1), Centella (4), Cnidium (1), Heracleum (1), Hydrocotyle (5),
Pimpinella (1), Sanicula (1).
Apocynaceae (28) – genus not defined (5), Aganosma (3), Alyxia (5), Melodinus? (1), Hollarrhena
(5), Ichnocarpus (4), Ochrosia (1), Parameria (4).
Araceae (8) – Alocasia (2), Amorphophallus (3), Raphidophora (2), Scindapsus sp.? (Raphidophora
sp.?) (1)
Araliaceae (43) –Acanthopanax (1), Aralia (1), Brassaiopsis (1), Dendropanax (3), Macropanax (1),
Schefflera (35), Trevesia (1).
Arecaceae (25) – Arecaceae (2), Calamus (4), Caryota (4), Korthalsia (4), Licuala (2), Pinanga (9).
Asclepiadaceae (11) – Calotrophis (4), Dishidia (3), Hoya (1), Streptocaulon (3).
Aspleniaceae (8) – Asplenium (7), Diplasium (1).
Asteraceae (24) – Asteraceae (3), Ageratum (4), Ainsliaea (2), Anaphalis (1), Anisopappus (1),
Bidens (2), Blumea (1), Erigeron? (1), Eupatorium (2), Gnaphalium (1), Gynura (1), Vernonia (4),
Wedelia (1).
4
Begoniaceae (1) – Begonia? (1).
Blechnaceae (3) – Woodwardia (3).
Bombacaceae (4) – Bombax (3), Ceiba (1).
Campanulaceae (4) – Codonopsis (1), Lobelia (2), Pratia (1).
Cannabaceae (1) – genus not defined (1).
Capparaceae (6) – Capparis ?(1), Capparis (4), Cleome (1).
Caprifoliaceae (1) – Viburnum (1).
Caricaceae (1) – Carica (1).
Celastraceae (4) – Euonymus (4).
Clusiaceae (6) – Cratoxylon (3), Garcinia (3).
Combretacae (11) – Combretum (5), Galygopteris (3), Lumnitzera (2), Terminalia (1).
Commelinaceae (7) – genus not defined (1), Commelina (1), Cyanotis (2), Floscopa (1), Forrestia (1),
Palisota (1).
Connaraceae (4) – Cnestis (1), Connarus (3).
Convolvulaceae (24) – Argyreia (5), Ipomoea (16), Merremia (3).
Cornaceae (2) – Cornus (1), Swida (1).
Cucurbitaceae (14) – Coccinia (3), Gymnopetalum (1), Hodgsonia (1), Melothria (1), Momordica (1),
Momordica? (2), Thadiantha (4), Trichosanthes (1).
Cupressaceae (1) – Fokienia (1).
Cyatheaceae (3) – Cyathea (3).
Cyperaceae (68) – genus not defined (1), Carex (13), Cyperus (21), Fimbristylis (6), Gahnia? (1),
Hypolytrum (6), Scirpus (10), Scleria (10).
Dennstaedtiaceae (2) – Sphenomeris (2).
Dilleniaceae (9) – Dillenia (4), Tetracera (5).
Dioscoreaceae (10) – Dioscorea (10).
Dipterocarpaceae (17) – Dipterocarpus (13), Hopea (1), Shorea (3).
Dryopteridaceae (6) – Tectaria (6).
Ebenaceae (13) – Diospyros (13).
Equisetaceae (4) – Equisetum (4).
Ericaceae (3) – Agapetes (1), Gaultheria (1), Vaccinium (1).
Erythroxylaceae (2) – Erythroxylon (2).
Euphorbiacaeae (83) – genus not defined (2), Aleurites (2), Antidesma (16), Breynia (3), Cleistanthus
(2), Excoeraria (7), Glochidion (1), Macaranga (9), Mallotus (22), Manihot (2), Microdesmis (5),
Phyllanthus (1), Sapium (5), Sauropus (5), Trigonostemon (1).
Fabaceae (Caesalpinioidae) (39) – Afzelia (1), Bauhinia (15), Caesalpinia (6), Cassia (2), Delonix
(2), Lasiobema (6), Mezoneurum (1), Pterolobium (3), Saraca (3).
Fabaceae (Mimosoideae) (40) – Acacia (13), Acacia? (1), Albizia (14), Archidendron (2), Mimosa
(9), Parkia (1).
Fabaceae (Papilionideae) (49) – Aeschynomene (2), Centrosema (2), Crotalaria (6), Dalbergia (4),
Derris (1), Desmodium (14), Dunbaria (4), Erythrina (2), Indigofera (2), Mucuna (4), Rhynchosia (1),
Uraria (6), Vigna (1).
Fagaceae (5) – genus not defined (1), Lithocarpus (3), Quercus (1).
Flacourtiaceae (2) – Casearia (1), Homalium (1).
Flagellariaceae (1) – Flagellaria (1).
Gentianaceae (3) – Centaurium (1), Gentiana (2).
Gesneriaceae (3) – Paraboea (3).
Gleicheniaceae (1) – Gleichenia (1).
Gnetaceae (3) – Gnetum (3).
Goodeniaceae (3) – Scaevola (3).
Hamamelidaceae (1) – Rhodoleia (1).
Hemodoraceae (2) – Ophiopogon (1), Pelyosanthes (1).
Icacinaceae (11) – Gonocaryum (11).
Illiciaceae (1) – Illicium (1).
Ixonanthaceae (2) – Irvingia (2).
Juncaceae (1) – Juncus (1).
5
Lamiaceae (3) – Anisomeles (1), Gomphostemma (1), Nosema (1).
Lardizabalaceae (1) – Stauntonia (1).
Lauraceae (2) – Cinnamonium (1), Litsea (1).
Lecythidaceae (8) – Barringtonia (8).
Leeaceae (8) – Leea (8).
Liliaceae (4) – genus not defined (1), Asparagus (1), Dianella (2).
Linaceae (1) – Reinwardtia (1).
Lythraceae (7) – Lagerstroemia (7).
Loganiaceae (4) – Strychnos (4).
Loranthaceae (3) – Taxillus (1), Viscum (2).
Lycopodiaceae (16) – genus not defined (2), Diphasiastrum (1), Huperzia (1), Lycopodium (12).
Magnoliaceae (2) – Magnolia (1), Michelia (1).
Malvaceae (33) – genus not defined (3), Abutilon (1), Hibiscus (10), Sida (6), Thespesia (5), Urena
(8).
Moraceae (18) – Artocarpus (4), Ficus (14).
Melastomataceae (29) – genus not defined (1), Medinilla (1), Melastoma (9), Memecylon (18).
Meliaceae (6) – Amoora (3), Chisocheton (2), Walsura (1).
Menispermaceae (1) – Menispermum (1).
Musaceae (1) – Musa (1).
Myristicaceae (11) – Knema (10), Maesa (1).
Myrsinaceae (31) – Aegiceros (8), Ardisia (22), Embelia (1).
Myrtaceae (16) – Callistemon (1), Psidium (2), Rhodomyrtus (5), Syzygium (8).
Nelumbonaceae (1) – Nelumbo (1).
Nyctagenaceae (1) – Bougainvillea (1).
Oleaceae (7) – Jasminum (7).
Onagraceae (2) – Jussiaea (1), Oenothera (1).
Ophioglossaceae (3) – Botrychium (2), Ophioglossum (1).
Orchidaceae (17) – genus not defined (4), Anoectochilus (1), Apostasia (1), Arundina (1), Calanthe
(1), Dendrobium (2), Habenaria (2), Haemaria (1), Ludisia (2), Microstylis (1), Pecteilis (1).
Orobanchaceae (1) – Aeginetia (1).
Oxalidaceae (1) – Oxalis (1).
Pandanaceae (4) – Pandanus (4).
Passifloraceae (3) – Passiflora (3).
Pinaceae (3) – Pinus (3).
Piperaceae (5) – genus not defined (1), Piper (3), Piper? (1).
Platanaceae (2) – Platanus (2).
Poaceae (61) – genus not defined – 1, Andropogoneae – genus not defined (1), Andropogon (2),
Arundo (1), Axonopus (Paspalum) (1), Botriochloa? Microstegium? (2), Centotheca (5), Chloris (1),
Diplachne (1), Eleusina (1), Eragrostis (2), Erianthus? (5), Eulalia (1), Exotheca (1), Imperata (9),
Oplismenus (3), Panicum (6), Paspalum (1), Pennisetum (8), Pogonaterum (1), Sacciolepis (2), Setaria
(1), Setaria? (1), Sporolobus (1), Thysanolaena (3).
Podocarpaceae (4) – Dacrydium (2), Podocarpus (2).
Polygalaceae (7) – Polygala (2), Salomonia (5).
Polygonaceae (8) – Fagopyrum (1), Polygonum (7).
Polypodiaceae (17) – Colysis (2), Colysis? (1), Drymoglossum (2), Drynaria (9), Pyrrhosia (3).
Proteaceae (1) – Helicia (1).
Restionaceae (1) – Leptocarpus (1).
Rhamnaceae (8) – genus not defined (1), Gouania (1), Ventilago (1), Zizyphus (5).
Rhizophoraceae (16) – Bruguiera (2), Carallia (7), Kandelia (1), Rhizophora (6).
Rosaceae (3) – Rubus (3).
Rubiaceae (91) – genus not defined (5), Canthium? (4), Geophila (1), Hedyotis (4), Ixora (14), Ixora?
(6), Keenania (1), Lasianthus (10), Morinda (3), Mussaenda (1), Mussaenda? (8), Ophiorrhiza (2),
Pavetta (5), Psychotria (8), Randia (9), Randia? (1), Saprosma (1), Saprosma? (3), Wendlandia? (5).
Rutaceae (23) – Clausena (5), Euodya (7), Glycosmis (5), Micromelum (2), Microsorium (4).
Santalaceae (3) – Dendrotrophe (3).
6
Sapindaceae (12) – genus not defined (4), Allophyllus (7), Xerospermum (1).
Sapotaceae (1) – Xantolis (1).
Schizeaceae (6) – Lygodium (6).
Scrophulariaceae (15) – genus not defined (3), Alectra (1), Angelonia (1), Lindernia (1), Mimulus
(1), Scoparia (1), Sopubia (1), Torenia (5), Veronica (1).
Selaginellaceae (13) – Selaginella (13).
Simarubaceae (13) – Eurycoma (8), Harrisonia (5).
Smilacaceae (5) – Smilax (5).
Solanaceae (8) – Brugmansia (2), Datura (2), Solanum (4).
Sonneratiaceae (10) – Duabanga (2), Sonneratia (8).
Sterculiaceae (13) – genus not defined (1), Helicteres (3), Pterospermum (5), Pterospermum? (1),
Scaphium? (2), Sterculia (1).
Symplocaceae (1) – Symplocos (1).
Taccaceae (4) – Tacca (4).
Taxodiaceae (1) – Cunninghamia (1).
Telipteridaceae (5) – Cyclosorus (5).
Theaceae (4) – Euria (3), Ternstroemia (1).
Thymeleaceae (2) – genus not defined (2).
Tiliaceae (33) – Colona (19), Grewia (14).
Ulmaceae (17) – Celtis (1), Trema (16).
Urticaceae (2) – Boehmeria (1), Pellionia (1).
Verbenaceae (70) – genus not defined (1), Avicennia (17), Callicarpa (6), Clerodendrum (16),
Clerodendrum? (1), Congea (4), Gmelina (5), Lantana (1), Sphenodesma (10), Vitex (9).
Vitaceae (18) – Ampelopsis (7), Cayratia (3), Cissus (4), Parthenocissus (1), Tetrastigma (3).
Xyridaceae (5) – Xyris (5).
Zingiberaceae (25) – Alpinia (3), Ammomum (2), Costus (2), Globba (5), Hedychium (1), Zingiber?
(8), Zingiber? Alpinia? (2), Alpinia? (2).
Family not defined (66).
Thus, the collection of Vietnamese flora in the Moscow University Herbarium numbers 1607
specimens collected in the period of 1957 till 2009. As a whole, the herbarium of Vietnamese
flora includes representatives of 382 genera, attributed to 135 families, and 66 specimens whose
family identity is not known. Families Fabaceae (128 sheets), Rubiaceae (91), and Euphorbiaceae
(83) are presented by most numerous specimens.
Acknoledgements
The authors are very much obliged to direction of Russian-Vietnamese tropical scientific-research and
technological center in the name of professor V.S. Rumak and A.N. Kuznetsov for help in organization
of field works in Vietnam and also to the chief of Geobotany Department of Biology Faculty of
Moscow State University, professor, corresponding member of Russian Academy of Sciences V.N.
Pavlov for help in identifying of some plants.
Biblioography
1. Balandin S.A. Herbarium of Moscow University during the last quarter of century (in the
context of world traditions and tendencies of herbarium management development) // Bulletin
of Moscow Society of Naturalists. – Moscow, 2004. – Vol. 109, issue 6. – P. 73-81.
2. Balandin S.A. Introduction // Herbarium of Moscow University (MW): history, modern state
and development perspectives / Editor-in-chief: S.A. Balandin / Moscow, 2006. – P. 5-9.
3. Kuznetsov A.N. Modern state of tropical forests of South Vietnam and their reaction on
treatment by herbicides during the Second Indochinese War // Investigation of terrestrial
ecosystems of Vietnam. Moscow – Hanoi: Geos, 2003a. – P. 11-44.
4. Kuznetsov A.N. Tropical dipterocarpus forest. Moscow: Geos, 2003b. – 140 p.
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5. Prilepsky N.G. Comparative characteristics of secondary and some primary plant communities
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6. Pham-Hoàng Hô. Cây-Có Miėn Nam Việt-Nam. An Illustrated Flora of South Vietnam. –
Saigon, 1970-1972. – Quyen I, 1970, 1114 p.; quyen II, 1972, 1139 p.
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