Ecosystems (Ecology is the study of ecosystems) ECOSYSTEMS ARE OPEN SYSTEMS Ecosystem – The interactions between all of the biotic components (the community) and the abiotic component (the habitat). The biotic components affect the abiotic components (oxygen in the atmosphere is a result of photosynthesis; the water cycle is strongly affected by transpiration in plants, etc.), and vice versa. The two (biotic and abiotic) are 100% interdependent. Community (biotic) – ALL of the life forms within the system, from microbes to plants, animals, and everything else… all means all. Producers are autotrophic… they make their own food (most are photoautotrophs, a few are chemoautotrophs). Consumers get their energy molecules form producers (herterotrophs). Decomposers are a sub-set of consumers… they recycle nutrients. Detriovores eat dead stuff, and they are sometimes considered decomposers, but the ‘real’ decomposers are bacteria and fungi. Habitat (abiotic) – temperature, moisture, minerals in the soil/water, atmospheric gasses, sunlight exposure, etc… ENERGY FLOWS THROUGH ECOSYSTEMS….MATTER IS RECYCLED. Energy flow through ecosystems. Most ecosystems are based on solar energy (photosynthesis). The photosynthesizers are PRODUCERS. Everything else gets its energy from another carbon source besides making their own carbon-based fuel from CO2. As energy moves from one trophic level to the next, most (90%) is lost. This can be attributed to the energy organisms use as they live, and to the 2 nd law of thermodynamics. The lost energy leaves the planet as heat and contributes to the entropy of the universe. The sun is still working, so ideally, the energy in = the energy out. This might be out of balance at the moment. TROPHIC LEVELS (Energy levels) The producers form the base trophic level of all ecosystems. Only 10% of the energy collected by producers is passed to the next level, primary consumers. Only 10% of the energy in primary consumers makes its way to the secondary consumers, and so on. From this we can calculate and/or infer the following: 1. There is a limit to the number of trophic levels (about 4-10, with most systems having about 5). 2. The total biomass energy of at each level is decreased by 90% as it moves to the next level. 3. There are fewer individuals at each increasing level (especially in the consumers), thus… 4. Top level consumers are the fewest in number. 5. Tropic levels can be represented by an “energy pyramid” with top-level consumers at the top level… duh. Feeding (energy acquisition) can be viewed as either a food chain or a food web. Food webs more accurately portray the complex feeding interactions in an ecosystem. Humans are top-level consumers. We are at the top of the food chain. This is a good thing in my opinion. Some species are more significant in ecosystems than others. Think of a house of cards… some cards can be removed but the house will stand. Others are essentially holding the house of cards up. The most critical species are called “keystone species.” When they go, the ecosystem may collapse. Humans are NOT keystone species. DIVERSITY is measured by the number of species in an ecosystem. As a general rule, the greater the diversity, the more stable the system. Succession: Like everything else, ecosystems evolve… they change over time. Especially when something alters the ecosystem in a dramatic way. When an ecosystem experiences a catastrophic event (fire, flood, volcanic eruption, etc..) many species may be lost. This creates a “biological void.” Species rush in to fill the vacated niches. The recovery of severely damaged ecosystem is called secondary succession. ‘Secondary’ because not everything was killed off. Primary succession occurs when an ecosystem is obliterated and completely sterilized. The eruption of Krakatoa in the early 1800s made an island disappear…. Poof. Magma rising from underwater then created a new island that was completely devoid of life. This set the stage for primary succession. Primary succession is rare. Secondary succession is common. When an ecosystem evolves to a point where all of the interactions are in balance and sustainable, we say the ecosystem has reached its climax state. Species Interactions: The relationship between any two (or more) species in an ecosystem can usually be identified. Here are the major examples: 1. Herbivory – an animal eats a plant. 2. Predatory/prey – an animal eats another animal 3. Symbiosis – a long-term, intimate relationship. a. Parasitism – parasite/host b. Mutualism – both partners benefit c. Commensalism – one partner benefits, the other is unaffected. (Moss grows on trees… the moss benefits from getting into the light, the treat is neither helped nor harmed.) Niche: An organism’s niche is defined as the sum of its interactions with the biotic and abiotic components of an ecosystem. Any organism’s niche is limited by competition from other organisms. This is called competitive exclusion. All organisms compete in some way. You might even think of competition as a category of interaction, but it is common to all organisms in one way or another. Population growth: Since all organisms produce more offspring than can possibly survive owing to competition, predation, herbivory, disease, etc… We can model population growth mathematically. If an organism is placed in a habitat with (theoretically) unlimited resources and no competition, it’s growth will be exponential. We call this exponential growth. At some point, the population size becomes selflimiting owing to competition – even from within. Exponential growth begins to level out. This is logical. Therefore it is called logistic growth. If we look at the population growth curves of two species in a predator/prey relationship, we see that it is cyclic. Human population growth is currently exponential. Exponential growth is NEVER sustainable. The carrying capacity for any organism is the population that the ecosystem can support indefinitely. It is a topic of debate, but many scientists calculate that human population (currently between 6 and 7 billion) has already passed its carrying capacity. Cycles: Matter cycles, and cycles matter. You know about the oxygen/carbon dioxide cycle. You need to also understand the nitrogen cycle and the water cycle (we have alluded to these earlier). All of these go through a phase that involves atmospheric gases. The phosphorous cycle does not involve an atmospheric stage. Phosphorous cycles through water and soil.