AP Environmental Science Ecosystems: Relationship - chapter 5 text web site | main class page | Gonzaga page I. Community structure refers to the physical appearance, species diversity and abundance, and ecological relationships in an ecosystem (p. 144) A. biodiversity varies across different ecosystems 1. types of biodiversity: ecological, species, genetic, functional (Miller p.91) 2014 Bioblitz at Kenilworth gardens in DC 2. in general, diversity increases as you move toward the equator 3. diversity also varies with depth in water 4. pollution has a significant effect on diversity in aquatic systems (such as streams and the Chesapeake) 5. Alpha, beta, and gamma diversity 6. Diversity indeces | richness vs. evenness B. island biodiversity varies with island size and distance to the mainland 1. number of species on an island depends on rates of immigration and extinction 2. larger islands have more species 3. nearer island have more species 4. research supports these theoretical projections 90% - 50% rule C. types of species – ecological roles 1. species can be described by their mode of obtaining energy (predator, decomposer, etc) – see chap 3 2. non-native species cause problems (more on this later) 3. indicator species are particularly sensitive to environmental problems (amphibians are a good example; also certain stream insects) 4. umbrella species: A recent article in National Parks calls the Greater Prairie Chicken an umbrella species , because each type of habitat required by this bird also serves as important habitat for many other species, including deer and other prairie birds 5. are all species vitally important? Compare the rivet and redundancy models 6. keystone species have a disproportionate effect on the rest of the ecosystem a. top predators such as wolves, cougars, or grizzly bears Do cougars still exist in the eastern US? check out these sites: Michigan cougar | save the cougar | Eastern Cougar study suggests that predators exert "topdown control" on ecosystem structure learn more about the "top-down control" from Where the Wild Things Were the effects of wolves in Yellowstone | Spring 2010 Nat Geo article Ecological effects of cougars Lords of Nature video explores this phenomenon b. ecosystem engineers such as elephants, beavers, buffalo, prairie dogs, and even ants | more on buffalo as keystones beavers causing problems in suburban areas c. providers of important services, such as pollinators | more here - match pollinators with their flowers | Another pollinators page the starfish Pisaster is a classic example from rocky tidal zones of the western North America coast more on wolves American Chestnuts d. Miller suggests that the term "Foundation Species" is broader D. competition 1. intraspecific competition (within the same species) leads to natural selection, but does not affect community structure 2. interspecific competition refers to competition between different species for resources such as food and habitat 3. interference competition – individuals chase each other away or otherwise physically exclude them 4. exploitation competition – individual vary in how efficiently they use a resource 5. the competitive exclusion principle states that two species cannot occupy the same niche (classic experiment with mussels described in class) lynx and bobcats in North America 6. species reduce competition through resource partitioning - fig. 5-2 and also 4-16: they use different parts of the same habitat or forage at different times E. predator-prey relationships – fig. 5-6 deer have become a major pest in suburban areas because of lost predators. simulation of predator-prey relationships using wolves and rabbits Oak trees, mice, ticks, and deer have complex relationships (explore with this interactive) F. non-predatory ecological relationships 1. interactions between species can be positive, negative, or neutral overview of the types these relationships are examples of co-evolution | more here on co-evolution 2. parasitism is a relationship in which one species benefits and the other is harmed parasites and the red queen hypothesis -- running just to stand still derive nutrients and energy from the blood/body fluids of another organism (note examples) Can have very complicated life cycles – often two or more hosts (example – deer ticks) Cryptosporidium and Giardia are parasites found in water -even in the wilderness!! slide show of parasites that take over their host bodies 3. mutualism is a relationship in which both species benefit (fig. 59) a. symbiosis – organisms living on or in each other i. lichens a professor in Wisconsen has created a list of fungi of the month ii. myccorhizal fungi pictures from the microbe zoo iii. termites b. non-symbiotic i. herbivores and grasses ii. dispersal of fruits by birds and small mammals iii. pollination flowers display many interesting designs adapted to pollination the forgotten pollinators -many are going extinct or getting rare! iv. cleaner fish v. animals that are normally enemies will sometimes form mutual relationships 4. commensalism - one organism benefits and the other is not harmed. (fig 5-10) a. ephiphytes (distinguish from parasitic plants) b. barnacles c. remora and shark II. Ecological succession A. Primary succession 1. Occurs in an area previously unoccupied by organisms 2. Examples – volcanic islands, places exposed by retreating glaciers, etc. Glacier Bay National Park has glaciers that have retreated dramatically in the past 200 years (more on Glacier Bay) vegetation at Mt. St. Helens is growing back after the 1980 eruption occasionally a flood can be severe enough to expose rock. One example is found at the spillway of a dam near DeMoines, Iowa | 3. example of typical progression shown for Isle Royale in fig. 511 B. Secondary Succession 1. Occurs in areas that have been disturbed by some natural phenomenon 2. examples of disturbances: fire, storms, floods, agriculture C. Examples: 1. field to forest (fig. 5-12) 2. pond to bog to meadow 3. fire-disturbed areas 4. sand dunes D. Climax communities 1. type of climax community is determined by local climate, soil, rainfall, etc. 2. Succession is not necessarily a predictable event, and some ecologists prefer the term “mature” to “climax” – see discussion p. 110 E. The ecological importance of fire 1. In many ecosystems, regular fires are an important part of the community structure 2. benefits of fires (see also links on right) | 3. history of fire management in the US Healthy Forest Initiative Ecosystems adapted to fire fire management on US public lands including a map o current fires and National Fire Plan | Incident Fire Management - an interactive list of current fires US Forest Service fire-fighting page fire-fighting jobs how to conduct a prescribed burn | more on prescribed fire Yellowstone fires of 1988: Yellowstone NP page on fir Fire Wars - NOVA special on fire o global maps of fires in 2000 o how plants are adapted to fire Global Fire Monitoring Center Fire ecology - plants and ecosystems that benefit from fire 2007 California wildfires and more on wildfire in general Ashes to Wildflowers - before and after photos and text describing ecological succession after a fire Southern California Chaparral | more here Google Earth view of current fires Life returns to the blast zone (Mt. St. Helens) Chaparral Southeastern coastal pine forest Grasslands