EVOLUTION DEFINITIONS AND EXAMPLES Fitness: An organism’s ability to survive and reproduce in a particular environment Gradualism: States that small evolutionary changes occur slowly and steadily over long periods of time- the process yields many transitional forms Punctuated Equilibrium: states that a population tends to remain relatively stable for long periods of time interrupted by periods of relatively rapid evolutionary change- explains gaps in the fossil record Speciation: evolving of a species Inbreeding: crossing of genetically similar organisms to maintain the presence of certain desirable traits- tends to increase homozygous genotypes for both desirable and undesirable traits- Ex. Purebred dogs are more likely to have joint deformities and blindness Outbreeding (hybridization): crossing of genetically dissimilar organisms to bring together the best of both organisms- tends to increase heterozygous genotypes and vigor (health). Microevolution: occurs when there are relatively small-scale changes in the gene frequency of a population- may result in new phenotypes or adaptations, but not a new species (changes within the species level) Macroevolution: occurs when there are relatively large-scale changes in the gene frequency of a population- results in a new species (or higher taxonomic group) Variation: Differences that exist between members of the same species (the raw material of evolution)- Ex. Size, color, blood type Adaptation: An inherited trait that helps an organism survive in its environment- a beneficial variation- Ex. Structural, behavioral, physiological Artificial Selection: selective breeding- technique that only allows those organisms with desirable traits to mate and produce the next generation Natural Selection: process in which organisms with favorable variations are more likely to survive and reproduce. These variations are passed onto the next generation- Ex. Antibiotic resistance in bacteria Population: a group of organisms of the same species that live in the same area Radiometric dating: rock or fossil may contain elements that are radioactive. The decay (breakdown) of these radioisotopes occurs at a constant rate and can therefore be used a geologic “time clock”. The parent isotope breaks down, through a series of steps, to form a daughter product. Scientists use the ratio of parent isotopes to daughter product to determine how many half-lives have passed and then calculate how old the same is. The age is given in years- A half life is the length of time it takes for exactly one-half of the parent atoms to decay to daughter atomsEx. Carbon 14 dating, Potassium 40 dating, Rubidium 87 dating, Uranium 238 and 235 dating- Axioms: the rate of decay of a radioactive isotope is measurable and constant; the isotopes of an element appear in a mineral in precisely the same ratio as they occurred in the environment; when an atom undergoes radioactive decay, its internal structure and chemical behavior change- Assumptions: we can determine the amount of parent isotope present when it was formed; leaching of the parent isotope out of the rock did not occur; the daughter product was not present when the rock formed nor was the rock contaminated by an infusion of the product- x=ln (n+) x= number of half lives passed ____No N+=parent isotope remaining ln (.5) No= parent isotope originally ln= natural log Stabilizing Selection: favors individuals with the intermediate phenotype (most common variation in the population) because they are more fit than those with extreme phenotypes (less common variations in populations)- evolution is minor or absent- Ex. Weight of human infants Directional Selection: favors individuals with an extreme phenotype because they are more fit than all the other phenotypes- evolution more in a specific direction- Ex. Size of human brains, size of horses, color of peppered moths Disruptive Selection: favors individuals with the extreme phenotypes that deviate in both directions from the population average. Both of the extreme phenotypes have higher fitness than the intermediate phenotypes- evolution into 2 populations with distinct traits- Ex. Beak size in African finches Analogous structure: refers to the part of 2 different species that are similar in function, but not in structure- Ex. Wings of a bird and wings of a butterfly SCIENTISTS, etc.: Cuvier: 1769-1832- Prominent scientist who studied fossils and was instrumental in the development of paleontology (particularly vertebrate). He also plays a key role in proving that extinctions did occur, He believes these are a result of periodic catastrophes (revolutions). He does not believe in evolution, he believes species are immutable (fixed and unchanging). He explains that after a catastrophe, species immigrate from other regions to repopulate those areas. Does not go as far as to suggest new creations after each catastrophe Bonnet: 1720-1793- (discovered parthenogenesis in aphids). Observed that fossilized organisms did not resemble modern organisms. He supported the idea of “evolution”, however he believed that there was an “evolutionary ladder”. Bonnet believed organisms perished in periodic, worldwide catastrophes that wiped out all life- organisms were brought back after “stepping up” a rung the “evolutionary ladder” Ex. Apes became humans, humans became angels Malthus- 1798 wrote an “Essay on the Principle of Populations”, Essay stated that human babies were being born faster than people were dying. He noted that the human population could outgrow their resources, such as living space and food supply. States that factors such as war, famine, and disease help keep population in check. Darwin will apply these principles to all organisms- not all of those born will survive, those that survive will have to compete for limited resources Lamarck: 1809- Believes the fossil record shows evolution has occurred and presents a theory to explain it. Theory includes Tendency Toward Perfection, Use and Disuse, and Inheritance of Acquired Characteristics Lyell: 1830-1833 book entitled “Principles of Geology”- Lyell studied geologic deposits and determined that geologic forces (erosion, glaciers, etc.) showed that earth was shaped by slow, progressive changes, not catastrophic events (e.g. floods). Therefore, Earth much be millions of years old (necessary for Darwin’s explanation of evolution)- He stressed that scientists must explain past geologic events in terms of processes that can actually be observed today (uniformitarianism)- Although Lyell rejects the idea of evolution for most of his life, he accepts it in the early 18h60’s and becomes a close friend and defended of Darwin Darwin: 1809-1882- sailed on the ship HMS Beagle as its naturalist from 1831-1826. He collected numerous samples and fossils. In the Galapagos Islands, he studied finches, tortoises, and iguanas. The diversity of these species greatly influenced his thinking on evolution. In 1859, he published the book “On the Origin of Species by Means of Natural Selection”- never uses the word ”evolution”, instead Descent With Modification Wallace: 1823-1913- developed a theory of evolution nearly identical to Darwin’s (“On the Tendency of Varieties to Depart Indefinitely from the Original Type”). He sent a copy to Darwin, asked for some advice, and told him that he was going to present his theory. This motivated Darwin to also present his research and theory to the Linnaean Society in 1858 EVIDENCE OF EVOLUTION: Fossils: the preserved remains or traces of ancient organisms- most accurate representation of the history of life on earth, the fact that the types of fossils present have changed over time implies that life on earth has changed over time (evolution)- many species that are alive today have no corresponding fossils during much of earth’s history. Implies that more recent species must have evolved from older species- younger fossils are usually more similar to modern organisms than are the older fossils. Often the fossil record shows transitional forms that reveal a progression of evolutionary changes- the earliest fossils are all unicellular organisms and the most complex are found most recently in fossil record- many species that were alive in the past are not alive today Homologus Structures: compares structures found in different species that are similar in structure, but not necessarily similar in functionstructures are similar to each other because they evolved from a common ancestor for both species Vestigial Structures: structures that have no or a reduced function in organisms- these structures evolved from an ancestor that had a greater need for them- difficult to explain their existence without evolution Comparative Biochemistry: compares the DNA and proteins of different species to see how similar they are- closely related species have similar DNA because they evolved from a common ancestor- the genetic code is universal because organisms have a common ancestor- at some point in time, chromosome number 2 combined 2 chromosomes to make the human chromosome 2. there is a remenance of telomeres. Now humans have 23 airs, but chimps have 24 Biogeography: studies the distribution of organisms in different parts of the world- explains why regions that are separated by large physical barriers, but that have similar environments, wound contain different species rather than the same species- this occurred because population evolved differently in one region than they did in another CONCEPTS: Lamarck’s Theory of Evolution: organisms constantly strive to improve themselves and become more advanced- the effort to improve cause the most used body structures to develop, while unused structures waste away- once a structure is modified by use or disuse, the modification is inherited by the organism’s offspring Darwin’s Theory of Evolution: there is variation within populations. Many variations are inherited and such traits are passed from parent to offspring- some variations are favorable, improving the organism’s ability to function and reproduce in its environment- more young are produced in each generation than can survive. Only a few survive long enough to reproduce (competition)- organisms that survive and reproduce are those with favorable variations. Because the offspring of these individuals will inherit favorable variations, a larger and larger proportion of each new generation will have these variations (natural selection)- over enormous periods of time, small changes accumulate, and population change (gradualism).