SECTION 3 THE UPPER PARAGUAY RIVER AND PANTANAL OF MATO GROSSO This section describes the physiographic and hydrologic characteristics of the Upper Paraguay River and the adjoining Pantanal of Mato Grosso. It is divided into four subsec tions: 1.Geologic setting, 2.Geomorphologic setting, 3.Hydrologic setting, and 4.Ecologic setting. The information of this section serves as the basis for the analysis of Section 4. The references sources are listed in Appendix 1. The figures and tables are included in Appendices 2 and 3, respectively. 3.1 Geologic Setting The geology of the Upper Paraguay river and environs is described in Almeida (1945); Projeto Bodoquena (1979), Projecto RADAMBRASIL (1982) (Volumes 27 and 28), and Godoi Filho (1986). The predominantly Upper Precambrian formations underlie exten sive quaternary deposits, with significant rock outcroppings. Geomorphologic evidence reveals the presence of substantial tectonic activity in the form of subsidence and uplift (DNOS, 1974; Orellana, 1979; Ab'Sáber, 1988). Table 5 shows the stratigraphic units present in the Upper Paraguay river. The Up per Precambrian is represented by the groups Alto Paraguay, Corumbá, and Jacadigo, and their respective formations; the Paleozoic by the Coimbra formation; the Mesozoic by the Alkaline Fecho dos Morros formation; and the Cenozoic by the Xaraiés and Pan tanal formations. Table 6 lists primary rock types, including sandstones, siltstones, lime stones, conglomerates, dolomites, calcareous dolomites, syenites, trachytes, calcareous tufa, and travertines. Table 7 shows the general location of rock outcroppings in the vicinity of the Upper Paraguay river. Navigation charts confirm the existence of numerous rock outcroppings on or along the Upper Paraguay river (Marinha do Brasil, 1974; and later dates). A list of these out croppings is shown in Table 8, and their possible rock types in Table 9. Four of these outcroppings are located in the middle of the channel (Passo Simão Nunes Inferior, Cór rego Bonfim, Farolete Balduíno, and Passo Mucunã), effectively functioning as grade controls. The rock outcroppings have been mentioned in INTERNAVE (1990), and rec ognized for more than a century (Leverger, 1862a; 1862b). These facts confirm that the Upper Paraguay river bed is substantially con trolled by the prevailing geology. While the sediments of the Pantanal for mation (quaternary) are the primary surficial geologic/geomorphic feature of the landscape, the longitudinal slope of the river is controlled more by the rock outcroppings than by the alluvium. As shown in Table 8, there are thirty-two (32) rock outcroppings within 1270 km of river, an average of one every 40 km. Other outcroppings along the river, particularly that of canga, may have been less thoroughly documented. According to Dorr (1945), canga is colluvium or talus, com posed in large part of fragments of iron-rich rock, which has been recemented into a coherent mass by limonite. In the vicinity of the Morro de Urucum, these outcroppings tend to have a linear distribution, suggesting the possibility of a fault (Dorr, 1945; Al meida, 1945). Moreover, it is noted that the Internave report mentions the occurrence of canga in several places along the river, including Porto Rabicho (km 2740), entrance to Passo do Conselho (km 2607), and upstream of Passo Piuva Inferior (km 2573) (IN TERNAVE, 1990). Table 10 describes three geologic faults in the vicinity of the Upper Paraguay river (Projeto RADAMBRASIL, 1982). The Falha da Lagoa, which crosses Gaíba Lake, may have a significant effect on the flow of the Upper Paraguay river. Likewise, the Falha da Penha, which also crosses Gaíba Lake, is strategically located in close proximity to the river (Fig. 5). According to DNOS (1974), it appears that the northern portion of the Pantanal has subsided in relation to the Serra do Amolar (Section 3.2). This would explain the elevated position of the older rocks in the Serra do Amolar. The essentially tectonic character of the Upper Paraguay basin has been admirably discussed by Freitas (1951). In this regard, Orellana (1979) has stated that active faults in a direction contrary to the regional runoff have created local sills (soleiras) which act to impede runoff. More recently, Ab'Sáber (1988) has attributed the genesis of the Pan tanal to "a tectonic character dominated by a system of faults which are geomorphologi cally contrary." A historical note of interest regarding rock outcroppings: In describing his passage through Lagoa Gaíba more than one hundred years ago, Captain Augusto Leverger noted the following (Leverger, 1862a): 3.2 Geomorphologic Setting The geomorphology of the Upper Paraguay river is described in Projecto RADAM BRASIL (1982) and Da Silva (1986). The geomorphic units in the vicinity of the Upper Paraguay river are: 1.Serras of Urucum-Amolar, 2.Serrana Province, 3.Depression of the Paraguay River, and 4.Plains and Swamps of Mato Grosso. The Serras of Urucum-Amolar comprise two groups of residual hills/mountains located to the right of the Upper Paraguay river, close to the Bolivian border. These serras are important because of their strategic location, on the right margin, and often right next to the river. The Serra do Urucum, southeast of Corumbá, comprises the morrarias of Urucum, Santa Cruz, São Domingos, Grande, Rabichão, and Tromba dos Macacos; further south, about 40-50 km, are the morrarias of Zanetti, Albuquerque, Mato Grande, Saiutã, and Pelada. The Serra do Amolar, about 60-160 km north of Corumbá, comprises the Serra do Amolar proper and the morrarias of Insua, Novos Dourados, Santa Tereza, Castelo, and others of smaller size. The elevation of these mountains varies from 300 to 900 m, with the highest point at Morro Grande (1065 m). The contact between the Serra of Urucum-Amolar and adjacent geomorphic units is often sudden, revealing the presence of faults of probable Cenozoic age (Projeto RADAMBRASIL, 1982). The Serrana Province consists of a group of serras (mountain ranges) of roughly parallel crest alignment, located east of Cáceres, in a predominantly SWNE direction. From SW to NE, these serras are: (1) Simão Nunes, (2) Colonia, (3) Acorizal, (4) Facão, (5) Primavera, (6) Ponta do Morro, (7) Quilombo, (8) Morro Grande, (9) Jacobina, (10) Barreiro Preto, (11) Chapola, (12) Boi Morto, (13) Bocainão, (14) Campina, and (15) Retiro (Folha SE.21-V-B, Projeto RADAMBRADIL, Volume 27). The Serra Simão Nunes, the most southwestern of the group, comes in direct contact with the Upper Paraguay river in several locations along 45 km of river, on the left mar gin, from Passo Acuri, at Km 2128 (Km 3368 of the Hidrovia), to Passo Morro Pelado, at Km 2083 (Km 3323 of the Hidrovia) (Table 20). The elevations in the Serrana prov ince vary from 300 to 700 m, the rock formations being those of the Alto Paraguay group (Table 5). The Depression of the Paraguay River comprises extensive plain surfaces sur rounding the Serrana province and to a lesser extent, the Serra of Urucum-Amolar. The relief is mild, slightly undular, with elevations varying between 150 to 250 m, and under lying Precambrian and Cambrian formations, which often outcrop through the quater nary deposits. The Plains and Swamps of Mato Grosso (Planícies e Pantanais Matogros senses) resemble an amphitheater of roughly semicircular shape, with its approximate center at Corumbá, comprising 28 percent of the Upper Paraguay river basin (Fig. 2). The plains are a series of mutually coalescing alluvial fans, surrounded by the Depression of the Paraguay River almost continuously to the east, and discontinuously to the north and south. They comprise an extensive surface of accumulation, of extremely flat relief, with elevations varying from 80 to 150 m, and subject to seasonal flooding by the Upper Paraguay river and its tributaries. Valley slopes are about an order of magnitude greater along the tributaries than along the mainstem. The tributaries cross the plains in a predominantly eastwest direction, with a slight orientation towards the center of the amphiteather, with valley slopes of 12 to 50 cm/km. The high sediment production of the tributaries has forced the mainstem Upper Paraguay, which runs predominantly from north to south, to closely parallel the western edge of the plains (Schumm, 1977). Moreover, its unusually mild slope (0.7-6.5 cm/km) causes it to regularly overflow its banks during the wet season. Sánchez (1978) has identified ten geomorphic subunits within the Plains and Swamps of Mato Grosso (Table 11). The Pantanal is a complex mosaic of geomorphic features or subunits, with a regional base level at the Upper Paraguay river, along the western edge of the plains. In turn, the base level of the Upper Paraguay river is at its mouth, at the confluence with the Apa river. The sequence of events leading to the formation of the Pantanal is de scribed in Projeto Bodoquena (1979) and more recently, by Ab'Sáber (1988). During the Jurassic period, the prevailing arid climate led to substantial sedi ment deposition, mostly in the form of sand dunes. The Cretaceous period that followed saw a change to a more humid climate, which transformed the desert into a flood plain with numerous lakes and swamps. The end of the Mesozoic era signaled the end of sedi mentation and the beginning of slow epirogenetic movements, which were generally positive. The subsidence of the Pantanal region occurred later, probably in the Pleistocene (Tri cart, 1982). A clear evidence of subsidence is the great depth of the quaternary depos its, which at Fazenda Piquiri reaches beyond 320 m, and at Fazenda São Bento, beyond 420 m (320 m below sea level). The maximum depth of the Pantanal sediments has been estimated at 500 m at the center, near the apex of the Taquari fan (Godoi Filho, 1986). Tectonic uplift to the south strangled the exit of the depression. In the last 6 million years, under the effects of subsidence and uplift, the Upper Paraguay river has been forced to carve an exit through the basal rocks to the south, a situation which ex plains its unusually mild slopes. 3.3 Hydrologic Setting The hydrology of the Upper Paraguay river is described in DNOS (1974) and EDIBAP (1979). The Upper Paraguay river flows from its headwaters in the Serra de Tapirapuã, in Mato Grosso, to its mouth at the confluence with the Apa river, in Mato Grosso do Sul. At Cáceres, where it meets the first human settlement of importance, the river drains an area of 33 860 km2; at the Apa river confluence, downstream of Porto Murtinho, it drains an area of 496 000 km2. The distance along the river, from Cáceres to the Apa river confluence is 1270 km. Throughout most of this distance, the Upper Paraguay river ad joins parts of the Pantanal of Mato Grosso, flooding the Pantanal during the wet season and draining it during the dry season. Thus, the hydrology of the Upper Paraguay river is effectively connected to that of the Pantanal. This relationship tends to blur the distinction between surface water and groundwater, significantly increas ing the complexity of hydrologic analysis. In fact, the peculiar geologic and geomorpho logic setting of the Upper Paraguay river and its neighboring Pantanal results in a unique hydrologic behavior, unmatched in the American continent. The prevailing climate is dry subhumid, grading to wet subhumid along a narrow strip paralleling the mountain ranges which delimit the basin to the north, east, and south, and humid in limited northernmost areas bordering with the Amazon basin (EDIBAP, 1979). The spatially averaged mean annual rainfall in the Upper Paraguay basin ranges from 1180 mm (Projeto RADAMBRASIL, 1984) to 1380 mm (EDIBAP, 1979), de pending on the data source. This fact alone would dictate that the mean runoff coeffi cient (the ratio of mean annual runoff to mean annual rainfall) should be close to the global average, estimated at 0.39 (for peripheral continental areas) by L'vovich (1979) or 0.46 by Berner and Berner (1987). Instead, the mean runoff coefficient of the Upper Paraguay is quite low, varying in the range 0.07-0.10, depending on which data is used in the calculation (Section 3.3.4). The abnormally low runoff coefficient of the Upper Paraguay river is a di rect result of its hydrologic interaction with the Pantanal. The latter functions as an inmense surface/subsurface reservoir which stores water annually and multiannu ally. In the annual time frame, the Pantanal stores water during the wet season and re leases it back to the mainstem river during the subsequent dry season. In the multiannual timeframe, the Pantanal stores water in a wet year and releases it back to the mainstem (primarily as baseflow) in a dry year. In this process, the slowness of which is com pounded by the unusually mild relief, large amounts of would-be runoff are instead returned to the atmosphere through evaporation and evapotranspiration. The latter helps to sustain a thriving ecosystem, characteristically a marsh or wetland, its biotic pro ductivity closely linked to the annual flood pulse (Junk et al, 1989). 3.3.1 Hydrography The hydrography of the Upper Paraguay river, from Cáceres to the Apa river conflu ence, is shown in schematic form in Fig. 6. At Cáceres, the Paraguay river has already received the contributions of two of its most important right-margin tributaries: the Sepo tuba and Cabaçal. From Cáceres, the river flows south towards Descalvados, a distance of 139 km along the river. At a point 71 km downstream of Cáceres, the Paraguay receives the contribution of its third important right-margin tributary, the Jauru. From Descalvados, the river flows first southeast and then turns south towards Porto Conceição, a distance of 121 km. At a point 46 km downstream of Descalvados, the river branches into two channels: the mainstem Paraguay to the right and the Bracinho to the left. This bifurcation marks the beginning of the Pantanal proper; from this point on, as far downstream as Amolar, the Paraguay river crosses extensive areas of lakes (baías) and adjoining permanently flooded plains. The two branches form the island of Taiamã, and rejoin 43 km downstream (measured along the mainstem). From Porto Conceição, the river flows first south and then turns southwest towards Bela Vista do Norte, a distance of 135 km. At a point 40 km downstream of Porto Conceição, the river again branches into two channels: the mainstem Paraguay to the right and the Caracará to the left. The two branches form the great island of Caracará and rejoin further south close to Refugio das Três Bocas. The mainstem Paraguay flows southwest towards Bela Vista do Norte, at the base of Morraria da Insua. The Caracará branch flows south towards Refugio das Três Bocas, in the vicinity of Serra do Amolar. The island of Caracará constitutes a veritable inland delta, with its apex at the bifurcation (40 km downstream of Porto Conceição) and its base the Paraguay river itself, which turns from Bela Vista do Norte southeast to Refugio das Três Bocas, flowing for a distance of 53 km. During extraordinary and exceptional floods, most of the island of Caracará is completely submerged. In the vicinity of the Morraria da Insua and the Serra do Amolar, the Paraguay river interacts hydrologically with three large lakes: Uberaba, Gaíba, and Mandioré. The larger of the three, Lagoa Uberaba, located north of Morraria da Insua, receives overflows from the Paraguay as well as runoff from Corixa Grande, the last significant right-margin tribu tary of the Paraguay, and local streams. Lagoa Gaíba is located between Morraria da Insua and Serra do Amolar. It consists of three smaller lakes: 1.Gaíba, surrounded by mountains to the east and west, 2.Gaíba-Mirim, surrounded by mountains to the east, west, and south, and connected to Gaíba during extremely high flows, and 3.Pre-Gaíba, a continuation of Gaíba to the northeast. The link between the Upper Paraguay river and Lagoa Gaíba is the Riacho da Gaíba. However, a branch of the Paraguay drains into Pre-Gaíba. The Riacho da Gaíba is generally as deep as the Paraguay, excluding the exit of Lagoa Gaíba, where it is extremely shallow (a depth of 0.1-0.6 m and a width of 2000 m). Judging by the unmixed colors of its waters, the Riacho da Gaíba appears to drain Lagoa Gaíba to the right (reddish color: iron-rich dissolved solids), Lagoa Uberaba to the center (dark color: humic colloids), and the mainstem Paraguay river to the left (light brown color: suspended sediments) (Fig. 7). The link between Lagoa Gaíba and Lagoa Uberaba is the Canal Pedro II, with a length of about 100 km. The direction of the current in the Canal Pedro II is normally from Lagoa Uberaba to Gaíba, but it can change seasonally if the flow is considerably reduced (DNOS, 1974). From Refugio das Três Bocas, the river flows south towards Amolar, a distance of 28 km. Shortly before reaching Refugio das Três Bocas, the river branches into two chan nels: the Paraguay and the Moquém. Before rejoining the Paraguay, the Moquém river branches into the Ingazal, which joins together with the São Jorge, another branch of the Paraguay. In turn, the São Jorge rejoins the Paraguay immediately upstream of Amolar. These bifurcations reveal the extremely small gradient in this section of the river. From Amolar, the river flows south towards Porto São Francisco, a distance of 58 km. At a point 46 km downstream of Amolar, the river flows past Riacho da Mandioré, the inlet to Lagoa Mandioré. The Paraguay river flows into Lagoa Mandioré during high flows, and out during low flows. From Porto São Francisco, the river flows in a general southwestern direction towards Corumbá, a distance of 146 km, and then east to Ladario, a distance of 7 km from Co rumbá. At a point 16 km downstream of Porto São Francisco, the Paraguay river branches again into two channels: the Paraguay and the Paraguay-Mirim. The latter crosses the plains and rejoins the Paraguay river 20 km downstream of Ladario. Be tween Porto São Francisco and Corumbá, the river can overflow to the right into the Lagoa Conceição, Lagoa do Castelo, and Lagoa Cáceres. The connection between Lagoa Cáceres, in Bolivia, and the Paraguay river is the Canal Tamengo, with its mouth in the vicinity of Corumbá. From Ladario, the river flows first east and then southeast to Porto da Manga, a dis tance of 69 km. At a point 32 km downstream of Ladario, the Paraguay river receives the contribution of the Taquari Velho, an ancient channel of the Taquari river, to the left. About 2 km upstream of Porto da Manga, the Paraguay river receives the contribution of the Taquari river to the left. The Negro river, a tributary of the Taquari, flows into the latter right upstream of its confluence with the Paraguay. From Porto da Manga, the river flows southwest to Porto Esperança, a distance of 58 km. At a point 24 km downstream of Porto da Manga, the Paraguay river receives the contribution of the Miranda river. Together with the Aquidauana river, its principal tribu tary, the Miranda river drains extensive areas of Pantanal and Upper Paraguay river ba sin to the southeast. From Porto Esperança, the river continues to flow southwest to Forte Coimbra, a dis tance of 67 km. Between 2 and 40 km downstream of Porto Esperança, the Paraguay river overflows during floods to feed its Nabileque branch. The latter crosses the plains east of the Paraguay in a general southern direction for about 250 km, eventually rejoin ing the Paraguay river at a point located 217 km downstream of Forte Coimbra. From Forte Coimbra, the river flows in a general southern direction for 239 km to wards Barranco Branco; then an additional 51 km to Fecho dos Morros, and from there, 36 km to Porto Murtinho. From Forte Coimbra to Porto Murtinho, the Paraguay river receives the contribution of several left-margin tributaries, including the Aquidabã, Branco, Tereré, and Amonguijá rivers, as well as small surface contributions from the Paraguayan Chaco on the right margin. At Fecho dos Morros, 36 km upstream of Porto Murtinho, the Paraguay river passes through a group of hills, which effectively constitutes a grade control. This control has been referred to as a syenite sill (DNOS, 1974). From Porto Murtinho, the Upper Paraguay river flows south for another 63 km to reach its mouth at the confluence with the Apa river. Table 12 shows selected hydrologic data at gaging stations along the Upper Paraguay river: drainage area, channel length, channel slope, and mean annual discharge. The fol lowing observations are made: ·The subbasin drainage areas are not precisely defined in several points, particularly between Descalvados and Amolar. This is due to the extreme complexity of the drain age patterns, including channel bifurcations and endorheic (closed) drainages. ·There is a slight reduction in channel slope and mean annual discharge from Descal vados to Porto Conceição, as the Paraguay river enters the Pantanal proper. This is where the Paraguay river begins to flood the Pantanal. ·There is a marked reduction in channel slope and mean annual discharge from Porto Conceição to Bela Vista do Norte, as the Paraguay river flows through and around the island of Caracará. ·At Refugio das Três Bocas-Amolar, the channel slope is further reduced to 1.82 cm/km, while the mean annual discharge at Amolar increases sharply (to 943 m3/s). This indicates the substantial contributions of surface and subsurface runoff immedi ately upstream of this point. ·Downstream of Amolar, the channel slope increases somewhat to Porto Esperança, while the mean annual discharge continues to increases gradually all the way to Porto Murtinho, close to the basin mouth. ·Downstream of Porto Esperança, the channel slope decreases again, reaching a value of 0.83 cm/km in the reach between Fecho dos Morros and Porto Murtinho. ·The mean annual discharge of the Upper Paraguay river at its mouth is estimated to be 1565 m3/s, based on measurements at Fecho dos Morros and Porto Murtinho. The average channel slope along the Upper Paraguay river, from Cáceres to Porto Murtinho, is 3.2 cm/km. As shown in Table 12, the channel slope varies between 0.83 cm/km (Fecho dos Morros-Porto Murtinho) to 6.54 cm/km (CáceresDescalvados), and the bed profile alternates between convex and concave. According to principles of fluvial geomorphology, a river that is free to move its bed eventually carves a concave upwards bed profile (Leopold et al, 1964; Christofolleti, 1980; Leopold, 1994). Thus, the documented convexi ties in bed elevation of the Upper Paraguay river reveal the presence of sub stantial geologic controls. These controls are operating in at least three reaches: 1.Refugio das Três Bocas-Amolar, with 1.82 cm/km, 2.Ladario-Porto da Manga, with 2.06 cm/km, and 3.Fecho dos Morros-Porto Murtinho, with 0.83 cm/km. The extent of the geologic control can be assessed by calculating the size of the hump at locations where its presence is suspected. For instance, from Table 12, the average channel slope from Refugio das Três Bocas to Porto São Francisco can be calculated to be 3.51 cm/km. Therefore, the hump at Amolar is: (3.51 - 1.82) cm/km X 28 km = 47 cm Likewise, the average channel slope from Ladario to Porto Esperança is 2.70 cm/km. Therefore, the hump at Porto da Manga is: (2.70 - 2.06) cm/km X 69 km = 44 cm The extent to which these humps can cause backwater in these channels of extremely small slope is evaluated in Section 4.1.1. A similar calculation at Porto Murtinho is not possible because of lack of data at the river's mouth. However, it is noted that the downstream river (the Middle Paraguay) has an average slope of 6 cm/km throughout its 797-km length (Anderson et al, 1993). This much larger slope downstream reveals that there is a substantial geologic control in the vicinity of Fecho dos Morros. Significantly, this is precisely the location of the sy enite sill that is mentioned in the literature (DNOS, 1974). 3.3.2 Flood Hydrology The flood regime of the Upper Paraguay river is a result of complex climatic interac tions at the various atmospheric scales. The climate is determined primarily by the ba sin's geographic location (latitude and continental location) and secondarily by its topo graphic relief and surface features. Mean annual rainfall varies from as high as 1800 mm at Chapada dos Parecis, the northernmost part of the basin, to as low as 800 mm in the alluvial fan of the Taquari river, close to the basin center (Projeto RADAMBRASIL, 1984). Within these limits, mean annual rainfall increases towards the mountains and high plains (planaltos) in the basin perimeter and decreases towards the alluvial plains in the basin center. Rainfall is concentrated in the summer months. The wettest threemonth period is December-February; the driest is June-August. The temporal distribution of rainfall has a tendency to vary spatially in a general north-south direction. The percentage of annual rainfall in the wettest three-month period is greatest in the north (48 percent at Cáceres), gradually decreasing toward the south (to 36 percent at Porto Murtinho). Thus, the northern portion of the basin is prone to flooding from tributary streams. On the other hand, the percentage of annual rainfall in the driest three-month period is smaller in the north (3 percent at Cáceres), gradually increasing toward the south (8 percent at Co rumbá, and 12 percent at Porto Murtinho) (EDIBAP, 1979). This indicates the possibil ity of local droughts recurring on an annual basis. In any stream, the number of flood peaks per year is a good indication of the extent to which surface runoff is being diffused (i.e., attenuated) by the prevailing geomorphic conditions. If the number of flood peaks per year is high, say more than 10, there is little runoff diffusion. Conversely, if there is only one flood peak, runoff diffusion is at its maximum. The Upper Paraguay tributaries have a number of flood peaks, following in tense storms that cover all or portions of their respective drainage basins. For instance, the Cuiabá river at Cuiabá has 15 flood peaks per year on the average; the Taquari river at Coxim has 18 flood peaks; the Miranda river at Miranda has 12 flood peaks (DNOS, 1974). A sequence of several flood peaks depicts the local nature of the floods as well as the absence of significant attenuation (the channel gradients vary from 7 to 50 cm/km, Table 3). Unlike its tributaries, the Upper Paraguay river behaves quite differently with respect to the number of flood peaks. This reflects both its milder gradient (0.7 to 6.5 cm/km) and the presence of the Pantanal, which stores and further attenuates the flood peak. The overall effect of this hydrologic process is a reduction in the number of flood peaks. At Cáceres, close to the entrance to the Pantanal, there is an average of five flood peaks per year. However, the number of flood peaks decreases markedly downstream, to one at Amolar, one at Ladario, and one to two from Ladario to Forte Coimbra. Downstream of Forte Coimbra, the number of peaks increases somewhat due to local contributions, but the locally-generated peaks tend to be much smaller than the peak propagated from upstream. The floods in the Upper Paraguay river have been classified as follows (DNOS, 1974; Carvalho, 1986): 1.Common floods, which are exceeded 75 percent of the time (return period < 2-yr), 2.Mean floods, which are exceeded 50 percent of the time (2-yr return period), 3.Extraordinary floods, which are exceeded 25 percent of the time (4-yr return pe riod), and 4.Exceptional floods, which are exceeded 10 percent of the time (10-yr return pe riod). The hydrographic records at Ladario (1900-95) show the strong attenuating capacity of the Pantanal upstream of this point. Throughout the 96-yr period of record, the flood wave at Ladario has always had a 12-month duration, i.e., one rise and one recession per year. The rise begins usually in December and finishes in June; the recession begins in June and finishes in December. The occurrence of the flood peak at Ladario varies with the flood level; it is accelerated (to early June, May, April, or late March) during extraordinary and exceptional floods, slowed down (to late June, July, or early August) during a typical flood year (common or mean flood), and again accelerated (to April or late March) during multiannual drought periods (Table 13 and Fig. 8). The latter behav ior is due to the drought flow being mostly contained within the river banks. Prediction of flood flows along the Upper Paraguay river using mathematical model ing was attempted by DNOS (1974) and EDIBAP (1979). Under a limited budget, the DNOS model continues to be operated to this date by the Companhia de Pesquisas de Recursos Minerais (CPRM), with offices in Rio de Janeiro. It is a difficult undertaking because of the temporal and spatial variability and complexity of the hydrologic proc esses, which includes high channel sinuosity, branching, overflows, endorheic surface drainage, and the presence of aquatic macrophytes in the surface waters. This is com pounded by the complex nature of the interaction between surface and subsurface water, since the Pantanal has a net gain of water in a wet year, and a net loss in a dry year (EDIBAP, 1979; Adámoli, 1986). The speed of propagation of the annual flood wave can be readily extracted from the discharge measurements. It has been established that a typical flood wave takes about 130 days to travel from Cáceres to Porto Murtinho. This represents an average speed of propagation of only 0.11 m/s. This extremely low value is due to the substantial contri bution of channel overflows to the overall flood wave propagation. Table 15 shows low flow, mean annual, and peak flood discharges along the Upper Paraguay river. Also shown are the ratio of peak flood to low flow discharge, and peak flood to mean annual discharge. The low values of these ratios (compared to other rivers in similar climatic settings) show that the Upper Paraguay river is very effective in de creasing the flood peaks, and correspondingly increasing the low flows. This is due to the presence of the Pantanal. Thus, the Pantanal is the geomorphic feature which provides the mechanism for the spreading (i.e., attenuation, diffusion) of flood floods and consequently, the increased permanence of low flows. 3.3.3 Low Flows and Droughts Table 14 shows the minimum seasonal water surface elevation at Ladario, from 1900 to 1994. The following conclusions can be drawn from these records: ·The minimum seasonal flow occurs usually late during the calendar year, in the month of November or December, although occasionally it may be advanced to Octo ber, or delayed to January or February. ·The minimum seasonal flow has been recorded at or below zero gage in the following years or drought periods: --In 1910 and 1915. --Five times during the nine-year drought period 1936-44. --In 1948. --Nine times during the ten-year drought period 1964-73. As shown by the Ladario gage records, the Upper Paraguay river has a tendency to ward multiannual droughts with a recurrence of 28-30 years. The lack of more extensive data precludes a more thorough analysis. Nevertheless, the tendency is borne out by the data and should be acknowledged. According to the EDIBAP study, the Ladario gage records, from 1900 through 1977, show a tendency for a decrease in both maximum and minimum seasonal flows (EDI BAP, 1979). In light of the wet period being experienced since 1974, this conclusion is no longer valid. 3.3.4 Basin Yield The complexity of the hydrologic setting in the Upper Paraguay river precludes a de tailed analysis of basin yield vs annual precipitation. However, an approximate analysis based on mean annual precipitation is possible. During a wet year, the annual precipita tion P is split in three ways: 1.Runoff R, 2.Vaporization V, which consists of evapotranspiration from vegetation, evaporation from water bodies, and evaporation from bare ground; and 3.Change in basin storage DS, which consists of surface, subsurface, and groundwater storage. Deep percolation is usually small or intractable, and can be neglected on practical grounds (L'vovich, 1979). Conversely, during a dry year, annual precipitation plus (a fraction of) basin storage go into runoff and vaporization. A simple water balance equation can be formulated as follows: P = R + V ± DS (3.1) where DS is positive during a wet year and negative during a dry year (Adámoli, 1986a). In an average year, where change in basin storage is reduced to a minimum, the above equation reduces to: P = R + V (3.2) from which average basin yield can be calculated. The spatially averaged mean annual rainfall in the Upper Paraguay basin is 1380 mm according to EDIBAP (1979), or 1180 mm according to RADAMBRASIL (1984). This estimation is based on isohyetal maps for the Brazilian portion of the basin (71 percent). Comparable isohyetal maps for the western portion of the basin, in Eastern Bolivia and Paraguay's Northern Chaco (29 percent), are not readily available. The mean annual discharge at the basin mouth (see Table 12) is 1565 m3/s. Accord ing to DNOS (1974), the mean of six years of annual discharge measurements (196571) at Porto Murtinho is 1212 m3/s. However, this value appears too low, since this was a particularly dry period (see Section 3.3.3). A longer and wetter period (1969-78) of measurement at Porto Murtinho (33 values) gives 2188 m3/s (Hidrologia S. A., un published data). The basin drainage area at its mouth is 496 000 km2 (Section 2.2). Given this information, the mean annual runoff coefficient Kr, i.e., the ratio of runoff to rainfall, can be calculated as follows: (1 565 m3/s) (86 400 s/d) (365 d/y) (1000 mm/m) (1 380 mm/y) (496 000 km2) (1 000 m/km)2 A similar calculation, assuming a mean annual discharge Qa = 1700 m3/s (for lack of better data), with P = 1180 mm/yr leads to: (1 700 m3/s) (86 400 s/d) (365 d/y) (1000 mm/m) (1 180 mm/y) (496 000 km2) (1 000 m/km)2 Thus, the mean annual runoff coefficient for the Upper Paraguay basin can be taken as Kr = 0.08. This value is interpreted to mean that on an average year, runoff at the basin outlet amounts to eight percent of rainfall, with the balance returned to the atmos phere as evaporation and evapotranspiration (i.e., the vaporization coefficient is Kv = 0.92). A similar analysis for the 11-year period 1965-76 showed a runoff coefficient varying in the range 7-14 percent, with a mean of 10 percent (EDIBAP, 1979). The above calculation confirms that the Pantanal functions not only as an attenuating mechanism for flood flows (and consequent increases in low flows), but also as an ab stracting mechanism for all flows, i.e., as an effective means of storing the would-be runoff and converting it instead into evaporation/evapotranspiration. Throughout mil lennia, this process has been responsible for sustaining the extraordinary biotic potential of the Pantanal (Section 3.4). By way of comparison, the mean annual runoff at Cáceres, at the northern entrance to the Pantanal, is 382 m3/s, and the contributing drainage area is 33 860 km2 (Table 12). The mean annual precipitation of the subbasin varies from 2000 mm at the head waters to 1300 mm at Cáceres (EDIBAP, 1979). This amounts to a runoff coeffi cient Kr = 0.22, which is 2.75 times that of the Upper Paraguay river at its mouth. Likewise, the mean annual runoff coefficient of the Paraná river at Corrientes (Argentina) has been calculated at Kr = 0.16 for the decade 1962-71, Kr = 0.19 for 1972-81, and Kr = 0.22 for 1982-91 (Ponce, 1994). These values are from 2 to 2.75 times that of the Upper Paraguay at its mouth. These calculations confirm that the substantial attenuating and abstracting property of the Upper Paraguay is due to the presence of the Pantanal, while the mean annual runoff coefficients of the Upper Paraguay river at Cáceres, immediately upstream of the Pantanal, and at the Paraná river at Corrientes, 964 km downstream of the Pantanal, depict more typical subhumid/humid basins. The annual potential evapotranspiration in the Pantanal varies spatially in the range less than 1100 mm to more than 1400 mm, according to the Thornthwaite method (Alfonsi and Camargo, 1986). However, measured pan evaporation data at Fazenda São João and Fazenda Rio Negro for 1971-72 show values of 1650 mm (Tarifa, 1986). The actual evapotranspiration in an average year can be estimated to be: Ea = 0.92 X 1180 mm = 1086 mm, or (3.5) Ea = 0.92 X 1380 mm = 1270 mm, (3.6) depending on which value of mean annual precipitation is used in the calculation (Projeto RADAMBRASIL, 1984; or EDIBAP, 1979). Thus, it is concluded that the Upper Paraguay basin is able to effectively evapotranspire the difference between rainfall and runoff in an average year. The calculation of runoff coefficient is based on surface runoff at the basin outlet, and does not include subsurface runoff at the basin outlet, which may be real but difficult to evaluate directly. The existence of a certain amount of subsurface runoff may be postulated on the basis that the mean annual discharge at Asunción, on the Middle Paraguay, 542 km downstream of the Apa river confluence, is 2700 m3/s (INTERNAVE, 1990). The increase of more than 1000 m3/s is difficult to explain, particularly since there are no major intervening drainages. For instance, OEA (1975) has calculated that the combined contribution of the Apa, Aquidabán, and Ypané rivers, which are measured, is about 180 m3/s. Based on areal comparisons, the contri bution of the Aguaray-Guazú and other unmeasured tributaries is estimated to be about 100 m3/s. The subsurface runoff, if it does exist in substantial amounts, would have the effect of reducing the vaporization coefficient Kv to a somewhat lower value, say, around 0.88-0.90, which is still high in comparison to other basins in similar climatic settings. 3.3.5 Sedimentology Measurements of suspended sediments, which include gravel, sand, silt, and clay parti cles, along the Upper Paraguay river have been scanty. The longest existing records are at Cáceres and at Porto Esperança. The Cáceres data, shown in Table 16, consists of 55 once-monthly measurements of sediment discharge taken between March 1977 and February 1982. The Porto Esper ança data, shown in Table 17, consists of 52 once-monthly measurements taken be tween April 1977 and November 1981. These measurements and related calculations were carried out by Hidrologia S.A. for the now defunct Departamento Nacional de Obras de Saneamento (DNOS). The methodologies utilized were the Modified Einstein and the FrijlingKalinske methods. The values shown in Tables 16 and 17 are total sediment discharge, consisting of: ·bed load, i.e., coarse particles (primarily gravel and sand) transported by rolling and sliding along the bed, ·suspended bed material load, i.e., coarse particles transported in suspension, and ·wash load, i.e., suspended fine particles, the concentration of which depends on source availability and not on the flow hydraulics. These measurements enable the following observations regarding total sediment trans port in the Upper Paraguay river: 1.The measured sediment concentration at Cáceres varies between 28 mg/liter and 507 mg/liter, with a mean value of 147 mg/liter, and a standard deviation 122 mg/liter (Table 18). In general, the low values are associated with low to average flows, while the high values are associated with high to exceptionally high flows. 2.The measured sediment concentration at Porto Esperança varies between 12 mg/liter and 897 mg/liter, with a mean value of 176 mg/liter, and a standard devia tion 183 mg/liter (Table 18). In general, the low values are associated with low to average flows, while the high values are associated with high to exceptionally high flows. 3.The correlation between sediment discharge and water discharge at both gaging sta tions is poor. This is partly due to the presence of the wash load, whose concentra tion does not depend on the water discharge. Generally, the wash load concentra tion is a function of the degree of natural or human-produced watershed disturbance upstream of the gaging station. Table 18 includes a summary of sediment discharge measurements along the main tributaries of the Upper Paraguay river: Cuiabá, Piquiri, Taquari, Aquidauana, and Mi randa rivers. Based on this limited by significant data, a preliminary sediment budget analysis is performed in Section 4.2. 3.4 Ecologic Setting The ecologic setting of the Upper Paraguay river basin and the Pantanal of Mato Grosso is unique in the American continent. The basin is strategically located contiguous to four major South American ecosystems, which surround it, exerting their influence on it (EDIBAP, 1979, Adámoli, 1986b): 1.The tropical Amazon rainforest to the north and northwest. 2.The subhumid savanna woodlands (cerrados) of Central Brazil to the northeast, east, and southeast. 3.The humid Atlantic forest (Floresta Atlántica) to the south. 4.The semiarid scrub forest (Chaco) of Eastern Bolivia and Western Paraguay to the west and southwest. The unusual combination of geology, geomorphology, and hydrology (see previous sections) has contributed to the richness and variety of the vegetation and associated mi croclimates of the Pantanal. In turn, this helps sustain a diverse ecosystem, where a complex assortment of permanent swamps, seasonal swamps, and terra firma is annually replenished with ample moisture, sediment, and nutrients. The entire process hinges on the high rate of vaporization (estimated at up to 92 percent on an average year) which characterizes the hydrologic budget of the Upper Paraguay river. 3.4.1 Flora In the existing literature, the Pantanal vegetation is often marked as a single unit and referred to as the "Pantanal" complex. Actually, the latter is a mosaic of many different communities, with frequent abrupt changes often correlated with topography, and many ecotones. The Pantanal has no endemic flora of its own; rather, it is made up of ele ments from mato (deciduous and semideciduous forests transitional to the Amazon tropi cal rainforest and the humid Atlantic forest), campo (open grassland), cerrado (savanna woodland), and caatinga (desert scrub). There are three main vegetation zones in the Pantanal (Veloso, 1947): 1.The aquatic and hydrophylous zone, 2.The hygrophylous zone, and 3.The mesophylous zone. The hydrophylous zone is permanently flooded. It is characterized by three vegeta tion types: (a) aquatics in flowing water (Eichhornia crassipes, Pistia, Elodea); (b) float ing aquatics in stagnant water (Eichhornia azurea, Marsiliea, Reussia subovata); and (c) aquatics largely rooted in shallow water (Echinodorus spp., Hydrocleis spp., Limnocha ris spp., Victoria amazonica). The hygrophylous zone is divided into: (a) permanently flooded, and (b) seasonally flooded swamps, with the latter usually dominated by one species. Plant communities of the seasonal swamps include the Thalietum (dominated by Thalia geniculata), the Cy peracietum (dominated by Cyperus giganteus), and the Ipomoeætum (dominated by Ipo moea fistulosa). The mesophylous zone coincides with noninundated alluvial soils. Many floristic asso ciations occur in this soil type, as well as many transitional areas (ecotones). Veloso (1947) classified associations in order of successional development, and concluded that the region is in an active state of change toward a more mesic forest. The most striking aspect of the Pantanal is its curious combination of mesic and xeric vegetation growing side by side, a result of its unique combination of climate and topog raphy. Toward the center of the Pantanal, close to Corumbá, the climate is markedly seasonal, with a clearly defined drought period. However, given the extremely flat to pography, a small difference in elevation is all that is needed to make a great difference in seasonal soil moisture, particularly when the underlying strata is coarse alluvium. Hoehne (1936) has referred to the Pantanal as a mixture of Amazonas (hylaea) and Ceará (caatinga), and provided examples of the two types of flora by contrasting the gi gantic candelabra cactus (Cereus peruvianus), and other cacti such as Opuntia stenar thra, with the aquatic Alismataceae and Victoria. Prance and Schaller (1982) have noted the strong cerrado element in the Pantanal. These cerrados are dominated by species such as Bowdichia virgiloides, Caryocar bra siliense, Curatella americana, and Qualea parvifora, which are typical of the savanna woodlands (Planaltos) of Central Brazil. Cerrado occurs mainly in the nonflooded up land, but also towards the eastern edge of the Pantanal, where the land is inundated for only short periods at the height of the flood season. Such wet cerrado tends to consists of numerous islands of cerradão (dense cerrado forest) on slightly elevated areas that are not flooded. Cerrado tree species which are most resistant to waterlogging (e.g., Cu ratella americana and Byrsonima crassifolia) are common near the boundary cer rado/campo, and on raised islands of ground in wet campos (Furley and Ratter, 1988). The distribution of these islands produces the campos de murundus, consisting of an expanse of wet campos dotted with a regular pattern of raised earthmounds bearing cer rado trees, shrubs, and often termitaria. The larger earthmounds, or capões, are circular or elliptical in shape, of lengths up to 300 m, and sparsely distributed across the season ally flooded campos (Ponce and Cunha, 1993). The sharpness of the campo/cerrado boundary has been documented by Eiten (1975). Within 1 m, or even 0.5 m, the change from the shrubs and low trees of the cerrado to the grassy layer without woody plants of the campo is complete. The reason for this abrupt change appears to be that the cerrado plants cannot establish themselves from seed in continually wet soil. In general, the campo occupies a site with a lower and more fluctuating water table, whereas the cerrado occupies the higher ground, where the soil seldom if ever remains saturated. In almost all cases, the cerrado stops suddenly at the edge of the campo, apparently due to the competition between the two vegetation types as whole plant communities. Cerrado species tolerant of waterlogging are able to grow in open campos in places where the soil level is only a few centimeters higher than elsewhere. The observation that larger islands on Pantanal landscape are thickly clothed with cerrado vegetation confirms that groundwater level exerts a precise control on the cerrado/campo boundary. In the Pantanal, the main trend of vegetational variation is highly corre lated with soil moisture and topography. The patent lack of trees in the wet cam pos is striking, particularly since a wide range of woody species successfully colonizes both the interfluves, which are drier than the campos, and the stream sides (riparian ar eas, or gallery forests) which are wetter. The absence of tall, woody species from areas which are intermediate in their physical characteristics is attributed to the fluctuating na ture of the water table and associated soil moisture. Thus, trees are able to tolerate both permanently wet (gallery forest) and moist-to-dry (cerrado) environmental conditions, but not an extreme alternation of saturation and desiccation (Cole, 1960). Areas subject to the latter are successfully colonized by the grassy elements (campos). In summary, the Pantanal is extremely rich in floristic diversity and physi ognomic composition. It floristic diversity is due to its location, in the middle and neighboring four great South American ecosystems: the tropical Amazon rainforest, the subhumid savannas of Central Brazil, the humid Atlantic forest, and the semiarid scrub forest of the Chaco. Its diverse physiognomic composition is due largely to its variety of geomorphic/topographic features, which include baías, barreiros, cordilheiras, vazan tes, corixos, capões, murundus, and aterros de bugre (Cunha, 1990; Ponce and Cunha, 1993). The annual flood pulse replenishes the Pantanal ecosystem with water, sediment, and nutrients, assuring its continuance and survival (Junk et al, 1989). 3.4.2 Fauna The ecological diversity of the Pantanal ecosystem has conditioned its suitability as habitat for a variety of animal species, among which are reptiles, mammals, birds, fish, and insects. The Pantanal remains a rich and unique repository for a variety of wildlife species, including 658 species of birds, 1132 species of butterflies, and 405 species of fish (Brown, 1986; Bucher et al, 1993). It also serves as the resting place for many species of migratory birds from the Northern Hemisphere. Terrestrial and amphibious species inhabiting the Pantanal include (EDIBAP, 1979; Bucher et al, 1993): ·caiman (jacaré, Cayman crocodilus) ·armadillo (tatú bola, Tolypeutes tricinetus) ·bush dog (cachorro do mato vinagre, Speothos venaticus) ·capybara (capivara, Hydrochoerus hydrochaeris) ·crab-eating fox (cachorro do mato, Dusicyom thous) ·giant anteater (tamandúa bandeira, Myrmecophaga trydactyla) ·giant armadillo (tatú canastra, Priodontes giganteus) ·giant otter (ariranha, Pteronura brasiliensis) ·jaguar (onça pintada, Pantera onça) ·maned wolf (lobo guará, Chrysocyon brachyurus) ·marsh deer (cervo do pantanal, Blastocerus dichotomus) ·neotropical river otter (lontra, Lutra longicaudis) ·ocelot (jaguaritica, Felis pardalis) ·pampas deer (veado campeiro, Ozotocerus bezoarticus) ·rhea (ema, Rhea americana), ·tapir (anta, Tapirus terrestris), and ·peccary (porco monteiro, Tayassu pecari) These species selectively inhabit the campos, capões, cordilheiras, gallery forests, and water courses (baías, vazantes, corixos) of the Pantanal. Such impressive biodiver sity is due in large measure to the unusual climatic, geologic, geomorphologic, and hy drologic setting of the Pantanal. Wildlife management in the Pantanal has been dis cussed by Dourojeanni (1980), Paiva (1984), and Alho (1986), among others. Wildlife conservation in the Pantanal, particularly with regard to the jaguar (Pantera onça), has been discussed by Quigley and Crawshaw (1992). _ _ _ "...Virando a Norte, tive de circumdar um como promontorio da margem do Poente da lagoa o qual é terminado por um cabeço pedregoso e co berto de mato, ao cual devese dar resguardo a fim de evitar as muitas pedras que o cercam, umas submergidas, outras a flor de agua ou pouco elevadas..."