1
Online Resource 5. Breeding history/origin summaries for 78 Agrostis, Apera, and Polypogon cultivars, accessions, and collections in
the current study. Entry order follows the order presented in the UPGMA diagram (Fig. 1), with the exception of the outgroup species
of Apera and Polypogon.
Entry
007 A. stolonifera
Breeding History or Origin
Developed using 23 parent plants – 19 plants originating from Rutgers University
breeding program and 4 plants originating from Blue Moon Farms (Lebanon, Oregon).
Rutgers clones: selections from ‘L-93’ and ‘Southshore’; collections tracing back to
putting greens from the Piping Rock Club (Locust Valley, NY), Spring Lake GC (Spring
Lake Heights, NJ), Northshore CC (Glen Head, NY), Rumson CC (Rumson, NJ), and
Harkers Hollow Golf and CC (Harmony, NJ); and collection from a golf course fairway
in Chicago, Illinois. Blue Moon Farms clones: trace to germplasm from L-93 and
collection from old putting green in Atlanta, GA. (Rutgers, the State University of New
Jersey and R.H. Hurley 2009)
Pinup A. stolonifera
Maternal progenies of 13 clones collected from old golf course putting greens and
fairways – 46% of the germplasm tracing to collections from the Piping Rock club
2
(Locust Valley, NY); 31% of the germplasm tracing to a plant collected from Harkers
Hollow Golf and CC (Harmony, NJ); and 23% of the germplasm tracing to a plant
collected from National Golf Links of America (Southampton, NY). The creeping
bentgrass plants selected from old golf courses were of unknown origin, however, it is
likely that these plants originated from South German mixed bentgrass as this was one of
the few commercially available sources for bentgrass seed in the early 1900’s. (Stacy
Bonos – personal communication)
AP18 A. stolonifera
Maternal progenies of 10 clones collected from old golf course putting greens and
fairways – 10% tracing to a plant selected from Harkers Hollow Golf and CC (Harmony,
NJ); 20% tracing to a plant selected from Rumson CC (Rumson, NJ); 20% tracing to a
plant selected from the Piping Rock Club (Locust Valley, NY); 10% tracing to a plant
selected from Spring Lake GC (Spring Lake Heights, NJ); and 40% tracing to L93-10 (a
dollar spot resistant clone selected from L93). (Stacy Bonos – personal communication)
3
13M A. stolonifera
Maternal progenies of 21 clones collected from old golf course putting greens and
fairways - 32% of the germplasm tracing to several plants collected from putting greens
and fairways from golf courses in Arizona; 19% of the germplasm tracing to a plant
selected from or related to ‘L93’; 19% of the germplasm tracing to a plant selected from
the Piping Rock Club in Locust Valley, NY; 10% tracing to a plant selected from Lake
Success Golf Club in Great Neck, NY; 10% tracing to a plant selected from Hollywood
Golf Club in Deal, NJ; and another 10% tracing to a plant selected from Barrington Hills
CC in Barrington, IL. The creeping bentgrass plants selected from old golf courses were
of unknown origin, however, it is likely that these plants originated from South German
mixed bentgrass as this was one of the few commercially available sources for bentgrass
seed in the early 1900’s. (Rutgers, the State University of New Jersey 2009a)
SRP1WM A. stolonifera
Traces to clones selected from long-term research plots at Michigan State University
(Leah Brilman – personal communication)
4
Barracuda A. stolonifera
Maternal progenies of 9 clones collected from old golf course putting greens and
fairways – 44.4% of the germplasm tracing to a plant collected from Spring Lake Golf
Club in Spring Lake Heights, NJ; 33.3% of the germplasm tracing to several plants
selected from the Piping Rock Club in Locust Valley, NY; 11.1% tracing to a plant
selected from Lake Success Golf Club in Great Neck, NY; and 11.1% tracing to a plant
collected from a golf course in Portugal. The creeping bentgrass plants selected from old
golf courses were of unknown origin, however, it is likely that these plants originated
from South German mixed bentgrass as this was one of the few commercially available
sources for bentgrass seed in the early 1900’s. (Peak Plant Genetics LLC and Rutgers, the
State University of New Jersey 2013)
Independence A. stolonifera
Maternal progenies of 14 clones collected from old golf course putting greens and
fairways – Attractive plants were collected from Harkers Hollow Golf and CC (Harmony,
NJ), Old Westbury Golf and CC (Old Westbury, NY) Suburban Golf Club (Union, NJ),
Canoe Brook Country Club (Summit, NJ), Shackamaxon Country Club (Scotch Plains,
NJ), the Piping Rock Club (Locust Valley, NY), and Lake Success Golf Club (Great
5
Neck, NY). The creeping bentgrass plants selected from old golf courses were of
unknown origin, however, it is likely that these plants originated from South German
mixed bentgrass as this was one of the few commercially available sources for bentgrass
seed in the early 1900’s. Additionally, approximately 7% of the parental germplasm
traces to plants selected from or related to ‘L93’. (Lebanon Seaboard Corporation and
Rutgers, the State University of New Jersey 2005)
King Pin A. stolonifera
Maternal progenies of 18 clones collected from old golf course putting greens and
fairways – 89% of the maternal germplasm traces to a plant designated ‘AZ-8’, selected
from an Arizona golf course. 11% of the maternal germplasm traces to a plant designated
‘AZ-144’, selected from an Arizona golf course. The creeping bentgrass plants selected
from old golf courses were of unknown origin, however, it is likely that these plants
originated from South German mixed bentgrass as this was one of the few commercially
available sources for bentgrass seed in the early 1900’s. (Stacy Bonos – personal
communication)
6
Luminary A. stolonifera
Maternal progenies of 6 clones with improved dollar spot resistance – 50% of the
germplasm tracing to collections made in 1998 from Lake Waramaug Country Club in
New Preston, CT; 33% of the germplasm tracing to outstanding clones from an L-93
research trial border planted at Rutgers Horticultural Research Farm II, in New
Brunswick, NJ in 1996; 17% of the germplasm tracing to collections made in 1996 from
the Piping Rock Club in Locust Valley, NY (Stacy Bonos – personal communication).
Focus A. stolonifera
Maternal progenies of 9 clones collected from old golf course putting greens and
fairways – 56% of the germplasm tracing to a plant collected from the Piping Rock Club
in Locust Valley, NY; 11% tracing to a plant designated ‘AZ-144’ from an Arizona golf
course; and 33% tracing to a segregating clone of a Penncross seeded research trial
border area in North Brunswick, NJ. The creeping bentgrass plants selected from old
golf courses were of unknown origin, however, it is likely that these plants originated
from South German mixed bentgrass as this was one of the few commercially available
sources for bentgrass seed in the early 1900’s. (Stacy Bonos – personal communication)
7
Alpha A. stolonifera
The origin of ‘Alpha’ traces to 15 lines: 3 of the 15 lines traced to a Massachusetts golf
course; 3 lines traced to a North Carolina golf green; and the remaining lines traced to
selections of low growing surviving plants from a herbicide damaged Foundation field of
‘Putter’ creeping bentgrass in the Willamette Valley of Oregon. Selections of segregated
patches from old putting greens were 30 to 100 years old and were likely sown to South
German Mixed Bentgrass or ‘Penncross’. (Brede 2007)
Authority A. stolonifera
Traces to 2 plants designated C952 and C953 selected from an experimental turf trial
established in 1992 at the Pinehurst Resort (Pinehurst, NC) (Novel Ag Inc 2009)
Shark A. stolonifera
Fifty percent of ‘Shark’ traces to 2 plants designated C952 and C953 selected from an
experimental turf trial established in 1992 at the Pinehurst Resort (Pinehurst, NC). An
additional 50% of Shark traces to clones collected from putting greens of old golf courses
– golf course collection sites include Gulph Mills Golf Club (Conshohocken, PA), Brae
Burn Country Club (West Newton, MA), Richmond County Country Club (Staten Island,
8
NY), and Green Brook Country Club (North Caldwell, NJ). The creeping bentgrass
plants selected from old golf courses were of unknown origin, however, it is likely that
these plants originated from South German mixed bentgrass as this was one of the few
commercially available sources for bentgrass seed in the early 1900’s. (Novel Ag Inc and
Rutgers, the State University of New Jersey 2009)
V8 A. stolonifera
V8 traces to plants from a collection that were tested in a turf trial at the Pinehurst Resort
in Pinehurst, NC in 1992. Three selected clones (PH-N, PH-S, PH-TC) were turf tested at
Rutgers NJAES in North Brunswick, NJ, along with twelve attractive progeny from
them, labeled R1-R12. Remnant seed saved from the PH-N and R2 plants were used to
establish a spaced plant breeder block of V8 in June 2010 near Connell, WA. (Tom
Brentano – personal communication)
Benchmark DSR A. stolonifera
Maternal progenies of 13 clones collected from old golf course putting greens and
fairways – 76% of the maternal germplasm traces to plants designated AZ-8 and AZ144,
selected from an Arizona golf course; 14% of the maternal germplasm traces to a plant
9
designated 7350-6, selected from the Piping Rock Club in Locust Valley, NY; and 8% of
the germplasm traces to a plant collected from a golf course fairway at Barrington Hills
CC in Barrington, IL. The creeping bentgrass plants selected from old golf courses were
of unknown origin, however, it is likely that these plants originated from South German
mixed bentgrass as this was one of the few commercially available sources for bentgrass
seed in the early 1900’s. (Rutgers, the State University of New Jersey 2009b).
Memorial A. stolonifera
Maternal progenies of 25 clones from old golf course putting greens and fairways – 27%
of the germplasm traces to plants selected from the Piping Rock Club in Locust Valley,
NY; 14% tracing to plants selected from a fairway on Lake Success Golf Club in Great
Neck, NY; 13% tracing to a plant designated AZ-8 from an Arizona golf course; 13%
tracing to a plant designated AZ-64 from an Arizona golf course; 13% tracing to a plant
designated AZ-144 from an Arizona golf course; 13% tracing to a plant selected from or
related to L-93; 7% tracing to a plant designated 7356-5 collected from Hollywood Golf
Club in Deal, NJ. The creeping bentgrass plants selected from old golf courses were of
unknown origin, however, it is likely that these plants originated from South German
10
mixed bentgrass as this was one of the few commercially available sources for bentgrass
seed in the early 1900’s. (Stacy Bonos – personal communication)
Century A. stolonifera
Polycross of 6 clones: clone TAES 2831 (collected from green #9 Brookhollow CC,
Dallas, TX in 1984); clone TAES 3153 (collected from Rome Acquasanta GC Italy in
1986); clone TAES 3250 (from Nimes Campagne GC France 1986); clone TAES 3307
(also identified as U6 from Firenze GC France 1986); clone TAES 3794 (individual plant
selected from Syn4 (Cato)); and TAES 3799 (individual plant selected from Syn4
(Cato)). (Texas Agricultural Experiment Station 2004a).
Imperial A. stolonifera
Seven clone polycross synthetic: clone TAES 2833 (collected from green #9
Brookhollow CC, Dallas, TX in 1984); clone TAES 2845 (collected from green #9
Brookhollow CC, Dallas, TX in 1984); clone TAES 2916 (collected from green #14
Walnut CC, East Lansing, MI in 1984); clone TAES 2922 (collected from green #5
Platteville CC, Platteville, WI in 1984); clone TAES 3106 (also identified as AGBP-1/#4
from Kevin McVeigh, Oregon in 1986); TAES 3293 (from Nimes Campagne GC France
11
1986); and TAES 3307 (also identified as U6 from Firenze GC France 1986). (Texas
Agricultural Experiment Station 2004b).
Unknown A. stolonifera
Unknown contaminate creeping bentgrass plants in ‘Greenwich’ velvet bentgrass seed lot
(M50-9-70351)
Southshore A. stolonifera
Broad based cultivar derived from the progenies of 203 selected clones. Clones were
collected from old golf course putting greens primarily in New Jersey, New York,
California, Arizona, and Pennsylvania. (Lofts Seed Inc 1994)
L93 A. stolonifera
Progeny of 19 clones. Plant material originally traces to 15 maternal clones of the 203
clone ‘Southshore’ nursery (see above). The 15 clones were from plants collected in
New Jersey, New York, and Arizona. Seed harvested from 15 clones was subjected to
additional cycles of selection: approximately 1000 seedlings from original 15 clones
panted in a nursery at Rutgers Plant Science Research Farm, Adelphia, NJ in 1987; 34 of
the most promising plants subsequently vegetatively established at Lofts Research Farm,
12
Martinsville, NJ in 1989; vegetative propagules of selected clones established in an
isolated polycross nursery at Lofts Great Western Research Farm in Lebanon, OR in
1990, with progeny subsequently planted at Martinsville NJ in 1991. L93 breeders seed
nursery established using 12 clones from the 1989 Martinsville NJ trial, 1 clone from a
‘Southshore’ production field in Oregon, and six of the progenies of the 1991
Martinsville NJ trial. (Richard Hurley – personal communication)
Penncross A. stolonifera
First generation synthetic produced by random crossing of three vegetatively propagated
clonal strains. Developed by H.B. Musser in 1954 (Hein, 1958). As one of the first
improved seeded varieties of creeping bentgrass, Penncross was used to seed or overseed
many golf courses throughout North America (Musser 1959; Bonos and Huff 2013).
Penneagle II A. stolonifera
Population improvement project from ‘Penneagle’. Plants that produced breeder seed
trace to the following sources: 53% to PST-OPN, of which 44% trace maternally to
Penneagle; 23.5% to ‘Pennlinks’ pollinated by Penneagle; 8.5% to Penneagle; 8.5% to
13
Pennlinks; 3.5% to PennA1 pollinated by Penneagle; and 3% to ‘Seaside II’ pollinated by
Penneagle. (Pure Seed Testing Inc 2006)
SR 1119 A. stolonifera
Five clone synthetic. Clones trace back to breeder block of ‘Providence’ (SR 1019)
creeping bentgrass. (Seed Research of Oregon Inc 2004)
Pennlinks A. stolonifera
Original parental clone selected from a putting green at The Country Club, Shaker
Heights, Ohio, in 1964, with The Country Club putting greens originally established to
South German Mixed Bentgrass in 1930. Original clone outcrossed with 9 Penn State
experimental bentgrass selections in 1966. Progeny subjected to two cycles of selection
and then backcrossed to original parental clone. Backcrossed progeny subjected to
additional selection, followed by first generation breeder seed of Pennlinks produced
utilizing approximately 100 selected parental clones. (Pennsylvania Agricultural
Experiment Station 1987)
14
PennA4 A. stolonifera
Polycross procedure where single parent selection A4 was crossed with experimental
selections of Penn G-series followed by two generations of crossing selected sib plants of
PennA4. Penn A-series collected from four putting greens of the main course at Augusta
National Golf Club where putting greens were originally seeded to Penncross creeping
bentgrass. Penn G-series collected from two putting greens of the Par 3 course at
Augusta National Golf Club where putting greens were originally seeded to Penneagle
creeping bentgrass. (Pennsylvania Agricultural Experiment Station 2004)
Proclamation A. stolonifera
Maternal progenies of 12 different clones with improved dollar spot (Sclerotinia
homoeocarpa F.T. Bennet.) resistance. Sixty-five percent of the germplasm used in the
development of Proclamation creeping bentgrass traces to collections made in 1996 from
the Piping Rock Club in Locust Valley, NY. Seventeen percent trace to plants collected
from several golf courses in Northern Italy. Ten percent trace to plants collected from
several golf courses in Eastern France. Eight percent trace to a plant collected from
Harker’s Hollow Golf course in Harmony, NJ in 1996. (Stacy Bonos – personal
communication)
15
T1 A. stolonifera
The origin of ‘T1’ traces to 11 clonal lines: one clone traced to a selection from the
cultivar ‘Providence’; one clone traced to a plant selected from a South Carolina putting
green; one clone traced to a selection from ‘Putter’; 3 clones traced to two Massachusetts
putting greens; and the remaining clones traced to Oregon polycross 94-8283 (originating
from a Massachusetts putting green) and ‘Putter’. Selections of segregated patches from
old putting greens were 30 to 100 years old and were likely sown to South German
Mixed Bentgrass or ‘Penncross’. (Brede 2007)
Carmen A. stolonifera
European cultivar
Penneagle A. stolonifera
Four clone synthetic with the following origin of parental clones: 1) 3rd generation
seedling selection from ‘Washington’ bent (1938 source); 2) 3rd generation seedling
selection from ‘Seaside’ (1930 source); 3) 3rd generation seedling selection from Cocoos
16
bent (1932 source); 3rd generation seedling selection from ‘Washington’ bent (1933
source). (Pennsylvania Agricultural Experiment Station 1980)
Crenshaw A. stolonifera
Six clone synthetic: 5 clones obtained from the University of Arizona and 1 clone
collected from Brookhollow CC (Dallas, TX). (Texas Agricultural Experiment Station
1996).
Providence A. stolonifera
5 clone synthetic: 1) clone P19 – 1965 selection of a creeping bentgrass plant from plot of
‘Piper’ velvet bentgrass in a putting green trial at the University of Rhode Island; 2) clone
P-7 - 1965 selection of a creeping bentgrass plant from plot of ‘Arlington’ creeping
bentgrass in a putting green trial at the University of Rhode Island; 3) clone P-27 - 1965
selection of a creeping bentgrass plant from a plot of ‘Highland’ bentgrass; 4) clone P-43
- 1976 selection from polycross progeny from University of Rhode Island collections and
selections; and 5) clone P-87 – 1972 selection from old putting green. (University of
Rhode Island 1991)
17
Putter A. stolonifera
Hybrid of two lines: AP-10 and pollen parent BPA-163. AP-10 is an improved line
developed through consecutive cycles of intercrossing 26 bentgrass clones collected from
around the US Northwest and the United States. BPA-163 was a single plant selection
from the Puyallup Research and Extension Center (Washington State University) that
exhibited high seed yield potential. (Brauen et al. 1993; Jacklin Seed Company 1991)
Emerald A. stolonifera
Single clone selection from ‘Congressional’ bentgrass (Alderson and Sharp 1994)
PI 531251 Hungary A. stolonifera
‘Keszthelyi 4’ – cultivar from Hungary
Seaside A. stolonifera
Landrace originally harvested for seed from naturalized stands in the Coos Bay, Umpqua,
and Siuslaw regions of Oregon (Schoth 1930; Warnke 2003).
PI 235440 Switzerland A. stolonifera
Collection from near Schuls, Switzerland
PI 494118 Ukraine A. stolonifera
‘Klonobaja’ – cultivar from Ukraine
18
PI 302902 Spain A. stolonifera
Collection from Spain
PI 318934 Spain A. stolonifera
Collection from Zaragoza, Spain
PI 608809 US A. stolonifera
Maternal progeny of MSCB-3 from Mississippi Agriculture and Forestry Experiment
Station, polycrossed with MSCB-4, MSCB-10, MSCB-11, MSCB-19, MSCB-37,
MSCB-52, MSCB-73, MSCB-81, MSCB-304, MSCB-330, and MSCB-388; where
MSCB-3 and MSCB-4 were derived from tissue culture regenerated callus of Penncross
on culture media with high salt concentration; MSCB-10, MSCB-19, MSCB-37, MSCB52, MSCB-81, MSCB-304, and MSCB-388 were developed from tissue culture
regenerated callus of Penncross exposed to high temperature; and MSCB-11, MSCB-73,
and MSCB-304 were ecotypic selections from Mississippi. (Krans et al. 2000).
PI 608810 US A. stolonifera
Maternal progeny of MSCB-4 from Mississippi Agriculture and Forestry Experiment
Station, polycrossed with MSCB-3, MSCB-10, MSCB-11, MSCB-19, MSCB-37,
19
MSCB-52, MSCB-73, MSCB-81, MSCB-304, MSCB-330, and MSCB-388; where
MSCB-3 and MSCB-4 were derived from tissue culture regenerated callus of Penncross
on culture media with high salt concentration; MSCB-10, MSCB-19, MSCB-37, MSCB52, MSCB-81, MSCB-304, and MSCB-388 were developed from tissue culture
regenerated callus of Penncross exposed to high temperature; and MSCB-11, MSCB-73,
and MSCB-304 were ecotypic selections from Mississippi. (Krans et al. 2000).
Declaration A. stolonifera
Maternal progenies of 8 clones with improved dollar spot (Sclerotinia homoeocarpa F. T.
Bennet.) resistance. 72% of the maternal germplasm traces to several plants selected
from the Piping Rock Club in Locust Valley, NY in 1996. 14% of the maternal
germplasm traces to a putting green collection from Spring Lake Golf Club in Spring
Lake Heights, NJ in 1996, and another 14% traces to a plant selected from Harkers
Hollow Golf and CC in Harmony, NJ in 1996. The creeping bentgrass plants selected
from old golf courses were of unknown origin, however, it is likely that these plants
originated from South German mixed bentgrass as this was one of the few commercially
20
available sources for bentgrass seed in the early 1900’s. (Lebanon Seaboard Corporation
and Rutgers, the State University of New Jersey 2009a).
National A. stolonifera
Originated from plant selections from old turf areas made by A. C. Ferguson in 1963,
1967, and 1973 near Winnepeg, Manitoba. (Pickseed West Inc. 1991)
PI 221906 Afghanistan A. stolonifera
Collection from Afghanistan
PI 608812 US A. stolonifera
Maternal progeny of MSCB-11 from Mississippi Agriculture and Forestry Experiment
Station, polycrossed with MSCB-3, MSCB-4, MSCB-10, MSCB-19, MSCB-37, MSCB52, MSCB-73, MSCB-81, MSCB-304, MSCB-330, and MSCB-388; where MSCB-3 and
MSCB-4 were derived from tissue culture regenerated callus of Penncross on culture
media with high salt concentration; MSCB-10, MSCB-19, MSCB-37, MSCB-52, MSCB81, MSCB-304, and MSCB-388 were developed from tissue culture regenerated callus of
Penncross exposed to high temperature; and MSCB-11, MSCB-73, and MSCB-304 were
ecotypic selections from Mississippi. (Krans et al. 2000).
21
PI 608811 US A. stolonifera
Maternal progeny of MSCB-10 from Mississippi Agriculture and Forestry Experiment
Station, polycrossed with MSCB-3, MSCB-4, MSCB-11, MSCB-19, MSCB-37, MSCB52, MSCB-73, MSCB-81, MSCB-304, MSCB-330, and MSCB-388; where MSCB-3 and
MSCB-4 were derived from tissue culture regenerated callus of Penncross on culture
media with high salt concentration; MSCB-10, MSCB-19, MSCB-37, MSCB-52, MSCB81, MSCB-304, and MSCB-388 were developed from tissue culture regenerated callus of
Penncross exposed to high temperature; and MSCB-11, MSCB-73, and MSCB-304 were
ecotypic selections from Mississippi. (Krans et al. 2000).
PI 269838 Germany A. stolonifera
Collected material from Germany
PI 659907 Kyrgyzstan A. stolonifera
Collection from Ysyk-Kol, Kyrgyzstan
PI 204390 Turkey A. stolonifera
Collection from near Kayseri, Turkey
22
PI 423036 Spain A. stolonifera
Collection from Rio Guadarrama at Route 6, west of Madrid
PI 235541 Sweden A. stolonifera
Collection from natural pasture near Landskrona, Sweden
PI 283174 A. transcaspica
Collection from former Soviet Union
PI 255869 A. gigantea
Collected material from Poznan, Poland
PI 619558 A. gigantea
Collection from Xinjiang, China
PI 439027 Soviet Union A. stoloniferaa
Collection from former Soviet Union
PI 608813 US A. stoloniferaa
Maternal progeny of MSCB-19 from Mississippi Agriculture and Forestry Experiment
Station, polycrossed with MSCB-3, MSCB-4, MSCB-10, MSCB-11, MSCB-37, MSCB52, MSCB-73, MSCB-81, MSCB-304, MSCB-330, and MSCB-388; where MSCB-3 and
MSCB-4 were derived from tissue culture regenerated callus of Penncross on culture
23
media with high salt concentration; MSCB-10, MSCB-19, MSCB-37, MSCB-52, MSCB81, MSCB-304, and MSCB-388 were developed from tissue culture regenerated callus of
Penncross exposed to high temperature; and MSCB-11, MSCB-73, and MSCB-304 were
ecotypic selections from Mississippi. (Krans et al. 2000).
PI 222073 Afghanistan A. stoloniferaa
Collection from east base of Sibar Pass, Kabul, Afghanistan
PI 230235 Iran A. stoloniferaa
Collection from Dastana, Iran
W6 21826 Ukraine A. stoloniferaa
Collection from Krym, Ukraine
Alister A. capillaris
Origin traces to 10 colonial bentgrass plants collected from bluegrass roughs at the Piping
Rock Club, Locust Valley, NY. (Pure Seed Testing Inc 2004)
FT12 A. capillaris
FT12 colonial bentgrass was selected from the maternal progenies of 11 clones – 32%
tracing to a few plants collected from Hebo Cemetery in Hebo, Oregon in 1996; 32%
24
tracing to East border of creeping bentgrass research trial at Horticultural Research Farm
II, in New Brunswick, NJ in 1996. 23% tracing to a plant collected from a mountain
meadow in Eastern Europe in 1997; and 13% tracing to a few plants collected from Brae
Burn Country Club in West Newton, MA in 1997 (Stacy Bonos – personal
communication).
Capri A. capillaris
Maternal progenies of 24 clones – 64% tracing to several plants selected from the
Connecticut Agricultural Experiment Station; 26% of the germplasm tracing to
segregating plants from the border area of a putting green research trial in North
Brunswick, NJ; and 10% tracing to plants collected from Hebo Cemetery in Hebo,
Oregon. (Stacy Bonos – personal communication)
EBM A. capillaris
EBM colonial bentgrass was selected from the maternal progenies of 51 clones – 42%
tracing to a few plants received from the Connecticut Experiment Station in 1996; 34%
tracing to a few plants collected from Hebo Cemetery in Hebo, Oregon in 1996; 13%
tracing to a plant collected from Century CC in Purchase, NY in 1996; and 11% tracing
25
to several plants collected from mountain meadows and old golf courses in Eastern
Europe in 1997 (Stacy Bonos – personal communication).
SR7100 A. capillaris
Origin traces to 90 plants from the European colonial bentgrass ‘Bardot’ selected for
improved seed yield. (Seed Research of Oregon Inc 1994)
SR7150 A. capillaris
Progeny of four selected plants tracing to plants selected from a PI collection planted at
USDA Farm, Beltsville, MD. (Leah Brilman – personal communication)
Tiger II A. capillaris
Traces maternally to: 33.3% from the cultivar Bardot, and 66.7% from the cultivar Denso
(Steve Reid – personal communication)
CDE composite A. canina
CDE velvet bentgrass was selected from the maternal progenies of 29 clones. Sixtyseven percent of the maternal sources trace to several plants collected from Lake Success
Golf Club in Great Neck, NY in 1996. Thirty-three percent of the maternal sources trace
26
to plants selected out of the 1985 velvet bentgrass test located at Hort Farm II, acquired
by Jerry Pepin (Stacy Bonos – personal communication).
SMM composite A. canina
SMM velvet bentgrass was selected from the maternal progenies of 25 clones. Sixty-four
percent of the maternal sources trace to several plants collected from Lake Success Golf
Club in Great Neck, NY in 1996. Thirty-six percent of the maternal sources trace to
plants selected out of the 1985 velvet bentgrass test located at Hort Farm II, acquired by
Jerry Pepin (Stacy Bonos – personal communication).
SME composite A. canina
SME velvet bentgrass was selected from the maternal progenies of 21 clones. Seventyone percent of the maternal sources trace to several plants collected from Lake Success
Golf Club in Great Neck, NY in 1996. Twenty-four percent of the maternal sources
traces to plants selected out of the 1985 velvet bentgrass test located at Hort Farm II,
acquired by Jerry Pepin. Five percent trace to plants collected from Lake Sunapee CC in
New London, NH in 2004 (Stacy Bonos – personal communication).
27
VTP composite A. canina
VTP velvet bentgrass was selected from the maternal progenies of 18 clones. Seventyeight percent trace to plants collected from Lake Sunapee CC in New London, NH in
2004. Eleven percent trace to plants collected from Winchester CC in Winchester, MA in
2004. Another 11% trace to plants collected from Pound Ridge Golf Club in Pound
Ridge, NY in 2004 (Stacy Bonos – personal communication).
Legendary A. canina
Selected from the maternal progenies of nine clones. 20% of the germplasm traces to
selections from ‘Emanuel Francis’ velvet bentgrass acquired from Dr. Jerry Pepin; 50%
of the maternal sources trace to several plants collected from Lake Success Golf Club
(Great Neck, NY); 20% traces to plants collected from the National Golf Links of
America in Southhampton, NY; and 10% traces to a plant collected from Rock Ridge CC
in Newtown, CT. (Lebanon Seaboard Corporation & Rutgers, The State University of
New Jersey 2009b)
28
Villa A. canina
Material traces to germplasm acquired from Dr. Jerry Pepin, plants collected from Lake
Success Golf Club (Great Neck, NY), and plants collected from the National Golf Links
of America in Southhampton, NY. (DLF International Seeds 2012)
SR 7200 A. canina
Traces back to 4 clones collected from New England area putting greens by Dr. Richard
Skogley (University of Rhode Island). (Seed Research of Oregon Inc 1998)
PI 203444 Apera interrupta
Collection from Turkey
PI 204389 Apera interrupta
Collection from Turkey
PI 372227 Polypogon chilensis
Collection from Uruguay
PI 204395 Polypogon viridis
Collection from Malatya, Turkey
a
indicates putative Agrostis gigantea PI accessions originally labeled as Agrostis stolonifera by the USDA NPGS.
29
References
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Josh A. Honig*, Christine Kubik, Vincenzo Averello, Jennifer Vaiciunas, William A. Meyer, and Stacy A. Bonos. *Dep. of Plant Biology and Pathology, School
of Environmental and Biological Sciences, Rutgers, The State University of New Jersey, 59 Dudley Rd., New Brunswick, NJ 08901-8520,
honig@aesop.rutgers.edu