Natural and Artificial Selection Biology H Lamarck’s Theory of Evolution Lamarck hypothesized that species evolve through use and disuse and the inheritance of acquired traits The mechanisms he proposed are unsupported by evidence The Voyage of the Beagle During his travels on the Beagle, Darwin collected specimens of South American plants and animals He observed adaptations of plants and animals that inhabited many diverse environments His interest in geographic distribution of species was kindled by a stop at the Galápagos Islands near the equator west of South America Darwin made two major points in his book: Many current species are descendants of ancestral species Natural selection is a mechanism for this evolutionary process Darwin developed two main ideas: Evolution explains life’s unity and diversity Natural selection is a cause of adaptive evolution Descent with Modification The phrase descent with modification summarized Darwin’s perception of the unity of life The phrase refers to the view that all organisms are related through descent from an ancestor that lived in the remote past In the Darwinian view, the history of life is like a tree with branches representing life’s diversity Observation #1: For any species, population sizes would increase exponentially if all individuals that are born reproduced successfully Observation #2: Populations tend to be stable in size, except for seasonal fluctuations Observation #3: Resources are limited Inference #1: Production of more individuals than the environment can support leads to a struggle for existence among individuals of a population, or competition, with only a fraction of their offspring surviving Observation #4: Members of a population vary extensively in their characteristics; no two individuals are exactly alike Observation #5: Much of this variation is inherited Inference #2: Survival depends in part on inherited traits; individuals whose inherited traits give them a high probability of surviving and reproducing are likely to leave more offspring than other individuals “Fitness” in evolutionary terms is the individual who produces the most offspring that live to reproduce Inference #3: This unequal ability of individuals to survive and reproduce will lead to a gradual change in a population, with favorable characteristics accumulating over generations Natural selection The major microevolutionary process that results in differential survival and reproduction (works on phenotypes) There is variation among individuals More are born than can survive There is competition for resources Those individuals that are most fit for their environment survive, reproduce and pass on their alleles. Natural selection produces an increase over time in adaptation of organisms to their environment If an environment changes over time, natural selection may result in adaptation to these new conditions Organisms cannot “choose” to adapt to changes in their environment; the variation must already be in the gene pool. Those with the variation that helps them, survive and reproduce. Those that don’t have the variation, die. Selection of gene frequencies Directional selection: shifts allele frequencies in a certain direction in response to environmental pressures; moths Stabilizing selection: favors the most common forms of a trait in a population; human babies Disruptive selection: favors forms at both ends of the range of variation and selects against the intermediate forms; finches Peppered moths Artificial Selection In artificial selection, humans have modified other species over many generations by selecting and breeding individuals with desired traits Dogs Farm animals Corn