Continuity and Contrast: parameters in neutral point pattern landscapes. Spectral density has been used frequently for time series and lattice data (see Mugglestone & Renshaw 1996 for relevant references). Here we present how to, by a spectral representation, calculate two measures which capture essential characteristics in point pattern landscapes. The basis of the spectral method follows from the Fourier theorem, where it is stated that continuous time series, or surfaces and higher dimensions, can be represented by a combination of sine and cosine terms with different frequencies and amplitudes. Thereby, analysis of the time series, or surfaces, can be performed on the sine and cosine terms. From physics we use the notation Power Spectral Density Function, PSDF, for this representation of a time series or surface by the amplitudes of sine and cosine terms of different frequency. Continuity Large values of Continuity (γ) in point pattern reflects that nearby areas have similar density and hence πΎ is a measure of autocorrelation. It is defined from the spectral representation of the landscape such that if the spectral density is dominated by the amplitudes of low frequencies there is a higher degree of autocorrelation. The measure is a point pattern measure, akin to a two dimensional measure of 1/f noise (Halley et al. 2004, Keitt 2000). A higher value on Continuity is then analogous to a redder 1/f noise. We analyze, as is done when determining the color of 1/f noise, the autocorrelation as a linear regression fitted to the log(frequency) vs. log(amplitude) and thereby also infer self similarity. The slope of the regression line is the parameter Continuity, πΎ. In our analysis we have assumed isotropy which implies identical patterns in all directions. Hence, we may collapse the spectral analysis into an analysis in one dimension, a regression line instead of a surface, after the Fourier transform of the two dimensional point pattern. To calculate the Power Spectral Density Function of the point pattern distribution, denoted PSDFpp, we use the work of Mugglestone & Renshaw (1996) where they present an efficient way to calculate PSDFpp as 2 2 πππ·πΉππ = (∑ πππ (πΆππ π₯π )) + (∑ π ππ(πΆππ π¦π )) π (A.1) π Here, (μp,μq) are frequencies for Fourier transform of grid data, (xi,yi) are coordinates in the point pattern, and C is a constant that determines the number of frequencies. Mugglestone & Renshaw (1996) state that no more frequencies should be used than there are points in the point pattern, Np, to keep them independent. Therefore we chose C to be and the number of frequencies used is nF =2C×2C. Contrast N p 2 rounded down to nearest integer As a point pattern equivalent of coefficient of variation we introduce Contrast, πΏ, to measure difference of density between sparse and dense areas. To formulate an equation for that measure, we first have to relate it to how variance in the continuous case can be measured using spectral representation. Note that when time series and surfaces are represented by sine and cosine terns after the Fourier transform it is the amplitudes of the frequencies that determines the variance. If M is the number of frequencies in the PSDF the variance, π2, is calculated as π2 = 1 ππππ(πππ·πΉ − πππ·πΉ(ππππππ)) π (A.2) The mean is represented by the amplitude in the origin1 of the frequency plane, PSDF(origin), and hence the coefficient of variation, standard deviation divided by the mean, is πΆπ = π 1 √ ππππ(πππ·πΉ − πππ·πΉ(ππππππ)) πππ·πΉ(ππππππ) π (A.3) Substituting PSDF with PSDFpp from equation A.1 we end up with an equation of the point pattern measure Contrast, πΏ: πΏ = πΆπππ = ππΉ 1 √ ππππ(πππ·πΉππ − πππ·πΉππ (ππππππ)) πππ·πΉππ (ππππππ) π (A.4) Generating neutral point pattern landscapes To get NPPL with given characteristics we generated lattice landscapes of size m×m. The density defines the probability of a point in the landscapes. We first generated 2-dimensional 1/f-noise (denoted LG) using a method similar to that presented by Halley et al. (2004). Hence this is still a representation of a lattice landscape not a point pattern. The values in LG are normally distributed and hence not suitable for describing probabilities because it may include negative values. While this could be solved by truncating we found that it would not allow for generating sufficiently high values of δ. We therefore transformed LG using spectral mimicry. This method was introduced by Cohen et al. (1999) and has been used to construct stationary time series with different Fourier spectra. Cohen et al. presents the method for transformation to normally distributed values with specific mean and variance. We instead transformed LG to LΓ using a 1 Note that if implemented in Matlab using built in functions of Fourier transform, for example the fft function, the amplitude in the origin, PSDF(origin), represents the sum instead of the mean. gamma distribution (which contains no values <0) with mean=1/m2 and coefficient of variation δL. Point locations were distributed according to the probabilities given by LΓ. While γ and δ of the spectral point pattern is determined by γL and δL, they are altered by both the gamma transformation of the grid values and the distribution of points. Hence we measured these quantities in the generated landscape (see method given above). The relationship between spectral point pattern values of γ and δ used in the study and the γL and δL required to generate them was found iteratively. Furthermore, we found that the linear relationship in the power spectra was maintained better for large grids (values of m) and hence we used m=2000. The autocorrelation parameter Continuity generates a general aggregation pattern while the variance within the system is reflected by the Contrast parameter. High Contrast parameter will impose more isolated clusters of aggregated points onto the aggregation structure defined by the Continuity parameter.