Unit 01: Life Science - Sustainability of Ecosystems What are the Factors Affecting the Sustainability of an Ecosystem?: Introduction to Ecology Ecology: Ecology is the scientific study of the interactions of organisms and their environment. As a scientific study, ecology involves observations and experiments to test hypothetical explanations of ecological phenomena. Ecology has a long history as an observational or descriptive science, but is relatively young as an experimental science. Ecology is a multidisciplinary field of study involving all areas of biology as well as the physical sciences. Interactions of organisms and their environment refers to the way the organism affects the environment as well as how the environment affects the organism. An Ecosystem is a community of organisms and the physical environment in which it lives. When an ecologist studies the organisms living in a forest and includes a study of the physical factors that affect the organisms in the forest, then the ecologist is studying an ecosystem. Each of the following concepts describes the organisms within their environment and their interactions with other organisms in their environment. You may recognize some of these concepts from your earlier studies in science. Abiotic Factors Biotic Factors Symbiotic Relationships Trophic Structure Herbivore, Carnivore, Omnivore and Saprobes Abiotic Factors Abiotic factors are the nonliving factors which affect life in any ecosystem. Some abiotic factors are described below: Space - All organisms require enough space or territory to insure adequate resources to food, water, shelter, and mates. Temperature - Environmental temperature affects biological processes and the ability of most organisms to regulate their temperature. Few organisms have active metabolisms at temperatures below 0oC or above 45oC because enzymes function best within a short range of temperature and become denatured if the temperature is too high. Most organisms are ectotherms and can maintain their body temperature only within a few degrees of ambient temperature. Even endotherms function best in a temperature range for which they are adapted. Oxygen - Most living organisms require oxygen for cellular respiration, which is a process that releases energy from food. Terrestrial organisms obtain oxygen from the atmosphere as they breath (usually with lungs). Aquatic organisms generally have gills and extract oxygen which is dissolved in the water. Sunlight - Sunlight is the ultimate source of energy for all photosynthetic organisms which in turn provide the resources for other living things (in most ecosystems). Light also affects the development and behaviour of many organisms which are sensitive to photoperiod. Water - Water (humidity) is necessary for all life. The ability to find water, to maintain water balance, and to conserve water help determine the habitat range for each species. Inorganic and Organic Soil Nutrients - Inorganic soil nutrients include minerals such as phosphates, nitrates, potassium, magnesium and a host of other minerals derived from rocks. Organic nutrients include organic compounds in humus which promote the growth of bacteria, fungi, and a host of other organisms beneficial to the soil. The physical structure, water holding potential, pH, and nutrient level of soil limit the distribution of plants and in turn the animals that inhabit a terrestrial region. The composition of the substrate in a stream affects the chemistry of the water above and in turn affects plant and animal life living there. the type of substrate also influences what animals can attach to or burrow in (especially in the intertidal zone). Biotic Factors Biotic factors refer to the living environment and include all other organisms that interact with the individual both of the same species and all other species. Biotic factors also includes decomposing animals and plants (detritus), disease, predator/prey interactions, competition, and symbiotic relationships (symbiosis). Detritus refers to decomposing plant and animal materials including their dead bodies as well as their wastes. Bacteria and fungi living in the ecosystem help to break down the materials within the detritus and recycle these materials back to the plants. Disease is the result if infection by fungi, bacteria, virus, and other pathogens. Disease is an important biotic factor because disease tends to reduce the number of organisms within the community. Predator/prey interaction is another important biotic factor which helps to limit the size of populations within an ecosystem. A predator is an animal that kills and eats another animal for food. The prey is the hunted animal. An example is the lion and the zebra. When a lion kills a zebra for food, the lion helps to prevent the overpopulation of the zebra. If the number of zebra declines too much the lion will starve. There is a balance between the number of predator and prey in any ecosystem. Competition is a struggle for survival that occurs between two organisms either of the same or different species. Birds often compete for nesting space. Lions and hyenas compete for food. Competition tends to limit the size of the population keeping it in balance with the available resources. Symbiotic Relationships Symbiotic relationships are biotic relationships in which two different organisms live in close association with each other to the benefit of at least one. There are five types of symbiotic relationships including: mutualism, commensalism, parasitism, parisitoidism. and predation. Mutualism is the type of symbiosis resulting in mutual benefit to both of the organisms in the relationship. An example of this would be the relationship between the algae and fungus of lichens. The fungi penetrate the roots of the plants and make soil nitrogen available to the plant, receiving carbohydrates in return. This allows them to live in an environment in which neither could survive alone. An example is a polyp found in the deep water off the coast of Newfoundland; it attaches itself to the shell of a certain species of the hermit crab, and by budding, covers the entire shell with a colony that dissolves the original shell. Because the colony grows at the same rate as the crab, it furnishes continuous protection, and the crab does not shed its shell at periodic intervals as it normally would. The polyp in turn benefits by moving about with the crab, thus obtaining a greater food supply then it would obtain if attached to a stationary object. Commensalism is a relationship in which one organism benefits from the relationship but the other organism seems to neither be harmed nor benefited. One example to illustrate commensalism is the beaver and the fish. A beaver builds a dam to regulate water level which helps the beaver survive winter. Fish often inhabit the beaver pond. The fish benefit from the beaver, but the beaver is neither harmed nor gains benefit from the fish. Another example of commensalism is the relationship between trees and nesting birds. The tree provides a nesting space for the bird but the bird may neither harm nor benefit the tree. On the other hand, if the bird eats insects that normally cause harm to the tree or if the bird produces wastes that are absorbed by the tree, one might argue that the tree and the bird both benefit. In this case the example would be considered mutualism. It is often difficult to determine the complete nature of any relationship and as a result it is often difficult to distinguish between mutualism and commensalism. Parasitism is a symbiotic relationship in which one organism benefits and the other is harmed. The organism that benefits is called the parasite, the organism that is harmed is called the host. Some parasites only cause slight damage to their host, while others kill them. Therefore one organism is injured in satisfying the needs of the other. An example would be the tapeworm. they live in the digestive tracts of various organisms, while there they are provided with nutrient and an environment in which to grow and reproduce. However, the host is harmed by the presence of the tapeworm. Parisitoidism is similar to parasitism. One organism benefits but the other is eventually killed - a sort of slow death. An example of parisitoidism is when a female wasp stings a spider causing paralysis but not death. The wasp then lays a single egg on the spider. When the egg hatches into a larva, it slowly eats the body of the spider eventually killing it - but slowly. Predation is where the interaction is beneficial to one species and detrimental to the other. This is not always considered a symbiotic relationship, although it is quite similar to parasitism, except for the degree of harm to the host or prey. With predation, the prey is killed. An example of predation is when a lion kills a zebra and eats it as its source of food. Trophic Structure: Trophic structure refers to the feeding relationships within the ecosystem. These feeding relationships are generally divided into five trophic levels based on their source of nutrition and include primary producers, primary consumers, secondary consumers, tertiary consumers, and decomposers (also known as detritivores). The various organisms that comprise each of the trophic levels determines the flow of energy and the cycling of materials within the ecosystem. Feeding relationships are generally viewed as a food web consisting of all the possible food chains that exist within the ecosystem. Since the primary producers are the only photosynthetic organisms, they determine the amount of energy (trapped from the sun) that can be passed up through the food web. The detritivores enable the recycling of materials and are therefore the organisms that are the foundation for biogeochemical cycling. Producers or autotrophs are organisms, such as green plants, that produce their own food. They make organic compounds (food such as sugar) from inorganic compounds (carbon dioxide and water) by photosynthesis. The ultimate source of energy for life on Earth is the sun. Solar energy is trapped during the process of photosynthesis and converted into a chemical form that we normally call food. Food contains both materials (the elements carbon, hydrogen, nitrogen, and the other essential elements) as well as stored energy. The materials within the food are recycled. They pass from the producers to the consumers and finally are recycled back to the producers by the action of the decomposers. Energy, unlike the materials, is not recycled. As the food is passed through the food web, most of the energy is lost. In general terms, about 10% of the energy stored in one trophic level (such as producers) is actually transferred to the next trophic level (for example the herbivores). This is known as the pyramid of energy. Eventually there is so little energy remaining in the top trophic level that no higher trophic level can be supported. This is why there are so few if any fourth order consumers in an ecosystem. Consumers or heterotrophs are organisms that obtain nutrients from other organisms. They cannot synthesize their own food so they must obtain it ready made. They eat other living things. There are two different ways to classify carnivores. The first group are known as predators. Predators hunt and kill other animals for food. Animals like lions, wolves, snakes, and sharks are all classified as predators. Another group of animals that are considered carnivores are scavengers. Scavengers are animals that feed on the dead bodies of other organisms. Animals such as vultures, hyenas, and griffins and all classified as scavengers. Scavengers eat the food that has been killed and left behind by predators. Scavengers are a very important group because they dispose of the carcass's of animals that have been left to decompose. Decomposers are organisms of decay. These are also called saprobes. They are generally fungi or bacteria that break down the complex compounds in the remains of dead animals and plants, producing simple substances that can be used again by the producers. Decomposers are therefore very important because they recycle materials within the ecosystem. The decomposers are the final consumers in any ecosystem. Herbivores, Carnivores, Omnivores, and Saprobes: There are several groups of heterotrophs. Heterotrophs are all organisms that obtain their food from the environment. All animals and certain types of micro-organisms are heterotrophs. These organisms must take in, or ingest, food containing already made organic nutrients from other plants or animals. It all depends on what they eat and how they obtain it. Heterotrophs include herbivores, carnivores, omnivores and saprobes. Herbivores are animals that feed only on plants. Rabbits, cattle, horses, sheep and deer are all herbivores. Carnivores are animals that feed on other animals. Some carnivores may be predators (such as lions, hawks, and wolves who attack and kill their prey and feed on their bodies) and some may be scavengers (they feed on dead animals that they find). Omnivores are animals that feed on both plants and animals. Examples of omnivores are humans and bears. Saprobes are organisms that get nutrients by breaking down the remains of dead plants and animals, or their wastes. Examples of saprobes are bacteria and fungi. Saprobes are also known as decomposers are an essential component of any ecosystem. Their main role is to recycle nutrients in dead organisms and their wastes. Without the decomposers to recycle nutrients, there could be no life since plants would run out of nutrients.