Biological Classification Core Objectives: Understand that there is great variety in life Classifying Species o Understand the importance of classification o Scientific Name / Binomial Nomenclature o 4 Main ways to classify a species from another o Why scientist sometimes change the classification of a species Hierarchical Classification o Memorize Level Order o Understand how levels can be used to infer similarity and shared ancestry o Disadvantages Phylogenic Trees & Cladograms o Trees of life show evolutionary history (branching points / speciation events / derived characters) o Dichotomous Keys Major Taxa o Archae o Bacteria o Eukarya Protista Fungi Plantae Animalia Domains Kingdoms Archae Bacteria Eukarya Protista Fungi Animalia Plants Prokaryotic vs. Eukaryotic Unicellular vs. Multicellular Type of Metabolism (How they get energy) Examples Major Characteristics Key Terms Biodiversity 13 Billion Species estimated 5% of actually named Tropics have most Many of unnamed are microscopic New organisms found every day Evolution still happening Species Ecological / Biological Test (Interbreeding) Morphological Test Niche Test Molecular Test Early Taxonomy Aristotle Classification by complexity and role/habitat John Ray Latin names Linnaeus Binomial Nomenclature Genus Species Scientific Names Rules Genus first, then species Genus is capitalized Species is lowercase Latin or sometimes Greek Always italicized or underlined If anything greater than genus is specified, it is capitalized, but not italicized Acceptable Structure Genus species G. species Genus species G. species Examples International Code for Binomial Nomenclature International Naming Congress Reasons to Use Scientific Names Avoiding Confusion (Accuracy & Uniformity) Avoiding Ambiguity (Misnomers) in Classification “Fishes”: jellyfish (cnidarian), crayfish (crustacean), silverfish (insect) Standardization Provide insight into relationships between animals Example: Animals sharing genus are closer than animals sharing family Hierarchical classification Taxons Linnean Taxonomy Kingdom Phylum Class Order Family Genus Species Modern Taxonomy Subcategories Domains Advantage Taxa & Hierarchical similarity Disadvantage Differences in levels of similarities / differences for animals in different equivalent levels Lack of insight into evolutionary process / history Phylogeny Phylogenic Trees Branching Points Dichotomous Keys (Two-way split guides) Phylocode (Path on tree) vs. Taxa Relationships Rooting Basal Taxon Sister Taxa Polytomy Advantages Establish evolutionary relationships & history Links to hierarchical taxa Consideration of derived characteristics Disadvantages No information on rate of evolution Age of animals Easy to confuse common origin with origin from each other Hypothesis of Evolutionary Relationships Systematics Morphology / Physiology Ecological Role / Behaviors Paleontology Molecular Biology (DNA / RNA / Polypeptide sequence) Analogy (Homoplasies) vs. Homology (Molecular Organism) Fossil Evidence If common ancestor does not have trait = analogous Great degree of similarity in extremely complex structures homologous Phylogenetics Cladistics Clades Common Ancestry Monophyletic Paraphyletic Polyphyletic Shared characteristics (homologous structures) Shared derived character (Branching points) Ingroup vs. Outgroup Proportional Phylogeny & Cladistics Genome as Evolutionary Library Orthologous genes Paralogous Gene variety vs. phonotypical variety Molecular Clocking Using mutation rates to estimate time since evolutionary splits Calibration with fossil record Disavantage Possibility for variable rates or burst evolution Rate is an estimation based on average only Fossil record limitations Natural selection & mutation rates Genes & Evolution vs. clock speed Neutral = constant rate and fast Crucial = slower rate and variable Examples of Ticking Speeds rRNA Slowest to mutate (ancient splits) mtDNA Faster to mutate (recent speciation) Using multiple genes to clock Branch Lengths & Evolutionary History Constructing Trees Maximum Parsimony (Fewest evolutionary splits) Maximum Likelihood (Most common) Hypothesis of Evolutionary Relationships The Three Domains System & The Tree Life The Problem with Prokaryotic vs. Eukaryotic Cells Taxonomy Changes to old system Old Kingdom Monera Archae & Eubacteria Current Domains Archea Bacteria Eukarya Protista; Fungi; Plants; Animals Tree of Life vs. Ring of Life Horizontal Gene Transfer Conjugation Viral Transduction Organism Fusion Endosymbiosis The Main Taxa of Life [Know any major characteristics discussed in the videos] Archea Bacteria Eukarya Protista Fungi Plants Moss-like (Nonvascular) Ferns-like (Seedless vascular) Gymnosperms (Seed vascular) Angiosperms (Flowering plants) Animals Invertebrates Porifera Cnidarians Platyhelminthes Nematoda Rotifera Annelida Mollusca Arthropoda Echinodermata Chordata Invertebrate Chordates Tunicates Lancelates Vertebrata Jawless fish Cartilaginous fish Bony fish (Ray & Lobe finned) Amphibia Reptilia Mammalia Aves Major Animal Derived Characters Multicellularity Ingestive Heterotrophs Levels of Organization (Atom, Molecules, Organelles, Cells, Tissue, Organ, Organ System, Organism) Embryonic Development (Cleavage, Morula, Blastula, Gastrulation, Gastrula, Blastopore, Blastocoel, Archenteron, Mesoderm) Tissue Layers: No Tissue vs. Diploblastic (Endo vs. Mesoderm) vs. Triploblastic (Endoterm, Mesoderm, Ectoderm) Gastro-cavities (Two-way vs. one way) Body Plans Acoelomate Pseudocoelomate Coelomate Symmetry Asymmetric Radial Bilateral Cephalation Deuterostomes vs. Protostomes Segmentation Appendages