Lesson Overview Viruses Lesson Overview 20.2 Prokaryotes Lesson Overview Viruses Classifying Prokaryotes The smallest and most abundant microorganisms on Earth are prokaryotes—unicellular organisms that lack a nucleus. Prokaryotes DNA is located in the cytoplasm. Lesson Overview Viruses Bacteria Bacteria live almost everywhere Lesson Overview Viruses Bacteria Bacteria are usually surrounded by a cell wall that protects the cell from injury and determines its shape. The cell walls of bacteria contain peptidoglycan. Some bacteria, such as E. coli, have a second membrane outside the peptidoglycan wall that makes the cell especially resistant to damage. Lesson Overview Viruses Bacteria Some also have flagella that they use for movement, or pili, which in E. coli serve mainly to anchor the bacterium to a surface or to other bacteria. Lesson Overview Viruses Archaea Under a microscope, archaea look very similar to bacteria. The walls of archaea lack peptidoglycan, and their membranes contain different lipids. The DNA sequences of key archaea genes are more like those of eukaryotes than those of bacteria. Based on these observations, scientists have concluded that archaea and eukaryotes are related more closely to each other than to bacteria. Lesson Overview Viruses Archaea Many archaea live in extremely harsh environments. Lesson Overview Viruses Size, Shape, and Movement Prokaryotes are much smaller than eukaryotic cells. Rod-shaped prokaryotes are called bacilli. Spherical prokaryotes are called cocci. Spiral and corkscrew-shaped prokaryotes are called spirilla. Lesson Overview Viruses Size, Shape, and Movement Prokaryotes can also be distinguished by whether they move and how they move. Lesson Overview Viruses Nutrition and Metabolism Energy is released from these fuel molecules during cellular respiration, fermentation, or both. Prokaryotes vary in the ways they obtain energy and the ways they release it. Lesson Overview Viruses Nutrition and Metabolism: Energy Capture Lesson Overview Viruses Nutrition and Metabolism: Energy Release Lesson Overview Viruses Growth, Reproduction, and Recombination When a prokaryote has grown so that it has nearly doubled in size, it replicates its DNA and divides in half, producing two identical cells. This type of asexual reproduction is known as binary fission. During favorable conditions some bacteria can reproduce in 20 minutes. Lesson Overview Viruses Growth, Reproduction, and Recombination When growth conditions become unfavorable, many prokaryotic cells form an endospore—a thick internal wall that encloses the DNA and a portion of the cytoplasm. Endospores can remain dormant for months or even years. Lesson Overview Viruses Mutation Mutations are one of the main ways prokaryotes evolve. Mutations are random changes in DNA that occur in all organisms. Lesson Overview Viruses Conjugation Many prokaryotes exchange genetic information by a process called conjugation. During conjugation, a hollow bridge forms between two bacterial cells, and genetic material, usually in the form of a plasmid, moves from one cell to the other. Lesson Overview Viruses Decomposers By decomposing dead organisms, prokaryotes, supply raw materials and thus help to maintain equilibrium in the environment. Lesson Overview Viruses Producers Photosynthetic prokaryotes are among the most important producers on the planet. Food chains everywhere are dependent upon prokaryotes as producers of food and biomass. Lesson Overview Viruses Nitrogen Fixers All organisms need nitrogen to make proteins and other molecules. The process of nitrogen fixation converts nitrogen gas into ammonia (NH3). Ammonia can then be converted to nitrates that plants use, or attached to amino acids that all organisms use. Nitrogen-fixing bacteria and archaea provide 90 percent of the nitrogen used by other organisms. Lesson Overview Viruses Human Uses of Prokaryotes Prokaryotes, especially bacteria, are used in the production of a wide variety of foods and other commercial products. Some bacteria can digest petroleum and remove human-made waste products and poisons from water. Other bacteria are used to synthesize drugs and chemicals through the techniques of genetic engineering.