How do fossils form? Biological materials Conditions needed for preservation Types of preservation, types of fossils Biological Materials Vertebrate Crustacean Mollusk Plant DNA Protein DNA Protein DNA Protein DNA Protein Collagen Collagen Collagen Cellulose Mineral shell Lignin Mineral bone Teeth Fossilization 2 Conditions required for fossilization 1. Retard decay and chemical and physical breakdown 2. Burial 3. Fixation Fossilization 3 Step 1: Retard decay and chemical breakdown • Anaerobic environment Reduces decay, but acidity may destroy DNA, proteins, and carbonate-bearing minerals O- • Clay films Mg+ Ca++ Fe++ Fossilization C Carbonyl … first product of bacterial breakdown 4 Step 2: Burial • Removes object from destructive environment • Paves entry to rock record Fossilization 5 Step 3: Fixation • Reduces vulnerability to breakdown • Drying or freezing • Clay minerals fix organic material • Minerals entering cells • Mineral alteration - e.g. aragonite to calcite or dolomite Fossilization 6 Kinds of Fossils • Unaltered remains • Altered Hard Parts – Compression – Permineralization – Recrystallization – Replacement • Traces – Impression – Mold-Cast – Tracks, trails, and burrows – Coprolites - that’s fossil poop, by the way • Molecular fossils Fossilization 7 Unaltered Remains • Original tissues preserved intact Fossilization – Drying, freezing, or preservation in 8 amber Compressions QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Fossilization • 2D remains of 3D material • Retain organic material 9 Permineralization • Mineral infilling of porous spaces in skeletal elements like wood, bone, and echinoderm plates • Can preserve cellular detail Fossilization 10 Recrystallization Loss or blurring of original shell structure as shell is converted to interlocking crystals. Fossilization 11 Replacement • Original material dissolved away and replaced by another mineral Fossilization Pyritization, Silicification, Calicification, others 12 Impressions QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. 2-dimensional imprints without any organic material 13 Fossilization Mold / Cast QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Fossilization • Physical characteristics of organisms are impressed onto rocks, especially coarse porous rocks like sandstones.14 Ichnofossils • • • • Tracks Trails Burrows Other evidence that organisms were present • Only evidence for behavior Fossilization 15 Coprolites • Fossil Excrement • Extremely informative - what were they eating? Fossilization 16 Molecular fossils • • • • • • • Chlorophyll Flavinoids Collagen DNA Lipids Proteins And their biochemical breakdown projects Fossilization 17