Chapter 28: Arthropods & Echinoderms QuickTi me™ and a T IFF (Uncompressed) decompressor are needed to see thi s pi cture. Qui ckTi me™ and a TIFF (Uncompressed) decompressor are needed to see this pictur e. Echinoderms 28–1 Introduction to the Arthropods A. What Is an Arthropod? B. Form and Function in Arthropods 1. Feeding 2. Respiration 3. Circulation 4. Excretion 5. Response 6. Movement 7. Reproduction C. Growth and Development in Arthropods Phylum Arthropoda “joint foot” What Is an Arthropod? • Arthropods have a bilateral symmetry, a segmented body, a tough exoskeleton (made of chitin), and jointed appendages. • Arthropods are the largest phylum of animals with more than 750,000 species. • Living arthropods generally have fewer body segments and more specialized appendages than ancient arthropods (600 mya). QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Crustacean QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Trilobite Evolution of Arthropods QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. The Anatomy of a Grasshopper Antennae Compound eye Brain Digestive Malpighian tract tubules Ventral View Legs Heart Reproductive organs Mouth Salivary glands Anus Ganglia Tracheal tubes Nerve cord Spiracles Tracheal tubes Spiracle s Arthropods feed on respire using All types of foods Tracheal tubes reproduce using Internal fertilization Book lungs have well-developed External fertilization Book gills Heart Brain Muscles Form and Function in Arthropods Body Plan Exoskeleton, jointed appendages, specialized appendages for feeding, bilateral symmetry, cephalization. Feeding Herbivores, carnivores, omnivores, blood suckers, filter feeders, detritivores, and parasites. Highly adapted mouthparts for eating (sponging, piercing, grasping, grinding, lapping, chewing). Respiration Terrestrial Arthropods: Tracheal tubes extend throughout the body. Air enters and leaves through spiracles (small openings). Spiders use book lungs (layers of respiratory tissue stacked the pages of a book. Horseshoe crabs use book gills. Aquatic Arthropods: Featherlike gills exchange oxygen & carbon dioxide. Circulation Open circulatory system. A well developed heart pumps blood through arteries and then enters tissues. The blood is collected in a large sinus near the heart. Excretion Malpighian tubules (saclike organs) extract nitrogenous wastes from the blood and add them to the feces, or digestive wastes that move through the gut. Response Well developed nervous system. All arthropods have a brain. Most haave sense organs, such as eyes and taste receptors. Movement Muscle cells contract when stimulated by nerves. Muscles pull on exoskeleton to allow for flying, walking, or swimming. Reproduction Terrestrial: Reproduce sexually through internal fertilization (sperm deposited inside female body). Aquatic: Reproduce sexaully by internal or external fertilization. Growth and Development in Arthropods QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. When they outgrow their exoskeletons, arthropods undergo periods of molting (shed their entire exoskeleton, and grow a new one.) 17 yr. Cicada video http://www.bio.indiana.edu/~hangarterlab/broodx/broodxmovies/NSFmovie.htm 28–2 Groups of Arthropods A. Crustaceans (Subphylum Crustacea) B. Spiders & Their Relatives (Subphylum Chilicerata) 1. 2. 3. 4. Horseshoe Crabs Spiders Mites and Ticks Scorpions C. Insects & Their Relatives (Subphylum Uniramia) 1. Centipedes 2. Millipedes 3. Insects Groups of Arthropods Subphylum Crustacea Subphylum Chelicerata Subphylum Uniramia crabs, lobsters, shrimp, crayfish, barnacles horseshoe crabs, spiders, mites & ticks, scorpions Centipedes, millipedes, insects QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. •Crustaceans have two pairs of antennae, two or three body sections, chewing mouthparts called mandibles. •Most are aquatic. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. •Chelicerates have mouthparts called chelicerae and two body sections, and nearly all have four pairs of walking legs. •No antennae. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. •Uniraminans have jaws, one pair of antennae, and unbranched appendages. •Largest group of animals. The Anatomy of a Crayfish (subphylum Crustacea) Tail Abdomen Cephalothorax Swimmerets Carapace First antenna Mandible Walking legs Cheliped Second antenna The Anatomy of a Spider (subphylum Chelicerata) Cephalothorax Brain Abdomen Pumping stomach Heart Intestine Ovary Malpighian tubules Eyes Poison gland Pedipalp Fanglike chelicera Anus Spiracle Bases of walking legs Airflow Book Lung Spinnerets Silk glands The Anatomy of a Grasshopper (subphylum Uniramia) QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. 28–3 Insects A. What Is an Insect? 1. Responses to Stimuli 2. Adaptations for Feeding 3. Movement and Flight 4. Metamorphosis B. Insects and Humans C. Insect Communication D. Insect Societies 1. Castes 2. Communication in Societies Class Insecta What Is an Insect? • Insects have a body divided into three parts - head, throax, and abdomen. Three pairs of legs are attached to the thorax. • There are more insects than any other animal on earth. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Section 28-3 Insect Diversity Nonarthropod invertebrates (11%) Noninsect arthropods (12%) Vertebrates (4%) Insects (73%) Response to Stimuli • Compound eyes detect changes in color and movement. • Chemical receptors for taste and smell on their mouthparts. • Sensory hairs detect slight movements in air or water. • Well-developed ears hear sounds far beyond human range. Adaptations for Feeding • Three pairs of appendages are used as mouthparts, including a pair of mandibles. • Variety of mouthparts including; grinding, tubelike, spongelike, piercing, etc. Movement and Flight • Three pairs of legs used for walking, jumping, or capturing and holding prey. • Flying insects typically have two pairs of wings made of chitin. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Metamorphosis Adult Eggs Adult Eggs Incomplete Metamorphosis Complete Metamorphosis Larva Adult Nymph Nymph Immature Nymph Adult Larva Pupa Insects and Humans Beneficial •Bees, butterflies, and other insects pollinate crops. •Bees produce wax and honey. •Some insects are eaten as delicacies in Africa and Asia. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Harmful •Termites destroy wood structures. •Wasps produce painful stings. •Mosquitoes are a nuisance and can spread disease like malaria and west nile virus. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Insect Communication • Insects communicate using sound, visual, chemical (with pheromones), and dances. • Much of their communication involves finding a mate. Ex. Fireflies light up to signal for a mate. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Insect Societies QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Queen Winged male Major worker Minor worker • Ants, bees, termites, and some of their relatives form complex associations called societies. Each caste has a body form specialized for its role. Reproductive castes are females called queens. Reproductive males are called workers. Communication in Societies • Honey bees convey information about the type, quality, direction, and distance of food by “dancing.” • The round dance tells the other bees that food is nearby. • The waggle dance tells the other bees that food is a longer distance away. The direction of the waggle indicates the direction of the food. QuickT ime™ and a T IFF (Uncompressed) decompressor are needed to see thi s pi cture. Bee Dance Video: http://www.pbs.org/wgbh/nova/bees/dances.html 28–4 Echinoderms A. What Is an Echinoderm? B. Form and Function in Echinoderms 1. Feeding 2. Respiration and Circulation 3. Excretion 4. Response 5. Movement 6. Reproduction C. Groups of Echinoderms 1. Sea Urchins and Sand Dollars 2. Brittle Stars 3. Sea Cucumbers 4. Sea Stars 5. Sea Lilies and Feather Stars D. Ecology of Echinoderms Phylum Echinodermata “spiny skin” What Is an Echinoderm? • Echinoderms are characterized by spiny skin, an internal skeleton (endoskeleton), a water vascular system, and suction-cuplike structures called tube feet. Most adult echinoderms have five-part radial symmetry. Sea star Sea cucumber Sea urchin QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. QuickTi me™ and a T IFF (Uncompressed) decompressor are needed to see thi s pi cture. The Anatomy of a Sea Star Echinoderm Active Art http://www.phschool.com/webcodes10/index.cfm?fuseaction=home.gotoWebCode&wcprefix=cbe&wcsuffix=8284 Eyespot Endoskeletal plates Anus Stomach Digestive glands Ring canal Radial canal Madreporite Reproductive glands Tube foot Sucker Form and Function in Echinoderms Body Plan Five-part radial symmetry, spiny skin, internal skeleton, water vascular system, deuterostome (blastopore develops into an anus). Feeding Sea star pushes stomach into prey’s shell, digests prey in its shell, retracts stomach.Sea urchin uses five-part jaw to scrape algae and eat kelp. Sea cucumbers filter detritus in sand like a bulldozer. Sea lilies use tube feet to catch plankton. Respiration Water vascular system carries oxygen through body. In most species, thin walls of tube feet provide a surface for respiration. Some species use skin gills. Circulation Water vascular system carries nutrients throughout the body. Excretion Digestive wastes are released as feces through the anus. Nitrogen waste is passed into the water through tube feet and skin gills. Response Nerve ring surrounds the mouth and radial nerves connect the ring with body sections. Most have sensory cells to detect light, gravity, and prey chemicals. Movement Tube feet fill with water from the water vascular system. Muscles move plates of the endoskeleton. Reproduction Most Reproduce Sexually by external fertilization (sperm and eggs are released into open water, where fertilization takes place. Larvae have bilateral symmetry, adults have radial symmetry. Groups of Echinoderms Qui ckTime™ and a TIFF (Uncompressed) decompressor are needed to see this pictur e. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Sea urchins & sand dollars QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Sea cucumbers QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Sea stars QuickT i me™ and a T IFF (Uncompressed) decom pressor are needed to see this picture. Brittle stars QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture. Sea lilies Groups of Echinoderms Comparing Groups and Major Characteristics of Echinoderms Characteristic Sea urchins and sand dollars Brittle stars Sea cucumbers Sea stars Sea lilies and feather stars Feeding Detritivores Detritivores Detritivores Most carnivores Herbivores Shape Movement Disc- or globeshaped, no arms Star-shaped, arms Burrow in sandy ocean bottom or Move rapidly wedge in rock along ocean crevices using floor using arms moveable spines attached to endoskeleton Cucumbershaped, no arms Move slowly along ocean floor using muscular body wall to crawl Star-shaped, arms Creep slowly along ocean floor using arms Stalk with feathery arms Cannot move; attached to ocean bottom Ecology of Echinoderms • Echinoderms control the distribution of algae and other forms of marine life. • Crown-of-thorns sea star feeds on coral and is a serious threat to coral reefs around the world. QuickTime™ and a TIFF (Uncompressed) decompressor are needed to see this picture.