Utah Flora BOT2100 Lecture 4 Monocots Including: Agavaceae Liliaceae Poaceae (Graminae) Utah Flora Monocots BOT2100 Flowering plants with a ovary wall (carpels) Corollas and calyx generally not distinguishable sometimes bract-like others petaloid, called tepals Flower parts in 3’s Leaves generally with parallel veins A single cotyledon (1st leaf) Utah Flora Monocots Major Families Agavaceae – Agave, Yucca, sometimes included in Liliaceae Amaryllidaceae – Amaryllis, Narcissus Commelinaceae – spiderworts Cyperaceae – sedges Iridaceae – Iris, Gladiolus Juncaceae – rushes Liliaceae – Lilies, onions Poaceae – grasses Typhaceae – cattails BOT2100 Utah Flora 14 families Asparagales 1,122 genera BOT2100 26,070 species Agavaceae Worldwide Utah 10 genera 340 species 3 genera 13 species (Native and Introduced) DNA analyses places this family within as expanded Asparagaceae (Asparagus Family) Utah Flora Asparagales Agavaceae BOT2100 Agave Worldwide Utah (Native) 210 species 1 species •endemic to North, Central, and South America •leaves spirally arranged •ovary inferior •perianth regular •tepals fused (at least basally) and whitish to yellowish •fruit a capsule Utah Flora Asparagales Agavaceae Agave utahensis •restricted in Utah to Washington Co. •inflorescence narrowly cylindric (appears spicate) •leaves armed with prominent teeth copyright Stan Shebs, 2006 Stan Shebs, 2006 BOT2100 Utah Flora Asparagales Agavaceae BOT2100 Yucca Worldwide Utah (Native) 50 species 13 species •endemic to North, Central, and South America •leaves spirally arranged •ovary superior •perianth regular •tepals seperate, fleshy, and whitish •fruit a capsule Utah Flora Asparagales Agavaceae BOT2100 Yucca breviflora •restricted in Utah to Washington Co. •one of 2 tree-like Yucca species in Utah, all others are shrubs •leaves less than 2.5 dm long copyright UBC Botanic Garden Utah Flora Asparagales Agavaceae Yucca harrimaniae •most widely distributed Yucca in Utah •leaves concave on one side •other species with leaves flat on one side BOT2100 Utah Flora 11 families Liliales 67 genera BOT2100 1,558 species Liliaceae Worldwide 19 genera Utah 28 genera* (Native or Introduced) 610 species 39 species *The Utah flora recognizes many more genera in Liliaceae than current DNA analyses support. These genera have been split into their own families. Utah Flora Liliales Liliaceae BOT2100 Allium Worldwide Utah (Native and introduced) 800 species 13 species •perennial herbs, +/- scapose, with bulbs •inflorescence umbellate •perianth regular, all segments alike & petaloid •tepals separate and whitish to pinkish •ovary superior •fruit a capsule Utah Flora Liliales Liliaceae Allium acuminatum •widespread in Utah •leaves 2 to 4, opposite •bulbcoats with prominent, thick- margined veins (reticulations) BOT2100 Utah Flora Liliales Liliaceae BOT2100 Calochortus Worldwide Utah (Native) 60 species 8 species •restricted to North and Central America •perennial herbs with bulbs, stems leafy •inflorescence generally solitary •perianth regular, •outer segments bract-like, inner segments petaloid •tepals separate and whitish, yellowish, pinkish •ovary superior •fruit a capsule Utah Flora Liliales Liliaceae Calochortus nuttallii •widespread in Utah •stems erect •perianth whitish to purplish •hairy gland at base of tepals BOT2100 Utah Flora 17 families BOT2100 Poales 997 genera 18,325 species Poaceae Worldwide Utah 668 genera 10,035 species 91 genera 203 species (Native and Introduced) 3rd largest family in Utah All members of this family have a bract-like perianth (lemma and palea) and a caryopsis for fruit Utah Flora BOT2100 The Grass Leaf Sheathing •leaf blade forms a sheath as it surrounds the stem stem blade sheath Utah Flora BOT2100 The Grass Flower 4 Parts: Glumes •1st whorl (bract-like) •2 glumes, subopposite •alike or unlike one another Lemma •2nd whorl (bract-like) •1 lemma •different than palea •may look similar to glumes Palea •3rd whorl (bract-like) •1 palea •often thin & papery •enclosed inside lemma, mostly outer husk=glumes Utah Flora Poales Poaceae BOT2100 Stipa Achnatherum Worldwide Utah (Native) •spikelets pedicillate with a single floret •spikelets disarticulating above the glumes •glumes longer than the lemma •lemmas with simple awns (not branched) 100 species 17 species Utah Flora Stipa comata BOT2100 Poales Poaceae Hesperostipa comata •one of the most widespread species of Stipa in Utah •terminal awn 5 cm long or longer with hairs less than 1 mm long glumes awned lemma Utah Flora BOT2100 Poales Poaceae Elymus Agropyron Worldwide Utah (Native and Introduced) 150 species ~18 species •inflorescence a solitary, terminal spike •spikelets 1-several at each node •spikelets sessile with 1 to many florets •spikelets with a disarticulating rachis •disarticulating above the glumes, also •glumes can be short-awned, lemmas awnless Utah Flora Elymus cinereus Leymus cinereus •widespread in Great Basin •clump-forming perennials (can have short rhizomes) •inflorescence spicate •spikelets 3-7 at a node Poales Poaceae BOT2100 Utah Flora Poales Poaceae BOT2100 Bromus Worldwide Utah (Native and Introduced) 100 species 16 species •spikelets pedicillate with 2 to many florets •spikelets disarticulating above the glumes •glumes longer than the lemma to shorter than the lemma •lemmas with simple awns (not branched) Utah Flora Poales Poaceae Bromus tectorum •widespread exotic weed •very flammable when dry, good tinder for man-made wildfires •awns of the lemma less than 13 mm •panicles loosely flowered BOT2100 Utah Flora BOT2100 Poales Poaceae Phragmites Worldwide Utah (Native) 3 species 1 species •large, rhizomatous, wetland grasses (reeds) •spikelets pedicillate with 2 to many florets •spikelets disarticulating above the glumes •glumes shorter than the lemma Utah Flora Poales Poaceae Phragmites australis •widespread saline marsh grass •inflorescence large •branches drooping Copyright Environment Canada BOT2100 Utah Flora BOT2100 Assignment •Draw and label species provided in lab •Make sure to label those morphological features that helped you identify the sample (the characters in the key) •Identify & label all samples to family and genus •For Practice: use A Utah Flora to identify & label all samples to species (other than any we determined in class together)