Cyanobacteria and Algae Prokaryotes and Eukaryotes Prokaryotes Eukaryotes “primitive nucleus” “true nucleus” Lack clear nucleus and other inclusions Clear nucleus and other inclusions All organisms with cells that lack a nucleus All organisms with cells that have a nucleus Nutritional Requirements of Prokaryotes Methods of obtaining carbon Autotroph (“self-feeding) incorporate carbon into organic molecules from inorganic sources Heterotroph (“other feeding”) derive carbon from breakdown of organic compounds Methods of deriving energy Chemotroph (“chemical feeding”) obtain energy from catalyzing inorganic reactions Phototroph (“light feeding”) obtain energy by absorbing light photons Symbiotic Relationships Between Prokaryotes & Plants Rhizobium lives in soil Synthesizes enzyme nitrogenase which converts unusable N2 to plant-available ammonium (NH4+) Forms mutualistic relationship with legumes within root nodules Plant → carbohydrates & protection Bacterium → nitrogenase & other enzymes Both benefit from supply of fixed nitrogen Cyanobacteria “blue green algae” Photosynthetic chlorophyll a, phycobilins fix nitrogen Symbiotic Relationships Between Prokaryotes & Plants Other examples of symbiotic nitrogen fixing Bacteria Frankia – within cells of root nodules of alders Anabaena – association with water fern, Azolla Nostoc – invades cavities in gametophytes of hornworts & cycads Figure 17.11 Photosynthetic Protists “Algae” Algae = Photosynthetic Protists Variety of life histories, body forms, ecological roles Often named for distinctive colors Unicellular, colonial, multicellular Phylum Chlorophyta “green algae” Unicellular, colonial, multicellular Most found in freshwater, also some in shallow marine habitats Pigments: chlorophyll a & b, carotenoid Food stored as starch Cellulose cell walls Green Algae 50 µm (a) (b) (c) Algal Reproduction Zygotic Gametic diploid phase of life cycle is single-celled zygote Examples: Volvox, green algae, Chlamydomonas Multicellular sporophytes haploid phase of life cycle is single-celled gamete Examples: some brown algae like Cystoseira Sporic multicellular gametophytes & sporophytes Examples: Ectocarpus, Ulva (isomorphic), Laminaria (heteromorphic) Zygotic l.h found commonly in green algae Gametic l.h. found in some Brown algae, especially rockweeds (Fucus, Pelvetia, Cystoseira) Sporic l.h. found in kelps, green algae, land plants Zygotic Life Cycles Chlamydomonas Figure 18. Ulothrix Life Cycle Figure 18. Spirogyra Sexual Reproduction Phylum Chromophyta (Stramenopiles) All have 2 unequally sized flagella (hairy & smooth) Photosynthetic ones include: Chrysophyceae (golden algae) Bacillariophyceae (diatoms) Phaeophyceae (brown algae) Chrysophyceae “golden algae” Carotenoid pigments Usually biflagellated Most unicellular, some colonial Cell wall of cellulose Figure 28.17 25 µm Bacillariophyceae “diatoms” Unicellular algae with silica tests (cell wall) Important phytoplankton Contain chl. a & c, fucoxanthin Reproduction in Diatoms Phaeophyceae “brown algae” abundant in cool, shallow marine waters Largest algae, multicellular thallus Life history sporic or gametic More Chlorophylls a & c, fucoxanthin Carbohydrates can move through sieve elements & be stored (“mannitol”) Cell wall of cellulose & algin Blade Stipe Holdfast Gametic (film) Figure 18. Sporic Life Cycle Phylum Rhodophyta “red algae” (b) (c) Widespread Shallow to deep water Warm to cool water . Chlorophyl a, phycobilins complex life cycle involving three types of thallus structures (“triphasic life cycle”) Cell wall of cellulose, agar or carrageenan Triphasic Life Cycle Figure 18. Phylum Euglenophyta “euglena” “mixotroph”, contains chlorophyll a & b No cell wall Contains eyespot Lack cell wall Phylum Dinophyta “dinoflagellates” Unicellular Contain two flagella. 1 trails from the cell 1 encircles the cell disc-shaped chloroplasts with xanthophyll pigments Many with tiny projectiles Many produce toxins (Red Tides) Dinoflagellates Ecological & Economic Importance Phytoplankton Help build tropical reefs Base of aquatic food chains Produce about 4x food as earth’s crops Coralline green & red algae Medicine, food, and fertilizer Uses of algal cell walls diatomite for filtering, bulking, abrasive agar as culture medium, gel electrophoresis & in baking carrageenan used as stabilizer algin used to absorb water & as additive Kelp movie