POPULATION ECOLOGY M.Sc. Botany 4th sem POPULATION ECOLOGY • Population - A group of individuals of the same species occupying a particular space at particular time. • Population Ecology – Study of organismenvironment interactions at the population level. It is concerned with fluctuations in size and regulatory factors. CHARACTERISTICS OF A POPULATION Population density Patterns of Dispersion Age Structures Age structure/Diagrams/Population Pyramids Natality Mortality POPULATION DENSITY It is the number of individuals of a species per unit area in terrestrial ecosystem or per unit volume in aquatic ecosystem. Crude density: the density per unit total space. Specific/Economic/Ecological density: The density per unit habitat space i.e. available area or volume that can actually be colonized by the population. PATTERNS OF DISPERSION RANDOM UNIFORM/REGULAR CLUMPED NO PATTERN UNIFORM SPACING WITHIN A POPULATION INDIVIDUALS AGGREGATE INTO GROUPS INDIVIDUALS SPACED IN A PATTERN LESS, UNPREDICTABLE WAY RESULTS FROM INTERACTIONS AMONG THEMSELVES (COMPETITION) IN RESPONSE TO SOCIAL AGGREGATION OR UNEVEN RESOURCE DISTRIBUTION PATTERNS OF DISPERSION CLUMPED DISTRIBUTION IS MOST COMMON TYPE BECAUSE: Suitable physical, biological, and chemical conditions are patchy, not uniform. UNIFORM DISTIBUTION IS RARE. It is rare in nature. Many aniamls form social groups Many offsprings are not highly mobile and are forced to live “where they landed”. If it does occur, it is usually the result of fierce competition for limited resources. AGE STRUCTURE It is the proportion of individuals in each age group and is usually shown graphically The three age groups are: • Prereproductive • Reproductive • Postreproductive The relative duration of these age groups in proportion to lifespan varies greatly with different organisms. The age structure of a population dictates whether it will grow, shrink or stay the same size. AGE STRUCTURE DIAGRAMS/ POPULATION PYRAMIDS • The graphical representation of the number or percent in each age class. UPRIGHT BROAD BASED/TRIANGULAR BELL SHAPED/RECTANGULAR URN POPULATION/INVERTED A high percentage of young individuals A moderate proportion of young to old A low percentage of young individuals rapid population growth (Growing population) Relatively stable population (Aging Population) Show decrease in population (Aged Population) AGE STRUCTURE DIAGRAMS/ POPULATION PYRAMIDS • The graphical representation of the number or percent in each age class. UPRIGHT BROAD BASED/TRIANGULAR BELL SHAPED/RECTANGULAR URN POPULATION/INVERTED A high percentage of young individuals A moderate proportion of young to old A low percentage of young individuals rapid population growth (Growing population) Relatively stable population (Aging Population) Show decrease in population (Aged Population) AGE STRUCTURE DIAGRAMS/ POPULATION PYRAMIDS NATALITY MAXIMUM/ABSOLUTE/POTENTIAL/PHYSIOLOGICAL NATALITY: The theoretical maximum production of new individuals under ideal conditions, also called fecundity rate. ECOLOGICAL/RELAIZED NATALITY: The actual population increase under existing specific conditions, also called fertility rate. CRUDE BIRTH RATE: Births per 1000. CLUTCH SIZE: the number of young produced in each occasion. GENERATION TIME: the average time between birth and reproduction for an individual. MORTALITY Mortality: Number of individuals removed through death. MINIMUM/POTENTIAL MORTALITY: The theoretical minimum loss under ideal or non-limited conditions. ECOLOGICAL/REALIZED MORTALITY: The actual loss of individuals under a given environmental condition. CRUDE DEATH RATE: Deaths per 1000 VITAL INDEX: Birth-death ratio expressed as percentage. SURVIVORSHIP CURVE • Mortality • Mortality rate • Survivorship is the percentage of an original population that survives to a given age. • A survivorship curve is the graphic representation of the number of individuals in a population that can be expected to survive to any specific age. • Survivorship curves plot the proportion of individual alive at each stage, thus reflect species differences in reproduction & mortality. • Three types of survivorship curve: • Type – I (Convex) • Type – II (Constant) • Type – III (Concave) TYPES OF SURVIVORSHIP CURVE TYPES OF SURVIVORSHIP CURVE Type – I: Convex Late Loss High parental care Humans, annual plants and most primates have a Type I survivorship curve. I n a Type I curve, organisms tend not to die when they are young or middleaged but, instead, die when they become elderly. Species with Type I curves usually have small numbers of offspring and provide lots of parental care to make sure those offspring survive. TYPES OF SURVIVORSHIP CURVE • Type – II • Late Loss • Many bird species have a Type II survivorship curve. • In a Type II curve, organisms die more or less equally at each age interval. • Organisms with this type of survivorship curve may also have relatively few offspring and provide significant parental care. TYPES OF SURVIVORSHIP CURVE • Type – III • Early loss • Trees, marine invertebrates, and most fish have a Type III survivorship curve. • In a Type III curve, very few organisms survive their younger years. • However, the lucky ones that make it through youth are likely to have pretty long lives after that. • Species with this type of curve usually have lots of offspring at once—such as a tree releasing thousands of seeds—but don't provide much care for the offspring. LIFE HISTORY STRATEGIES • Life history means how organism survives and reproduces over the period of time. • Life history traits are • • • • Number, size and sex ratio of offspring Timing of reproduction Age and size at maturity Growth pattern • Organism has to maintain balance between growing, surviving and reproducing. • Energy available shoeld be properly allocated between growth, survival and reproduction. • So, trade offs are inevitable. LIFE HISTORY STRATEGIES • Mcarthur divided organisms into two categories on the basis of “strategies” of growth, survival and reproduction: • R-strategist species, • K- strategist species, R-SELECTED SPECIES • • • • • • • • • • • • • R is the symbol of birth rate, so birth rate is critical factor Small adults, mature quickly Early reproductive stage Reproduce often, with many small offsprings Little or no parental care and protection of offsprings Short lived, most offsprings die before reaching reproductive stage Live in places where resources are temporarily abundant Adapted to unstable environment Population size fluctuates wildly above and below K The species are Opportunistic Generalist niche Low ability to compete Early successional species K-SELECTED SPECIES • • • • • • • • • • • • K is the symbol of carrying capacity, so carrying capacity is a critical factor Large size, mature slowly, most die old Later reproductive age Peproduce slowly, with fewer and larger offsprings High parental care and protection of offsprings Long lives, most offsprings survive to reproductive age Population size fairly stable and usually close to k Adapted to stable climate Good competitors Specialist niche Late successional species Examples- Trees, humans