BIOMES Biomes are very large ecological areas on the earth’s surface, with fauna and flora (animals and plants) adapting to their environment. Biomes are often defined by abiotic factors such as climate, relief, geology, soils and vegetation. A biome is NOT an ecosystem, although in a way it can look like a massive ecosystem. Plants or animals in any of the biomes have special adaptations that make it possible for them to exist in that area. Many units of ecosystems are found within one biome. There are five major categories of biomes on earth. In these five, there are many sub-biomes, under which are many more well defined ecosystems. The Desert Biomes: They are the Hot and Dry Deserts, Semi-Arid Deserts, Coastal Deserts and Cold Deserts. The Aquatic Biomes: Aquatic biomes are grouped into two, Freshwater Biomes (lakes and ponds, rivers and streams, wetlands) and Marine Biomes (oceans, coral reefs and estuaries). The Forest Biomes: There are three main biomes that make up Forest Biomes. These are; Tropical Rainforest, Temperate and Boreal Forests (also called the Taiga) The Grassland Biomes: There are two main types of grassland biomes: the Savanna Grasslands and the Temperate Grasslands. The Tundra Biomes: There are two major tundra biomes— The Arctic Tundra and The Alpine Tundra. Biomes play a crucial role in sustaining life on earth. For example, The Aquatic biome is home to millions of fish species and the source of the water cycle. It also plays a very important role in climate formation. The terrestrial biomes provide foods, enrich the air with oxygen and absorb carbon dioxide and other bad gases from the air. They also help regulate climate and so on. Applying the idea of an ecosystem The ecosystem concept can be applied to any scale from a small pool of water to a whole ocean, or even to the biosphere. Whatever its size, an ecosystem is relatively self-contained and tends to maintain itself by recycling minerals. Therefore in a pond, organisms die and decompose and the nutrients in them are returned to the water. In turn, the autotrophs remove these nutrients from the water and use them in growth. Ecosystems are often thought of as closed systems, but they are not really closed. No ecosystem is completely self-contained; they all need an outside source of energy, usually the sun Ecosystem goods and services This is the extremely vital life-support services ecosystems provide to human life, its well-being and future economic and social development. For example: The benefits ecosystems provide include; food, water, timber, air purification, soil formation and Pollination. Energy Flow of ecosystems Ecosystems maintain themselves by cycling energy and nutrients obtained from external sources. The populations in an ecosystem interact through their feeding relationships. A typical community has: Autotrophs (also known as primary producers) which produce their own food. Herbivores (primary consumers) which eat the autotrophs. Secondary consumers which eat the herbivores Tertiary consumers which eat the secondary consumers Decomposers, mainly bacteria and fungi, which obtain nutrients by breaking down the remains of dead organisms At the first trophic level, primary producers (plants, algae, and some bacteria) use solar energy to produce organic plant material through photosynthesis. Herbivores—animals that feed solely on plants—make up the second trophic level. Predators that eat herbivores comprise the third trophic level; if larger predators are present, they represent still higher trophic levels. Organisms that feed at several trophic levels (for example, grizzly bears that eat berries and salmon) are classified at the highest of the trophic levels at which they feed. Decomposers, which include bacteria, fungi, molds, worms, and insects, break down wastes and dead organisms and return nutrients to the soil. On average about 10 percent of net energy production at one trophic level is passed on to the next level. Processes that reduce the energy transferred between trophic levels include respiration, growth and reproduction, defecation, and nonpredatory death (organisms that die but are not eaten by consumers). The nutritional quality of material that is consumed also influences how efficiently energy is transferred, because consumers can convert highquality food sources into new living tissue more efficiently than lowquality food sources. The low rate of energy transfer between trophic levels makes decomposers generally more important than producers in terms of energy flow. Decomposers process large amounts of organic material and return nutrients to the ecosystem in inorganic form, which is then taken up again by primary producers. Energy is not recycled during decomposition, but rather is released, mostly as heat (This is what makes compost piles and fresh garden mulch warm). Figure shows the flow of energy (dark arrows) and nutrients (light arrows) through ecosystems. An ecosystem is made up of organisms, which have established themselves in the given area and have continued to survive and have not become extinct. The species hence possess genes, which fit the environment and are tolerant to disturbances like flood, fire, drought; and a reproductive rate that balances the natural catastrophes. The birth rate of organisms will have to be optimum to avoid overpopulation and hence starvation. The human population is a good example. As technological evolution brings down our normal death rate, social evolution lowers the birth rate to strike a balance. Biological evolution is however much slower than social or technological. In ecosystems, organisms constantly adjust themselves to geologic or climatic changes and to each other. As an example, the bats developed sonar to find the moths and the moths developed ears sensitive to the bat’s frequency. The behavioral adaptations are also reflected in the anatomy or the body structure of the organisms. This evolutionary pattern is very common and is called character displacement. The process of life evolution started from lower plants and progressing to higher plants, lower animals, higher animals and finally to man. Biogeochemical cycle A biogeochemical cycle or substance turnover is a pathway by which a chemical substance moves through both the biotic (biosphere) and abiotic (lithosphere, atmosphere, and hydrosphere) components of Earth. A cycle is a series of change which comes back to the starting point and which can be repeated. Water, for example, is always recycled through the water cycle, as shown in the diagram. The water undergoes evaporation, condensation, and precipitation, falling back to Earth. Elements, chemical compounds, and other forms of matter are passed from one organism to another and from one part of the biosphere to another through biogeochemical cycles. The term "biogeochemical" tell us about the biological, geological and chemical factors. The circulation of chemical nutrients like carbon, oxygen, nitrogen, phosphorus, calcium, and water etc. through the biological and physical world are known as "biogeochemical cycles". The element is recycled, although in some cycles there may be places (called reservoirs) where the element is accumulated or held for a long period of time (such as an ocean or lake for water). Important biogeochemical cycles The most well-known and important biogeochemical cycles, for example, the carbon cycle, the nitrogen cycle, oxygen cycle What is The Carbon Cycle? The carbon cycle is very important to all ecosystems, and ultimately life on earth. The carbon cycle is critical to the food chain. Living tissue contains carbon, because they contain proteins, fats and carbohydrates. The carbon in these (living or dead) tissues is recycled in various processes. Let's see how this cycle works using the simple sketch below: Human activities like heating homes and cars burning fuels (combustion) give off carbon into the atmosphere. During respiration, animals also introduce carbon into the atmosphere in the form of carbon dioxide. The Carbon dioxide in the atmosphere is absorbed by green plants (producers) to make food in photosynthesis. When animals feed on green plants, they pass on carbon compounds unto other animals in the upper levels of their food chains. Animals give off carbon dioxide into the atmosphere during respiration. Carbon dioxide is also given off when plants and animals die. This occurs when decomposers (bacteria and fungi) break down dead plants and animals (decomposition) and release the carbon compounds stored in them. Very often, energy trapped in the dead materials becomes fossil fuels which are used as combustion again at a later time. The nitrogen cycle Nitrogen is also key in the existence of ecosystems and food chains. Nitrogen forms about 78% of the air on earth. But plants do not use nitrogen directly from the air. This is because nitrogen itself is unreactive, and cannot be used by green plants to make protein. Nitrogen gas therefore, needs to be converted into nitrate compound in the soil by nitrogen-fixing bacteria in soil, root nodules or lightning. To understand the cycle better, let us consider the diagram below: 1. Nitrogen is introduced into the soil by precipitation (rain, lightning). 2. Nitrates don’t only come from Nitrogen in the air. They can also be obtained by the conversion of ammonia, commonly used in fertilizers by nitrifying bacteria in the soil. Some root nodules can also convert nitrogen in the soil into nitrates. 3. Plants build up proteins using nitrates absorbed from the soil. 4. When animals like cows, eat these plants, they, in turn, use it to build animal protein. 5. When these animals (cows) poop, pee or die, the urea, excreta or carcass are broken down by decomposers and the nitrogen is reintroduced into the soil in the form of ammonia. 6. Nitrates in the soil can also be broken down by denitrifying bacteria (in specific conditions) and sent into the air as nitrogen. This process can help make the soil infertile because it will lack the nitrates needed for plant use. Once nitrogen gets back into the air, the cycle continues. Food chains All living things need to feed to get energy to grow, move and reproduce. But what do these living things feed on? Smaller insects feed on green plants, and bigger animals feed on smaller ones and so on. This feeding relationship in an ecosystem is called a food chain. Food chains are usually in a sequence, with an arrow used to show the flow of energy. Below are some living things that can fit into a food chain. A food chain is not the same as a food web. A food web is a network of many food chains and is more complex. See the food web illustration below you can pick out a basic food chain from the web: Green plants Grasshopper Frog Bird Hawk In the diagram above, the arrows show the direction of energy flow. It points to the animal doing the eating. Energy transfer Energy is transferred along food chains from one level to the next. Some of the energy is used up in growth, reproduction repair, movement and other ways, and not made available to the next level. Shorter food chains retain more energy than longer chains. Used up energy is absorbed by the environment. ECOTONE Ecosystem boundaries are not marked (separated) by rigid lines. They are often separated by geographical barriers such as deserts, mountains, oceans, lakes and rivers. As these borders are never rigid, ecosystems tend to blend into each other. This is why a lake can have many small ecosystems with their own unique characteristics. Scientists call this blending “ecotone”. BIODIVERSITY This refers to a variety of living organisms in an area Areas with a high biodiversity are those with lots of different species. Biodiversity can be considered at the following levels; habitat diversity- the number of different habitats in an area Species diversity- the number of different species and the abundance of each species in an area. E.g a woodland having many different species of plants, insects, birds and mammals. Genetic diversity- the variation of alleles within a specie MEASURING BIODIVERSITY 1. Sampling A sample of the population is taken and estimates about the whole habitat is based on the sample. Sampling can be random or non-random. To avoid bias in the results, the sample should be random Sometimes it’s necessary to take a non-random sample. For example, when there’s a lot of variety in the distribution of species in the habitat and you want to make sure all the different areas are sampled. There are three types of non-random sampling Systematic-this is when samples are taken at fixed intervals Opportunistic- this is when samples are chosen by the investigator. Stratified- this is when different areas in a habitat are identified and sampled separately in proportion to their part of the habitat as a whole. Biodiversity is affected by species richness and species evenness. The greater the species richness and species evenness in an area, the higher the biodiversity. Species richness is the number of different species in an area. It’s measured by taking random samples of a habitat and counting the number of different species. Species evenness is a measure of the relative abundance of each species in an area. The more similar the population size of each species, the greater the species evenness. Its measured by taking random samples of a habitat, and counting the number of individuals of each different species Example: Habitat x and y both contain two different species and 30 individual organisms. Habitat x Habitat y Species 1 28 15 Species 2 2 15 Total 30 30 Species richness in the two habitats is the same -2 In habitat y the individual organisms are more evenly distributed between the different species-it has greater species 2. Index of diversity Species richness is a very simple measure of biodiversity. But species present in a habitat in very small numbers shouldn’t be treated the same as those with bigger populations. An index of diversity is another way of measuring biodiversity. It is calculated using an equation that takes into account both species richness and species evenness. Calculate an index of diversity (d) using this formula d = N (N-1) ⅀ n (n-1) N = total number of organisms of all species n = total number of organisms of one species ⅀= sum of When calculating an index of diversity using this formula, the higher the number you end up with, the more diverse the area is. If all the individuals are of the same species (i.e. no biodiversity) this index will have a value of 1. Example: There are 3 different species of flower in this field-a red species, a white and a blue. There are 11 organisms altogether, so N=11. There are 3 of the red species, 5 of the white and 3 of the blue. So the species diversity index of this field is: d= = 11(11-1) 3(3-1) + 5(5-1) + 3(3-1) 110 6+20+6 = 3.44 GENETIC DIVERSITY