HYDROLOGICAL PROCESSES Hydrol. Process. 27, 687–699 (2013) Published online 20 December 2012 in Wiley Online Library (wileyonlinelibrary.com) DOI: 10.1002/hyp.9626 Will catchment characteristics moderate the projected effects of climate change on flow regimes in the Scottish Highlands? R. Capell,* D. Tetzlaff and C. Soulsby School of Geosciences, University of Aberdeen, Aberdeen, AB24 3UF, UK Abstract: Regional climate models were used with the UKCP09 weather generator to downscale outputs from the HadCM3 General Circulation Model, to project climate change by 2050 in the Scottish Highlands. The resulting hydroclimatic data were used to drive a tracer-aided hydrological model to assess likely changes in flow regimes in three experimental catchments. These are located along a hydroclimatic transect from the wet, mild western Highlands (Strontian), through the colder, more continental central Highlands (Allt a’ Mharcaidh), to the drier eastern Highlands (Girnock). At all sites, temperatures are projected to increase by around 2 C, with associated increases in potential evapotranspiration. Precipitation is expected to increase by around 10–15% at Strontian but remains slightly changed at the Allt a’ Mharcaidh and Girnock. However, the seasonal distribution of precipitation is projected to change, increasing in winter and decreasing in summer. Cautious interpretation of model outputs indicates that flows are likely to change accordingly at all sites, though the characteristics of each catchment result in some subtle differences. At Strontian, marked increases in winter high flows are expected, at the Allt a’ Mharcaidh, reduction in winter snowfall and reduced snowpack will increase winter high flows and moderate the influence of spring melts. In the Girnock, decreasing summer low flows are the most notable element of change. These subtle differences in response to climatic drivers are consistent with the distinct storage characteristics of the catchments which in turn reflect their landscape evolution histories. Copyright © 2012 John Wiley & Sons, Ltd. KEY WORDS climate change; inter-site comparison; modelling Received 17 May 2012; Accepted 15 October 2012 INTRODUCTION Anthropogenic emissions of greenhouse gases (GHGs) have altered the composition of the atmosphere, thereby changing the global climate (IPCC, 2007). The resulting regional impacts are likely to increase in the coming decades, but there remains much debate on the precise nature of these changes and how well they can be predicted by climate models (Beven and Alcock, 2012). Consequently, whilst the potential effects on water resources are likely to be significant (Gleick, 2010), they are also highly uncertain (Beven and Alcock, 2012). Heterogeneity in hydroclimate and catchment characteristics in contrasting geographical regions also means that the hydrological functioning of catchments can be variable (Buttle, 2006; Tetzlaff et al., 2009; Carey et al., 2010), resulting in different likely changes to external hydroclimatic drivers and different intrinsic sensitivity of catchments to regional climate change (e.g. Capell et al., 2012b). Projecting the likely effects of climate change on the water balance and flow regimes in different catchments is a major challenge, yet it is a prerequisite for assessing the implications for water resources, stream chemistry (e.g. Laudon et al., 2011) and in-stream ecology (Kruitbos et al., 2012). Critical to such projections is the *Correspondence to: Rene Capell, School of Geosciences, University of Aberdeen, Aberdeen, AB24 3UF, UK E-mail: rene.capell@gmail.com Copyright © 2012 John Wiley & Sons, Ltd. derivation of reasonable forecast data from climate models and the appropriate use of rainfall-runoff models to assess hydrological change. The cascade of uncertainty here is potentially great, and several authors have urged extreme caution in interpretation of such studies (Hall, 2007). Changes in drivers such as precipitation and evapotranspiration are the primary factors that will determine hydrological responses to climate change at the catchment scale. However, the physical form and structure of catchments will determine their sensitivity to change (e.g. Tetzlaff et al., 2011). The way that catchments partition, store and then release water controls their flow regime. This in turn determines the resistance of the catchments flow regime to change induced by external forcing factors and its resilience or rate recovery following major perturbations (Carey et al., 2010). For example, region-specific partitioning of precipitation into different flow paths and subsurface storages and the intra-annual effects on the flow regime has been shown to mediate climate change impacts, resulting in regionally variable hydrological changes (Tague and Grant, 2009; McNamara et al., 2011). However, many climate change impact studies have been restricted to individual catchments in single geomorphic provinces (e.g. van Roosmalen et al., 2009; Poyck et al., 2011). Comparison across climate gradients and in different geomorphic regions has potential in differentiating climatically driven impacts and the degree to which these can be moderated by catchment structure and landscape organisation (Tetzlaff et al., 2008; Carey et al., 2010; Jones et al., 2012). 688 R. CAPELL, D. TETZLAFF AND C. SOULSBY Upland landscapes in the northern mid- and highlatitudes are sensitive to climate change (Viviroli et al., 2011). Small differences in temperature determine the frequency with which the 0 C temperature threshold is crossed and the longevity of periods above and below this critical value. This controls precipitation type, the magnitude and timing of snow accumulation and subsequent melt (Shinohara et al., 2009). Being able to project the likely consequences in northern headwaters is of critical societal relevance (Smith, 2011). Downstream flows from upland areas provide a suite of critical ecosystem services, including the provision of water supplies, maintenance of water quality and sustaining in-stream ecology (Viviroli et al., 2011). Climate warming may also interact with other agents of environmental change in ways that may compromise these ecosystem services. For example, it may have indirect effects by promoting land use changes, due to natural ecohydrological responses or adaptive management to changing temperatures and/or water availability. These land use changes themselves can modify the catchment water balance in ways that may mitigate or enhance climate change impacts, but such feedbacks are extremely difficult to assess (e.g. Brown et al., 2005; Quilbe et al., 2008). Nevertheless, they can have implications for water resource management with increased flood regimes and reduced dry weather flows being two potential results of climate non-stationarity (e.g. Tague and Grant, 2009; Milly et al., 2008). Using climate projections from General Circulation Model (GCM) to drive hydrological models is the most common method of assessing the likely consequences of climate change on catchment hydrology (e.g. Christensen et al., 2004; Fowler and Kilsby, 2007; van Roosmalen et al., 2009). Here, we examine three Scottish sites from the North-Watch (Northern Watershed Ecosystem Response to Climate Change: http://www.abdn.ac.uk/northwatch/) international inter-comparison project that explores the inter-linkages between climate, hydrology and ecology. We examine a west-east climatic transect in Scotland where the three upland catchments give a regional perspective on the responses to climatic change. Climate change projections are based on the UK Climate Projections 2009 (UKCP09; Murphy et al., 2009), which provide probabilistic climate change functions derived from perturbed physics regional climate model (RCM) ensembles. Synthetic time series derived from UKCP09 projections were used to force a tracer-aided conceptual hydrological model that was developed specifically for Scottish Highland landscapes. The model structure and parameter ranges were constrained to those identified as behavioural on the basis of their ability to simulate stream flow dynamics as well as both geochemical source area tracers and isotopic time domain tracers in upland catchments (Capell et al., 2012a). This intercomparison tests the hypothesis that catchment landscape properties can moderate climate change impacts. The specific objectives are (1) To assess the projected climate change impacts on precipitation and temperatures in three Scottish upland catchments along a west-east transect, (2) to Copyright © 2012 John Wiley & Sons, Ltd. use projected synthetic climate forcing (2050s time slice) in a hydrological model to examine the impact of climate change on the water balance and hydrological dynamics of these catchments at annual, seasonal and monthly scales and (3) to assess how the differences in hydrologic response of the catchments might reflect catchment characteristics that influence their resistance (i.e. the degree to which runoff is coupled with precipitation, relating to drainage efficiency and storage capacity), and resilience (i.e. a measure of the degree to which a catchment can return to normal functioning following perturbations from extreme hydrological events (Carey et al., 2010)). STUDY SITES Three upland catchments situated along a west-east climate transect across Scotland were compared to assess the likely impact of projected climate changes on flow regimes (Figure 1). The Strontian (STR, 8 km2) is located near the Atlantic coast, the Allt a’ Mharcaidh (MHA, 10 km2) in the more ‘continental’central highlands of the Cairngorm mountains, and the Girnock (GIR, 30 km2) in the eastern rain shadow. All three catchments are montane upland sites, characterised by broadly similar steep terrain, with >600 m of relief and low permeability metamorphic or igneous bedrock geology. However, they differ substantially in climate conditions and landscape evolution history. The latter factor results in differences the distribution and thickness of unconfined quaternary sediments and soils in the catchments (Figure 1, Table I). The westerly STR catchment rises from near sea level to over 700 m. Annual precipitation is around 2700 mm mainly brought by Atlantic frontal systems; the ocean moderates temperatures which have an annual mean of 8 C (Table II). Winter temperatures generally remain above freezing, and snow is typically <5% of inputs and restricted to short periods. Bedrock geology is comprised of schists and gneiss. Drift cover is largely restricted to low permeability sediments in lower slope areas; soil cover here is dominated by peaty gleys (histosols) with rankers (regosols) in elevated areas (Ferrier and Harriman, 1990). The Hydrology Of Soil Types (HOST) classification scheme (Boorman et al., 1995) describes these soils as being dominated by lateral flow paths, and thus hydrologically responsive with little dynamic storage change (Table I). The vegetation is characterised by coniferous forest plantations in the lower catchments and moorland grasses at high altitudes. The combination of intense precipitation and catchment topography, soils and geology results in a very flashy hydrological regime (Table II), with groundwater accounting for less than 20% of annual runoff (Soulsby et al., 2010). The MHA catchment is colder due to its location and higher altitude. Mean annual temperatures are 5 C, and annual precipitation is ~1100 mm with up to 30% falling as snow (Soulsby et al., 1997), the climate is sub-arctic in above 800 m. The bedrock is granite but largely covered by drift. At higher altitudes, periglacial drifts and bedrock Hydrol. Process. 27, 687–699 (2013) 689 EFFECTS OF CLIMATE CHANGE IN CATCHMENTS ALONG A HYDROCLIMATE TRANSECT Figure 1. Catchment topography, stream network and location of the three sample catchments on the Scottish mainland. Scotland map shows average annual precipitation from 1971 to 2000 (UK Met Office) Table I. Physical catchment characteristics of the three studied catchments Topography Geology (%) HOST soil classes (%) Land Cover (%) Strontian Allt a’ Mharcaidh Girnock Location West Coast Western Cairngorms Eastern Cairngorms Area (km2) Mean elevation (m) Relief (m) Median slope (degrees) Igneous Metamorphic 4,5,7: Alluvial soils 15: Peaty podzols and peaty gleys 17: Humus iron podzols 19, 22, 27: Rankers 29: Blanket peats Forest Grassland Wetlands Heather Moorland Montane/rocky 8 340 737 14 10 704 779 16 99.8 0.2 0 35 30 7 28 5 30 405 619 9.9 52 48 2 60 25 9 4 8 5 18 63 fractures form groundwater stores (Soulsby et al., 1999). In lower areas, glacial drift exceeds 10 m in depth and has relatively high porosity and permeability in places. At higher altitudes, freely draining alpine soils have formed beneath montane heath. In steeper areas, podzols support heather (Calluna) dominated moorland, grading to regenerating Scots Pine (Pinus sylvestris) forest on lower Copyright © 2012 John Wiley & Sons, Ltd. 100 0 71 0 29 0 48 52 59 36 slopes. On gentle slopes, deep peats support boreal blanket bog. Storm runoff is generated by overland flow from the peats, fed by shallow lateral flow from the podzols (Wheater et al., 1993; Jenkins et al., 1994). Drainage in the alpine soils and podzols recharges significant groundwater stores (Soulsby et al., 2009). Geochemical hydrograph separations showed that Hydrol. Process. 27, 687–699 (2013) 690 R. CAPELL, D. TETZLAFF AND C. SOULSBY Table II. Summary of water balance variables during the observation periods in the three study catchments Observation period Annual Q (mm) Annual P (mm) Annual ET (mm) Mean daily flow (mm d 1) Q5 (mm d 1) Q95 (mm d 1) Strontian Allt a’ Mharcaidh Girnock 1998/99 to 2001/02 2493 2751 258 6.8 1990/91 to 1992/93 824 1163 339 2.3 2005/06 to 2007/08 666 1138 472 1.82 25.2 0.33 5.9 0.79 5.1 0.17 groundwater provides over 50% of annual runoff with high baseflow (Table II) (Soulsby et al., 1998). The GIR catchment is in the eastern rain shadow; it is the driest sited with a lower mean elevation and high mean annual temperature of around 6 C and <10% of precipitation falls as snow. As with the other sites, the landscape history is characterised by past glaciations; here, the wide valley bottoms (Table I and II) are largely covered by low-permeability glacial drift. The underlying bedrock comprises low permeability metamorphic rocks with granite intrusions (Soulsby et al., 2007). Soils are mainly peats and gleys in the upper catchment, and humus-iron podzols in lower areas (Tetzlaff et al., 2007). Unlike the MHA, the vegetation is more managed mainly to maintain habitat for animals and birds that are quarry in field sports involving shooting. Thus, grazing by Red deer (Cervus elaphus) is intense in the lower catchment, and burning to create habitat for moorland birds (e.g. grouse (Lagopus lagopus)) occurs in the upper catchment. Forestry is restricted to small enclosed areas on the lower slopes. Overland flow and shallow sub-surface storm flow from the peaty soils dominate the storm hydrograph (Tetzlaff et al., 2007). Groundwater recharge is restricted to more freely draining soils (Soulsby et al., 2007) and geochemically based hydrograph separations suggest that groundwater accounts for ca. 30% of annual runoff with low base flow (Table II) (Soulsby et al., 2005). DATA AND METHODS Measured and projected data Precipitation, runoff and air temperature data were available for each catchment at daily resolution for threeyear or four-year observation periods between 1990 and 2008, each covering drier and wetter years (Table II). Monthly potential evapotranspiration (PET) was estimated using the Thornthwaite equation, then distributed to daily amounts using air temperature as weighting factor with PET equal to zero under freezing conditions, and a simple volumetric correction to actual evapotranspiration volumes from the average water balance (Thornthwaite and Mather, 1955). Precipitation and discharge data were used to calibrate conceptual rainfall-runoff models for the Copyright © 2012 John Wiley & Sons, Ltd. study catchments which were then forced with probabilistic climate change scenario data (see below). Climate change projections provided by the UK Climate Projection 2009 (UKCP09) network were used (Murphy et al., 2009). These provide probabilistic projections at a range of IPCC GHG emission scenarios and future time slices for the British Isles. Within the scope of this study, an average GHG emission scenario (IPCC scenario A1B) for the 2050s decade was selected. Previous modelling studies in the Scottish Highlands have already contextualised this relative to other ‘worse case’ (A1FI) and ‘best case’ (B1) scenarios (Capell et al., 2012b). The UKCP09 climate projections are downscaled projections from HadCM3 GCM projections and a 1960 to 1991 baseline period. The approach addressed uncertainty in the HadCM3 model due to imperfect process representation within a Bayesian framework and an ensemble of HadCM3 parameterisations. These were downscaled to a 25 25 km resolution using a perturbed physics ensemble of 11 HadRM3 RCMs, generating probability density functions of target climate variables at a monthly time scale (Murphy et al., 2009). A stochastic weather generator (WG) is accessible through UKCP09, which allows generation of further downscaled stochastic daily time series projections of baseline and future precipitation, PET and air temperature at a 5 5 km resolution. UKCP09 WG simulations are statistically downscaled RCM simulations which use monthly rainfall statistics for the 1960 to 1991 baseline period (Jones et al., 2009). The WG produces stationary synthetic time series. Here, 100 synthetic time series of 55 km grid cells covering each study catchments were generated, using randomly sampled RCM variants (encompassing uncertainty from RCM and GCM variants), thus giving a range of projections addressing the model uncertainty inherent in the climate model and down-scaling chain. Each of the 100 time series span a 30-year sequence (assuming stationary climate conditions) to sample extreme events with large return periods. The synthetic baseline data are also generated by the WG, reproducing the statistical properties of the 1960 to 1991 baseline data (further details on the WG in Jones et al., 2009). The synthetic time series reflect anticipated changes in GHG emissions under the chosen scenario and future time slice and also give an indication of the uncertainty in these projections. Hydrological impact assessment using tracer-aided models Changes in runoff were projected using hydrological models with a common structure that was calibrated for each catchment. The calibration period encompassed short periods (i.e. up to 3 years) within the record that included both wetter and drier years than average (Table II). The model had a priori constrained structures developed from application to other catchments in the Scottish Highlands where a range of catchment types can be modelled with a broadly similar structure (see Hrachowitz et al., 2011). This was based on the use of Hydrol. Process. 27, 687–699 (2013) EFFECTS OF CLIMATE CHANGE IN CATCHMENTS ALONG A HYDROCLIMATE TRANSECT geochemical and isotopic tracers as additional constraints in an iterative process that produced models with structures and parameter ranges that could simulate the catchment runoff and tracer response. Details are given by Capell et al. (2012b); briefly, this resulted in a model that could account for the different geographical sources of runoff (i.e. shallow and deeper flow paths) and their temporal dynamics (i.e. fast and slow flow paths) as indicated by the tracer responses, providing a firmer process-basis for impact assessment. In summary, the model comprises two linear stores and a saturation overland flow (SOF) conceptualisation (Figure 2). These conceptualise baseflows from a deeper groundwater zone and a more dynamic storage volume (e.g. in the upper soils and lower hillslopes) and SOF or similar saturationdependent lateral runoff generation related to responsive soil cover (Capell et al., 2012a). The model partitions precipitation between recharge into the soil store and direct SOF runoff depending on a non-linear function of the soil storage volume. Evaporation is drawn from and limited by the soil storage. The baseflow storage receives a constant recharge from the overlying soil. A degree-day snow module, driven by daily air temperatures, similar to the HBV model (Bergström, 1975) is used to estimate snow accumulation and melt. The model requires five 691 calibrated parameters in the storage, and two in the snow routines. To project runoff responses in the three catchments, behavioural parameter sets were selected by calibration to measured hydrometric data. The synthetic time series derived from UKCP09 projections were then used to force the models with parameterisations identified as behavioural to predict the runoff response. Parameter sets were generated using a Monte Carlo approach sampling wide initial parameter ranges; 5105 parameter sets were generated. Model performances were evaluated by three objective functions; the Nash-Sutcliffe efficiency (NSE), the NSE of log-transformed runoff (logNSE) and the coefficient of determination (R2) (cf Gupta et al., 1998). Parameter ranges were evaluated against performance peaks and threshold performances determined (Table III), above which, parameter sets were accepted as behavioural. From the accepted parameter sets, 100 were randomly chosen and used to generate runoff simulations with each synthetic projection and baseline time series, deriving a range of rainfall-runoff model uncertainty. For assessing the long-term changes, the projected daily time series of runoff from the hydrological model were then aggregated to monthly, seasonal and annual statistics, along with the characteristics and magnitudes of projected climate change for comparison. Additionally, the projected number in snow days, i.e. days during which the models projected a resident snow layer, were compared. RESULTS Projected climate change impacts on precipitation and temperatures Figure 2. Hydrological model structure showing water fluxes (dotted lines) and storage components of the lumped model structure. Precipitation (P) is routed to a day-degree snow routine on a temperature threshold condition. A linear shallow storage (S1) generates fast runoff responses (Q1). An additional saturation excess overland flow runoff (QSOF) is generated as a function of storage in S1. Baseflow (Q2) is generated from a linear storage S2 The projected changes in climate for the 2050s are visualised as annual summaries in Figure 3 and summarised in Table IV. The projected increase in annual average temperatures of about 2 C above the present is consistent for all sites (Figure 4). Inter-annual variation (shown as whiskers in the boxplots), however, increases in the projected average temperatures indicating the uncertainty (Figure 4a). However, in addition to the inherent projection uncertainty, this may also indicate an increase in extreme temperature patterns, which can in turn mean a greater impact on catchment hydrology. Annual precipitation in the STR catchment, already the highest in the baseline figures (2400 mm), is projected to increase by approximately 12% in the 2050s with increased inter-annual variability (Figure 4b). Such increase is not apparent in the more central and easterly Table III. Model performance for all three catchments, showing maximum performances and performance thresholds for behavioural parameter sets Maximum model Performance (performance threshold) Copyright © 2012 John Wiley & Sons, Ltd. NSE logNSE R2 Strontian Allt a’ Mharcaidh Girnock 0.83 (0.6) 0.85 (0.7) 0.83 (0.6) 0.77 (0.6) 0.75 (0.6) 0.78 (0.6) 0.71 (0.6) 0.85 (0.6) 0.78 (0.6) Hydrol. Process. 27, 687–699 (2013) 692 R. CAPELL, D. TETZLAFF AND C. SOULSBY Figure 3. Comparison of annual air temperature and water balance terms along the west-east transect as pie charts. Segment sizes of precipitation, evapotranspiration and runoff scaled individually, allowing inter-catchment comparison of each term (segment sizes not to scale for within-catchment water balances) Table IV. Summary of annual change statistics relative to baseline (1960–91) for projected climate and hydrological impact in the study catchments. Long-term annual averages and standard deviations (in parentheses) of air temperature, water balance terms and number of snow days Time period Temperature Precipitation Runoff Evapo-transpiration No. of snow days baseline 2050s baseline 2050s baseline 2050s baseline 2050s baseline 2050s Strontian 8.4 10.7 2421 2711 2070 2327 602 688 1 0 sites of the MHA and GIR where precipitation changes are within +2%. The inter-annual variability also seems more stable at these sites, especially compared to temperatures. PET largely follows the temperature trend, with projected increases of 86 mm, 69 mm and 70 mm for STR, MHA and GIR, respectively (Figure 4c). The increased temperatures result in decreases in predicted number of snow days (Table IV). These Copyright © 2012 John Wiley & Sons, Ltd. (0.3) (0.9) (262) (357) (251) (346) (19) (41) (1) (0) Allt a’ Mharcaidh 4.4 6.7 1296 1310 1043 1075 420 489 119 51 (0.4) (0.9) (174) (183) (163) (167) (20) (35) (23) (5) Girnock 5.6 7.7 1025 1022 826 805 479 549 6 2 (0.3) (0.9) (149) (162) (134) (148) (19) (34) (1) (2) numbers can only be considered as indicative, given the simplicity of the snow model used, but the greatest impact will be in the MHA. Snow influence under baseline conditions is highest (119 days) here, with only 6 days in the GIR and one or two days at STR. Snow days is projected to decrease to ~40% of the baseline in the MHA, so the potential hydrological influence is significant. In the other catchments, snow has a limited baseline Hydrol. Process. 27, 687–699 (2013) EFFECTS OF CLIMATE CHANGE IN CATCHMENTS ALONG A HYDROCLIMATE TRANSECT 693 Figure 4. Variation and projected change of annual air temperature and water balance components in the study catchments, derived from synthetic weather generator time series for a baseline period (1961 to 1990; baseline) and projections for the 2050s decade under IPCC scenario A1B (2050s). (a) Air temperature, (b) precipitation, (c) potential evapotranspiration and (d) runoff. Box plots showing median and inter-quartile range, whiskers extending to data extremes water balance influence. Thus, the hydrological impact is much less important. Seasonality of the long-term projections for hydroclimatic drivers is shown in Figure 5. Air temperatures increase at all sites in all seasons (Figure 5a). From a hydrological perspective, the changes in autumn (OND) and winter (JFM) in the sub-alpine MHA catchment are particularly notable as raising temperatures corresponds to decreasing winter snow. Changes in seasonal precipitation distributions also project increases during the autumn/winter half year (OND, JFM) at all sites, most notably at STR (Figure 5b). All sites show precipitation decreases during summer (JAS), but only the GIR shows a notable decline in spring (AMJ) (Figure 5b). All sites show increases in PET mainly in the spring (AMJ) and summer (JAS). The magnitude of projected increases is consistent with current differences with STR > GIR > MHA. Changes to long-term monthly temperature and precipitation regimes (Figures 6, 7, and 8, respectively) provide more detailed resolution. The long-term precipitation averages are expected to increase most notably on the west coast (STR) from September to May, and slightly decrease in July to August, though monthly totals remain high throughout the year. The projected changes are comparatively subtle in MHA and GIR, and rather a shift and the annual totals remain almost unchanged. Nevertheless, winter increases at MHA are evident from Copyright © 2012 John Wiley & Sons, Ltd. October to May. Decreases are apparent in July and August, consistent with STR. In contrast, GIR projects decrease between May and September. Monthly PET increases greatest in summer for all sites, being highest at STR (Figures 6c, 7c and 8c). Current hydrological dynamics and projected future impacts Annually, the runoff from the catchments reflects precipitation, with yields under baseline and projected conditions approximately twice as high in the STR catchment compared to MHA and GIR (Table IV, Figure 4d). Reflecting precipitation inputs, annual runoff at STR is projected to increase by over 10%, whilst slight increases (+1%) are projected for MHA, and small decreases in the GIR catchment ( 3%) indicate overall limited change. This is despite likely increases in PET, which probably reflects the greater proportion of annual precipitation in winter when PET is lowest. Seasonally (Figure 5d), spring and summer flows are projected to decrease, while autumn and winter flows increase (MHA, STR) or remain stable (GIR). However, the inter-annual variation remains fairly stable in the projected annual and seasonal values, with the exception of the winter flows in the STR, where variation increases similar to the precipitation pattern, Figure 5b. The monthly flow regimes reveal greater resolution (Figures 6d, 7d and 8d). Most dramatic changes are in Hydrol. Process. 27, 687–699 (2013) 694 R. CAPELL, D. TETZLAFF AND C. SOULSBY Figure 5. Variation and projected change of seasonal air temperature and water balance components in the study catchments, derived from synthetic weather generator time series for a baseline period (1961 to 1990; baseline) and projections for the 2050s decade under IPCC scenario A1B (2050s). (a) Air temperature, (b) precipitation, (c) potential evapotranspiration and (d) runoff. Box plots showing median and inter-quartile range, whiskers extending to data extremes Figure 6. Projected change in average monthly air temperature (a) and water balance components (b–d) in the Girnock catchment, derived from synthetic weather generator time series for a baseline period (1961 to 1990; baseline) and projections for the 2050s decade under IPCC scenario A1B (2050s). Error bars show standard deviations of the monthly aggregates of all projection time series Copyright © 2012 John Wiley & Sons, Ltd. Hydrol. Process. 27, 687–699 (2013) EFFECTS OF CLIMATE CHANGE IN CATCHMENTS ALONG A HYDROCLIMATE TRANSECT 695 Figure 7. Projected change in average monthly air temperature (a) and water balance components (b–d) in the Mharcaidh catchment, derived from synthetic weather generator time series for a baseline period (1961 to 1990; baseline) and projections for the 2050s decade under IPCC scenario A1B (2050s). Error bars show standard deviations of the monthly aggregates of all projection time series Figure 8. Projected change in average monthly air temperature (a) and water balance components (b–d) in the Strontian catchment, derived from synthetic weather generator time series for a baseline period (1961 to 1990; baseline) and projections for the 2050s decade under IPCC scenario A1B (2050s). Error bars show standard deviations of the monthly aggregates of all projection time series Copyright © 2012 John Wiley & Sons, Ltd. Hydrol. Process. 27, 687–699 (2013) 696 R. CAPELL, D. TETZLAFF AND C. SOULSBY winter at STR, where flows increase up to 25% (December). A general increase in autumn and winter flows at STR is apparent, with only a slight decline in July and August. In the MHA, winter flows increase between December and February. Decreases during the baseline snowmelt period (March–April) are consistent with loss of winter snow and the consequent runoff increases in the preceding winter. Only drier months from July to August result in decreases in projected runoff. In GIR, decreased precipitation and increasing evaporation lead to more consistent decrease in runoff from April to October, and the projections indicate new low-flow extremes in dry summers. DISCUSSION AND CONCLUSIONS Projected changes in climate We examined three upland sites across a climatic transect for a regional perspective on the hydrological impact of climatic change in the Scottish North-Watch catchments. The patterns of change projected are largely consistent with other studies in other UK uplands (Fowler and Kilsby, 2007; Capell et al., 2012b), with increasing temperatures throughout the year, increased winter and decreased summer precipitation. The three sites also highlight the influence of regional climate patterns and their relevance for the hydrological change. The visualisation in Figure 3 shows that the hydroclimate of STR is dominated by the large precipitation inputs, a feature which is likely to be accentuated in future with around 10% more precipitation by 2050. The contrast with unchanged or drier conditions in central (MHA) and eastern (GIR) Scotland suggests the continuation of a trend over the past decades (Werritty, 2002). Despite the lack of input change at MHA and GIR, subtle changes in precipitation distribution and type are likely. At MHA, increased winter temperatures are likely to decrease the size and longevity of snow accumulation, with more winter precipitation falling as rain, and more rapid melts (cf Soulsby et al., 1997). The main change expected at GIR is decreased summer precipitation, which, with likely increases in temperature and potential PET, may have implications for summer flows. Projected changes to the hydrological regime The likely changes in runoff regimes mainly reflect the influence of the changing climatic drivers. However, it is also to an extent mediated by the characteristics of the catchments themselves. At STR, the increased precipitation drives increased runoff which is likely to be much greater than losses from increased evapotranspiration. Higher winter flows may increase the frequency and magnitude of storm flows. Although reduced summer precipitation and increased temperatures are likely to cause small declines in summer flows, these will remain relatively high. For MHA, annual runoff changes little, though temperature increases will affect snowfall, an important factor in the hydrology of this area. This increases runoff Copyright © 2012 John Wiley & Sons, Ltd. in November, December and January as more precipitation falls as rain and snowpack accumulation is reduced. Although a March snowmelt peak is predicted to remain, it is less marked than previously, and snowmelt continuation into April and May is likely reduced though flows are little changed. Flows in July and August are projected to decline by ~10–15% as rainfall declines and evaporation increases. At GIR, the changes in interannual precipitation distribution, with reduced summer inputs and increased ET, are more likely caused by drier conditions, and flows decreased by ~ 20% between June and September. Catchment characteristics and sensitivity to change – the resistance and resilience of catchments Previous studies have shown how the form and structure of montane catchments which determine the magnitude of different water storages and the partitioning of precipitation along different flow paths can influence sensitivity to climate change (Tague et al., 2008). In the case of STR, the thin soils, impermeable geology and steep slopes result in limited catchment storage, so there is very little resistance – in terms of coupling between precipitation-runoff – to precipitation changes (Carey et al., 2010). Modelling studies indicate that dynamic storage in the catchment is typically in the order of <20 mm, thus precipitation increases directly increases runoff (Soulsby et al., 2011). Tracer studies have shown very short mean residence times in the catchment, typically in the order of a few months (Hrachowitz et al., 2010a), which infers total passive storage in the catchment of at least 300 mm (Soulsby et al., 2009). The projected climate changes with increasing precipitation are likely to change these storages little with little capacity to buffer effects on increased streamflows. The MHA, with deeper soils and more permeable drift deposits, has relatively larger groundwater stores which contribute around half of the annual discharge. Thus, the hydrological regime is highly attenuated compared to STR. Mean residence times in the catchment are in the order of 3 years (Hrachowitz et al., 2010a), inferring passive storage probably >2000 mm (Soulsby et al., 2009). In the drier climate of the MHA, summer soil moisture deficits may exceed 50 mm giving some dynamic storage (Soulsby et al., 2011). Increased winter precipitation results in moderate increases in winter runoff indicating little resistance to increased high flows. In summer, however, the sub-surface storages allow a certain resistance to reduced precipitation and increased PET because winter recharge into the deeper subsurface is able to sustain low flows (cf Table II). This resistance of the GIR in summer is lower, probably because the drift deposits here are less permeable. This results in a catchment with a high coverage of responsive peat and gleyed soils with only around 30% of annual flow being derived from groundwater. Consequently, the catchment already experiences low summer flows as groundwater inputs are lower than at MHA (Tetzlaff et al., 2007). Tracer-based Hydrol. Process. 27, 687–699 (2013) EFFECTS OF CLIMATE CHANGE IN CATCHMENTS ALONG A HYDROCLIMATE TRANSECT studies indicate mean residence times of about 18 months (Hrachowitz et al., 2010a), inferring a passive storage of around 1000 mm, though dynamic storage may vary by around 70 mm. Thus, the catchment storages have potentially less resistance to changing seasonal precipitation, and flows during periods of low summer precipitation are likely to decrease accordingly. These differences in the likely potential for catchments to buffer or mediate the impacts of change in climatic drivers are quite subtle. This contrasts with the large-scale differences that geology has been shown to have on sensitivity to environmental change in other montane regions (cf Tague et al., 2008; Tague and Grant, 2009). Nevertheless, it highlights likely intra-regional subtleties that will play out in the coming decades and may have management implications. Thus, although the general patterns of warming, reduced snow, increased winter runoff and reduced summer flows are similar to the recent trends and projections for many other northern temperate/subboreal regions (see Smith, 2011 for a review), the exact impacts may be catchment specific (cf Beven, 2001). Potential for improving projections with tracer-aided models Results showed that the ensemble projections and downscaling chain result in wide uncertainty ranges for the projected climate variables. This precludes the estimation of absolute values and makes evaluation of impact scenarios necessarily cautious. However, the longterm annual, seasonal and monthly projections were considered instructive for comparing the three catchment hydroclimates. The large sample size of RCM realisations provides a more stable basis for the climate change projections over the coming decades, even though only one GCM provided the global change conditioning, this shortcoming is likely to be offset by using several GCM in an ensemble (Teutschbein et al., 2011). However, this could increase uncertainty if different outcomes result. Additional uncertainties are associated with the hydrological modelling. The use of bespoke tracer-aided models for Scottish upland catchments helped constrain some of the structural and parameter uncertainty (Capell et al., 2012a). However, through calibration, the modelling the approach assumes stationarity in the way the catchment generates runoff (Wagener, 2007), which is a limitation considering that the projected climate changes might cause non-linear responses in the longer term. The probabilistic nature of the projection time series, the inclusion of several model parameterisations and the relatively short projection timescale somewhat offset this risk, but at the cost of wide projection ranges and sometimes ambiguous results. Evolving catchment hydrology – the greatest uncertainty? Perhaps the largest unknown in terms of nonstationarity in the catchment response is how the vegetation cover changes, which, in turn, is likely to affect the catchment soils. This will either result from natural vegetation adaptation to the changing climate, or, more likely, be dependent on how land management Copyright © 2012 John Wiley & Sons, Ltd. 697 strategies evolve in response. At all three sites, land management is deliberate. At STR, commercial forestry in the lower catchment and rough grazing for sheep in the upper catchment are the current characteristics. However, even with forest cover on thin, peaty soils in a wet climate increased evapotranspiration from forest cover probably has limited potential in increasing catchment scale water storage capacity (Tetzlaff et al., 2007). At MHA, nature conservation is the main aim of land management, and natural forest regeneration is active with Scots Pine (Pinus sylvestris), and given the soil cover and altitudinal limits of the tree line, perhaps as much as half of the catchment could be afforested. In rapidly growing, dense stands of Scots Pine, this has the potential to substantially increase evapotranspiration, mainly through interceptions losses which could reduce groundwater recharge and summer low flows, though this will be offset by an increased proportion of winter precipitation (Haria and Price, 2000). There will also be effects on snowmelt processes, though most snow in the catchment falls above the tree line. Although GIR currently has only small areas of forest cover, like the MHA, it lays within the Cairngorm National Park, where restoration of natural forests is a management objective (Hall, 2007). Expansion of forest cover could similarly exacerbate summer low flows decreases projected by climate change and the GIR could be less resistant to these than MHA. Interestingly, some of the potential conflicts and paradoxes may result from adaptive management of land and water resources to climate change. For example, in all three streams, salmonids (especially Atlantic salmon (Salmo salar) are important components of the freshwater ecosystem. Concerns have been expressed over likely increases in stream temperatures projected under climate change scenarios, as salmonids are cold water species that can suffer both lethal and sub-lethal physiological effects of elevated temperatures; reduced summer flows will reduce flow volumes and increase the risk of high temperatures (Malmqvist and Rundle, 2002; Hrachowitz et al., 2010b). A potential solution is increased riparian tree cover in catchments, which has a moderating effect on temperatures by reducing short wave inputs to the water column (Imholt et al., 2012). Clearly, extension of forest cover beyond the riparian zones to cover larger parts of catchments has potential to induce hydrological change. CONCLUSIONS Downscaled GCM climate projections were used to drive a tracer-informed hydrological model to assess the effects on stream flow in three long-term experimental sites in the Scottish Highlands across a west-east hydroclimatic gradient. Mean annual temperature increases of around 2 C are expected at all three sites, as is a shift to increased winter precipitation and reduced snow influence. All sites are expected to see reduced summer precipitation and increased PET, and only the most Hydrol. Process. 27, 687–699 (2013) 698 R. CAPELL, D. TETZLAFF AND C. SOULSBY westerly site is expected to see a change (increase) in precipitation totals. Projected hydroclimatic change will cause responses in stream flow regimes, with all three catchments showing relatively little resistance. Subtle differences in drift geology in the Allt a’ Mharcaidh will mean summer low flows will be more resistant to change than at the other sites, though in a climate that is expected to remain wet and cool, winter flows are likely to be resilient. Adaptive management of land use has the potential to affect flows as much as projected climate change; this is an area where future research will be directed (Viviroli et al., 2011). 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