UNIVERSITI MALAYSIA KELANTAN EGT2023- PALEONTOLOGY TOPIC: GROUP ASSIGNMENT: PALEOECOLOGY AND PALEOENVIRONMENT ESSAY SUBMITTED TO: DR. NURSUFIAH BTE SULAIMAN PREPARED BY: SEMESTER 3 SESSION OF 2021/2022 1.0 INTRODUCTION Definition of Paleoecology Paleoecology refers to an ecology of the past which is the study and understanding of the relationship between organisms and their environment. It is because the past organisms and environment could not be watched directly, much paleoecological research is concerned with determining past occurrences, distributions, and abundances of organisms and reconstructing past populations, communities, landscapes, environment, and ecosystems using geological and biological evidence. Paleoecology can be studied by everyone in any period of Earth’s history in which life was present. However, the main parts of paleoecology to ecology are from the most recent geological interval, the Quaternary period, covering the last 2.8 million years. The emphasis of this article is on Quaternary paleoecology because of its strong links with contemporary ecology. The research approaches and philosophy of paleoecology are outlined and different types of Quaternary paleoecological evidence and stages in a paleoecological study are summarized. Two major approaches to the interpretation of paleoecological data are discussed as paleoecological reconstructions and ecological paleoecological. Definition of Paleoenvironment A paleoenvironment is simply an environment that has been maintained in the rock record at some time in the past. Identifying a particular paleoenvironment from the data available in the sediments is not so simple. Determining the geochemical parameters associated with the environment can be very difficult largely due to diagenesis. Unique and complex conditions during deposition of the paleoenvironment may also contribute to the difficulty of deciphering the significance and meaning of the geochemical signatures in the rock record. The paleoenvironment that is commonly studied from a geochemical perspective includes a wide variety of marine and freshwater systems. Of particular interest, marine systems are filled with basins, lagoons, carbonate shelves, banks and reefs, and deltas. In freshwater systems, paleolacustrine deposits provide a wide range of geochemical studies. 2.0 METHODOLOGY FOR PALEOECOLOGY AND PALEOENVIRONMENT Paloecology Method Paleoecology aims to provide the perfect model of the living environment of previously existing species discovered as fossils today. Reconstructing past environments requires the consumption of archives, such as sediment sequences, proxies such as micro or megafossils, and other sediment characteristics that provide evidence of the biota and physical environment, and chronology, such as obtaining absolute or relative dating of events in the archive. This type of reconstruction takes into account complicated relationships between environmental elements including temperature, food supply, and sunlight. The fossilization or diagenesis of the encircling strata sometimes loses or distorts much of this information, making interpretation challenging. Then, Imbrie and Kipp (1971) proposed the factor analytic transfer function, a statistical tool for analyzing fossil assemblages that combines factor analysis and linear regression of data on current ecological requirements of organisms, which was published in the 1970s. This is a major step forward for paleoecological surveys from a combined qualitative description based on the numerical count of taxa to statistical data sets with accompanying errors and confidence levels. When it comes to integrating paleoecologic findings into decision-making, the capacity to assess confidence in interpretations is crucial. Furthermore, advances in stable isotope biogeochemistry and chronologic techniques enabled more precise comparison and interpretation of the physical and biological components of ecosystems preserved in geological archives. Refinement of the use of transfer functions and analog datasets allowed for more precise and statistically defensible estimations of historical climate conditions. These advancements allowed paleoecology to widen its reach and begin to address research concerns directly relevant to present resource management issues. Method for Paleoenvironment Human settlement and subsistence activities are based on interaction with the natural world. With the reconstruction of the broader environmental context, archaeologists can identify the main external catalysts of culture change. Modern paleo environmental reconstruction methods have used techniques developed over the last century. Archaeologists also need to explain the sedimentology of surface deposits to understand the evolution of landforms. Furthermore, soils formed in stable and weathered sedimentary deposits are an indicator of climate criticality in the past. Soils were also first described in an excavation test unit using a macro-scale classification scheme. However, growing microscopic techniques such as soil micromorphology in thin sections and DNA sequences of endemic microbiota are often also used. Various types of plant and animal communities also provide details about the environment that are currently critical as they are often dependent on precipitation and temperature. In general, smaller organisms are more limited in their habitat. Therefore, the remnants of flowers and microfauna often provide the highest level of accuracy in environmental re -conservation. Finally, nitrogen and oxygen can be tested from a wide variety of organic materials. Thus, it can provide precise information on precipitation and biotic communities which can be used for understanding the paleoenvironment. 3.0 CASE STUDY RELATED WITH PALEOECOLOGY AND PALEOENVIRONMENT Case Study from North Wadi Qena, Egypt’s Eastern Desert in the Context of Paleoecology, Cenomanian/ Turonian Mass Extinction of Macroinvertebrates The paleoenvironment of the North Wadi Qena area is very much important in assessing the effects on the Late Cretaceous biota. A paleoenvironment interpretation like this would include biotic evidence from invertebrate fossils as well as geologic or sedimentary features like paleosols and sequence stratigraphy interpretations. Several scientists have shown that characteristics of modern macrofaunas are strongly associated with environmental variables, implying that invertebrate fossils might be valuable as paleoenvironment indicators. Major extinctions are cyclical and may be caused by similar, if not identical, mechanisms raised the exciting idea that mass extinctions may be comprehended easily. However, the studies of numerous Phanerozoic mass extinctions (e.g., Kauffman 1988) have revealed that comparisons are not as straightforward as expected. These investigations have revealed considerable disparities in the biotic, lithologic, and geochemical fabrics of many of the mass extinctions. Despite these differences, the broad pattern of repopulation appears to be comparable between biotic crises, and certain events have a very correlative biotic response from a broad range of viewpoints. This investigation aims to establish limits on environmental changes that impact macroinvertebrates’ biota. Harries (1993, 1999), Harries and Kauffman (1990), Harries and Crispin (1999), Kauffman (1988, 1995) and Kauffman and Harries (1996) conducted substantial research on the Cenomanian/Turonian mass extinction. To uncover faunal variations between Cenomanian/Turonian assemblages and recreate how these assemblages are impacted by platform flooding and crises, detailed analyses of macroinvertebrate fossil groups are needed. Several benthic macrofossil groups are functional markers of environmental changes, such as fluctuations in water depth, oxygen content, nutrient availability, or water energy, and there are kinds of research on these groups as ecological proxies have been conducted. The emphasis is on the questions of why many bigger benthic animals vanished during the late Cenomanian periods and did not reappear on Turonian. Furthermore, oyster bivalve assemblages are environmental proxies, do their presence and distribution imply changes in water depth temperature and nutrient supply? Paleoecology and ecosystem restoration: case study from Chesapeake Bay Another case study that we can relate to is paleoecology and ecosystem restoration from the Chesapeake Bay. Generally, this case study tells us about the paleoecological and paleoclimatic reconstructions which are related to recovery planning. These happen in the eastern US, the Chesapeake Bay, the largest estuary in the nation. The first one is environmental degradation and restoration that occur in the Chesapeake Bay. There are vast agriculture, urbanization, and a large number of populations in the Chesapeake Bay. Unfortunately, all of these entities affect the water quality in rivers, tributaries, and the Bay. Near the watershed was lived by the Native American community. Usually, they consume the food acquired through foraging, hunting, and maize farming (Smith 1989). These lifestyles consist an effect on land cover compared to the changes made by European colonists. For instance, the forests there are being cleared for 80%. And, this clearance caused erosion and downstream sedimentation. Thus, the water clarity in the Bay is reduced. As soon as several farming lands emerged reforested, the fertilizer consumption and development worsens the water quality there (Willard et al. 2003). The restoration progresses to enhance the water quality values about more than $18 billion by the year 2010 (Chesapeake Bay Commission 2003). Fortunately, Chesapeake Bay is in the top of target for restoration planning since 1983. It started when the Chesapeake Bay Program (CBP) was established. This strategy focused on define water-quality criteria to protect aquatic living resources. Miocene Vertervrate and Invertebrate burrows defining compound paleosols in the Pawnee Creek Formation, Colarado, USA Two case studies from the Paleogene and Neogene of the North American midcontinent are presented in detail. The case studies involve sites in northeastern Colorado, U.S.A., within the late Eocene-early Oligocene White River Formation (Hembree and Hasiotis 2007a,b) and the middle Miocene Pawnee Creek Formation (Hembree and Hasiotis 2008). Both sites consist of deposits of alluvial systems dominated by localized channel conglomerate and sandstone and abundant, widespread paleosols. Despite their general similarities, the two units were deposited under very different climatic conditions with distinct landscapes and ecosystems. Much of this information is recorded in the trace fossils preserved within the paleosol. Whereas the Eocene-Oligocene transition is well documented in the marine record (Miller et al. 1987; Miller 1992; Ivany et al.2003), details of the transition in the continental environment are limited by the scattered distribution of sedimentary basins and the difficulty of correlating continental and marine strata from this interval of time. Terrestrial plant- and vertebrate-fossil assemblages of North America suggest the development of lower mean annual temperature and precipitation in the late Eocene and early Oligocene (Hutchin-son 1992; Leopold et al. 1992; Retallack 1992; Stucky 1992; Wolfe 1992; Sullivan and Holman 1996; Retallack et al.2004). Other than that, four Middle-Upper Jurassic sections from central Saudi Arabia have been investigated to evaluate microfacies types and macroinvertebrate paleo communities and to interpret their paleoecology and paleoenvironments. The studied Jurassic successions are part of the Middle-Upper Callovian Tuwaiq Mountain Limestone and the Middle-Upper Oxfordian Hanifa Formation. 4.0 USES OF PALEOECOLOGY AND PALEOENVIRONMENT A role for paleoecology in landscape management There is a role for paleoecology and other forms of long-term science in restoring productive also biodiverse landscapes. In many areas of Europe, rural land abandonment caused the degradation of semi-natural woodlands, meadows, and heathlands due to the discontinuation of customary practices (Weissteiner et al. 2011). Many species of high conservation value are specifically sociated with these cultural landscapes and necessitate continuous and active management. In Europe, 55 of the 231 listed habitat types of European interest depend on, or benefit from, continued customary practices (EEA 2011). Paleoecological and historical studies have consistently shown a decrease in biodiversity associated with the loss of customary land-management practices, and higher biodiversity associated with low-intensity land uses, which generate mosaics of wooded and grassy habitats. For example, Swedish semi-natural wood pastures and forest meadows are important examples of relict cultural landscapes in Europe. Paleoecological research showed a decrease in the diversity of forest taxa within the last centuries that comes from the reduction in grazing and mowing practices. Paleoecological knowledge is a key in the management of these landscapes (Bradshaw and Lindbladh 2005). Similarly, remnants - a function of old grazing practices, such as oak meadows (oak savannas) - are now threatened through the lack of grazing and management (Fig. 1). Dahlström et al. (2008) show that the highest species richness in plants in central Sweden is found in what today is semi-natural grassland subjected to a long continuity of grazing. It is estimated that since 1870, 99.7% of the semi-natural hay meadows have been lost, the few remaining semi-natural hay meadows are now managed mainly for conservation purposes. In this case, long-term data may provide knowledge on management practices that have now been lost. Historical analyses show that meadows and chestnut orchards are being replaced by forest expansion, with an overall loss in landscape heterogeneity, biodiversity, and ecosystem services. Pastoral customary practices often combine grazing with fire management to maintain wood pastures, but since the 18th century, many state authorities have actively used fire both in agriculture and forestry. Shakesby et al. (2011) showed an increase in wildfire and erosion following rural land abandonment in the Mediterranean region, associated with increased shrub cover and afforestation with flammable species. Paleoecological studies in the Pyrenees have shown that these pastoral landscapes have been systematically managed by fire over millennia (Mazier et al. 2006) and similar results have been shown for the Iberian Peninsula (Gil-Romera et al. 2010). In Scandinavia, the ecological role of fire in both semi-natural and oldgrowth forests is now being realized (Bradshaw and Hannon 2006). In the Finnish boreal forests, dominated by the conifers Scots pine (Pinus sylvestris) and Norway spruce (Picea abies), local slash-and-burn practices were common until the early 20th century. The mixed forests and grazing lands within the boreal forests are now experiencing degradation and reduction in species diversity as slash and burn practices have been discontinued (Myllyntaus et al. 2002). In other parts of Scandinavia, the historical importance of fires in the boreal forests is shown by the presence of fireadapted species (Bradshaw and Hannon 2006). In all cases listed above, paleoecological knowledge is essential in managing and restoring landscapes, and for the protection and management of rare species through sound management practices (Gillson 2015). Uses of Paleoenvironment Paleoenvironment shows us the environments in which the ancient organism plants and animals lived long ago. This work is called paleoenvironmental reconstruction. Paleoenvironmental reconstruction can be used to develop a model of a sedimentary environment that existed in ancient prehistoric times. Sedimentary environments encompass both the depositional environment was sedimentary rocks are laid down as layers called strata as well as the preservation environments where ancient organisms were fossilized. If the organisms stayed in the same environment and were not fossilized there after their death, then you can also think about a sedimentary environment as a sort of habitat where the organisms lived? Paleoenvironments also play an important role in the interpretation of archaeological data. It chronicles past changes in climate and vegetation and can be used to predict potential human subsistence patterns. It also gives us about the date of specific geological time in ancient history. Paleoenvironments in archaeology teach us about past cultures through learning about artifacts, animal bones, and occasionally human bones. The knowledge about these artifacts provides us with some insight into what life was like for people who left no written record. Paleoenvironmental data can be useful for understanding the environment before, during, and after site occupation, and it can reveal environmental changes that influenced technology, social structure, human subsistence, and settlement strategies in modern society. 5.0 CONCLUSION Unique and complex conditions during deposition of the paleoenvironment may also contribute to the difficulty of deciphering the significance and meaning of the geochemical signatures in the rock record. Paleoecology method to provide the perfect model of the living environment of previously existing species discovered as fossils today. This type of reconstruction takes into account complicated relationships between environmental elements including temperature, food supply, and sunlight. This is a major step forward for paleoecological surveys from a combined qualitative description based on the numerical count of taxa to statistical data sets with accompanying errors and confidence levels. Furthermore, advances in stable isotope biogeochemistry and chronologic techniques enabled more precise comparison and interpretation of the physical and biological components of ecosystems preserved in geological archives. Refinement of the use of transfer functions and analog datasets allowed for more precise and statistically defensible estimations of historical climate conditions. These advancements allowed paleoecology to widen its reach and begin to address research concerns directly relevant to present resource management issues. Several scientists have shown that characteristics of modern macrofaunas are strongly associated with environmental variables, implying that invertebrate fossils might be valuable as paleoenvironment indicators. Other than that, four Middle-Upper Jurassic sections from central Saudi Arabia have been investigated to evaluate microfacies types and macro-invertebrate paleo communities and to interpret their paleoecology and paleoenvironments. In many areas of Europe, rural land abandonment caused the degradation of semi-natural woodlands, meadows, and heathlands due to the discontinuation of customary practices. Historical analyses show that meadows and chestnut orchards are being replaced by forest expansion, with an overall loss in landscape heterogeneity, biodiversity, and ecosystem services. In the Finnish boreal forests, dominated by the conifers Scots pine and Norway spruce, local slash-and-burn practices were common until the early 20th century. Paleoenvironmental data can be useful for understanding the environment before, during, and after site occupation, and it can reveal environmental changes that influenced technology, social structure, human subsistence, and settlement strategies in modern society. 6.0 REFERENCES ⮚ Ahmed. A. (2008a, January). Cenomanian/Turonian mass extinction of macroinvertebrates in the context of Paleoecology; A case study from North Wadi Qena, Eastern Desert, Egypt. Academiaedu. Retrieved November 10, 2021, from https://www.academia.edu/3392436/Cenomanian_Turonian_mass_extinction_of_ macroinvertebrates_in_the_context_of_Paleoecology_A_case_study_from_North _Wadi_Qena_Eastern_Desert_Egypt Ekblom. A. & Gillson. L. (2017). The importance of paleoecology in the conservation and restoration of cultural landscapes. Pages Magazine, 25(2), 88– 89. https://doi.org/10.22498/pages.25.2.88 ⮚ H. Stephen T. & H. Daniel I (2008). Miocene vertebrate and invertebrate burrows defining compound paleosols in the Pawnee Creek Formation, Colorado, U.S.A. Palaeogeography Palaeoclimatology Palaeoecology, 230(3–4), 349–365. https://doi.org/10.1016/j.palaeo.2008.07.019 ⮚ W. Debra A. & C. Thomas M. (2007). Frontiers in Ecology and the Environment. Paleoecology and Ecosystem Restoration: Case Studies from Chesapeake Bay and the Florida Everglades, 5(9), 491–498. https://doi.org/10.1890/070015 ⮚ W. David K. (2017, July 27). Paleoenvironmental Science: Methods. African History. Retrieved November 11, 2021, from https://oxfordre.com/africanhistory/view/10.1093/acrefore/9780190277734.001.0 001/acrefore-9780190277734-e-211 ⮚ H.J.B. Birks and T. Webb (2008). Paleoecology. Paleoecology. Retrieved November 12, 2021, from https://www.sciencedirect.com/topics/agricultural-andbiological-sciences/paleoecology