1. Testes: the source of sperm • Usually paired • Usually abdominal • Exception: scrotal testes (some mammals) • Connected via paired sperm ducts to the outside • specialised regions: epididymis, vas deferens Evolution of the scrotum in mammals (Kleisner et al. 2010): 1) - Probably evolved twice blue & black lineages 2) Probably multiple pressures: - selection cooling hypothesis galloping hypothesis signalling hypothesis (testis by kidneys) (prepenial scrotum) One of two representations of the mammalian tree showing different states of testicular location (Kleisner et al. 2010, p 31) Testicular structure in mammals = interstitial cells (Norris 1997 p 373. Also in Norris 2007, p 337)