Fungi Evolution and Diversity Grade 10 Term 3 Fungi There are 100,000 species of fungi at the moment but mycologists (scientists who study fungi) think there are many more. Fungi are mostly multicellular eukaryotes. Fungi are saprotrophs – they send out enzymes that break down dead and decaying organic matter. Nutrients are then absorbed by the fungi. Figure 10.1 shows how the 6 groups of fungi are related. Microsporidia and chytrids are single-celled. Chytrids are aquatic and have flagellated spores and gametes. Evolution of Fungi Zygospore fungi, sac fungi and club fungi have similar structures. Protists evolved 1.5 billion years ago. Animals and plants are descendent from these but animals show more recent links. Animals and fungi from the super group Opisthokonta are more closely related to each other than to plants. It is thought that they share an aquatic, flagellated, single-celled ancestor. Multicellular forms came later after animals and fungi divided into different groups. Fungi don’t fossilize تتحجرwell so it is harder to date their evolution. The earliest fungi fossil found is 460 million years old but we think fungi may be older than that. Mycorrhizae are evident in plant fossils. Mycorrhizae are fungus that grow on plant roots. It is thought that fungi were so successful because of adaptive radiation when organisms began to colonise the land. Some fungi are single-celled. Example: yeast.خميرة Most are multicellular. Structure of Fungi The body of most fungi are known as mycelium (fungus filaments). The mycelium is made up of filaments called hyphae (web). This structure allows a high surface-to-volume ratio which maximizes the absorption of nutrients. Hyphae خيط فطريgrow from their tips. Some have septa حاجزwhich are walls that break the hyphae into cells. Fungi with septa are called septate. Pores in the septa allow cytoplasm and some organelles pass through from one cell to another. Septa that separate reproductive cells are nonporous.غير قابل لإلختراق Aseptate fungi الفطريات المطهرةare not divided into cells and have many nuclei in one hyphae. Some hyphae can اخترقpenetrate/break through rigid جامدsubstances like plant tissue. When a fungus reproduces, a specific part becomes a reproductive structure and the rest of the mycelium nourishes/feeds it. Fungal cells don’t have chloroplasts. They have a cell wall made of chitin. Chitin is made of glucose microfibrils and each glucose has a nitrogen-containing amino acid attached. Chitin is also found in insects and crustacea so it shows the link to animals. Fungi store energy as glycogen which is the same as animals. Except for chytrids, fungi are not motile as they lack basal bodies and flagella. They find food by growing hyphae – hyphae can grow up to 1 km a day. Fungi reproduce both sexually and asexually. Sexual reproduction of terrestrial نباتfungi occurs in 3 stages. Hyphae from 2 different mating fungi make contact and fuse. This can happen immediately or take months or years. Reproduction of Fungi The nuclei divide until every cell in the hyphae has one of each nuclei. A hyphae with paired haploid nuclei is called dikaryotic. The nucleus fuse and the zygote undergoes meiosis before spore formation. Fungi usually produce nonmotile spores that are windblown – this ensures the offspring are spread to new locations. Spores are produced during sexual and asexual reproduction. Spores are reproductive cells that develop into new organisms without needing to fuse with another reproductive cell. Large mushrooms can produce billions of spores in a few days. When the spore lands on a food source, it germinates and grows. Asexual reproduction usually occurs when spores are produced by a specialized part of the mycelium. It can also occur by fragmentation and by budding.