B othalia 29,1 (1 9 9 9 ) 207 ACKNOW LEDGEM ENTS The authors w ould like to thank the U niversity o f the Free State for their facilities and the Foundation for Research D evelopm ent for financial support. R EFERENCES B ASHAW , E.C & H A N N A , W 1990 A pom ictic reproduction In G P Chapm ann. Reproductive versatility in the grasses. 1 00-130. C am bridge U niversity Press, Cambridge B R O W N , W V & EMERY, W .H .P 1958 A pom ixis in the Gramineae: Panicoideae. American Journal o f Botany 45: 1 6 5 -2 5 2 . CA R N O Y , J B 1886. La C ytodierêse de 1’oeuf. Cellule 3: 1 -9 2 C O N N O R , H E. 1981. E volution o f the reproductive system s in the Gramineae. Annals o f the Missouri Botanical Garden 68: 4 8 -7 4 . DU PLE SSIS, H & SPIES, J.J 1992 C hrom osom e numbers in the g en u s Pen tasch istis (P o a cea e, D anth onieae). Taxon 41 7 0 6 -7 2 0 . G IB B S R U SSE L L , G .E., W ATSON. L , KOEKEM O ER. M , SM OOK. L„ B A R K ER , N.P., A N D E R S O N , H M. & DALLW ITZ, M J 1990. Grasses o f southern Africa. Memoirs o f the Botanical Survey o f South Africa No. 58: 1—437. G R E E N H A M , J. & C H A P M A N . G. P 1990 O vule structure and diversity In G .P Chapm an, Reproductive versatility in the grasses C am bridge U niversity Press. Cambridge H A N N A , W W & BA SH A W , E C. 1987 A pom ixis: its identification and use in plant breeding. Crop Science 13: 7 2 6 -7 2 8 JE N SE N . W.A 1962. Botanical histochemistry Freeman & Company, San Francisco JO H A N SE N , D A 1940. Plant microtechnique. M cG raw -H ill, N ew York. JO H N ST O N , G W 1953. O bservations on tw in embry o sacs in Sorghum vulgare. Phytomorphology 3: 3 1 3 -3 1 5 . KL.OPPER, K .C., SPIES, J.J & V ISSE R , B 1998 C ytogenetic studies in the genus Pentaschistis (P oaceae Arundinoideae) Bothalia 28: 2 3 1 -2 3 8 . L A B U SC H A G N E , I F 1990 'n Sitotaksonomiese ondersoek van die morfologiese ooreenstemmende spesies Pentaschistis airoides, P patula en P pallida (Poaceae: Arundineae). B .Sc. Hons the­ sis, U niversity o f the O range Free State LINDER. H P & ELLIS, R P. 1990. A revision o f Pentaschistis (A rundineae: P oaceae). Contributions from the Bolus Herbarium 12: 1-124. M OGIE, M. 1992. The evolution o f asexual reproduction in plants. Chapman & Hall, London N A R A Y A N , K .N 1951. Apomixis in Pennisetum . U npublished thesis, University o f California. N A R A Y A N A SW A M I, S. 1954. The structure and developm ent o f the caryopsis in som e Indian m illets. Ill Paspalum scrobiculatum L. Bulletin o f the Torrey Botanical Club 81: 2 8 8 -2 9 9 . N A R A Y A N A SW A M I. S 1955. The structure and developm ent o f the caryopsis in som e Indian m illets. IV Echinichloa frum entacea Link Phytomorphology 5 : 161-171 N O G LER. G A. 1984 G am etophytic apom ixis In B M. Johri, Em­ bryology o f Angiosperms. Springer-Verlag, Berlin. PH ILIPSON, M .N . 1978. A pom ixis in C ortaderiaju bata (Gram ineae). New Zealand Journal o f Botany 16: 4 5 -5 9 . PH ILIPSON, M .N. 1986. A reassessm ent o f the form o f reproduction in Danthonia spicata (L .) Beauv. New P hytologist 102: 2 3 1 -2 4 3 . S A SS . J.E 1951 Botanical microtechnique Iowa State C o lle g e Press. A m es. SPIES, J.J & D U PLE SSIS, H. 1986 The genus Rubus in South A frica III The occurrence o f apom ixis and sexuality South African Journal o f Botany 52: 2 2 6 -2 3 2 . ST E B B IN S, G.L. 1971 Chromosomal evolution in higher plants Edward Arnold, London T U R E SSO N , G 1929 Zur Natur und Begrenzung der Arteinheiten Hereditas 12: 3 2 3 -3 3 3 V E R B O O M . G A ., LINDER. H P & BA RK ER . R.P 1994 Haustorial synergids: an important character in the system atics o f danthonioid grasses (Arundinoideae: Poaceae). American Journal o f Botany 81 1601-1 6 1 0 . V ISSER . N .C . & SPIES, J.J. 1994 C ytogenetic studies in the genus Tribolium (P oaceae D anthonieae). IV. Section Uniolae. South African Journal o f Botany 60: 2 7 9 -2 8 4 . J.J S P IE S * t, K.C. KLOPPER* and B V ISSE R * * Department o f Botany and G enetics, U niversity o f the Orange Free State. P.O. B ox 339, 9 3 0 0 B loem fontein t To w hom correspondence should be addressed M S received 1998-04-08 PR O G R E SS W IT H TH E TR IA L PH A SE FOR R EG ISTRA TIO N O F N EW PLA N T N A M ES* $ In a spirit o f co-operation and to publicise this proposed innovation in plant nom enclature in southern A frica, this note has been subm itted sim ultaneously to the follow ing botanical journals: South African Journal o f Botany, Forum Botanicum and SABONET News. IN T R O D U C T IO N A num ber o f far-reaching proposals aim ed at the refine­ ment and sim plification of the International Code o f Botanical Nomenclature were discussed in some detail at the N om enclature Section o f the X Vth International B otanical C ongress (IB C ) held in A ugust 1993 in Yokohama, Japan. Three o f the main issues were the adop­ tion o f lists of Names in current use (Smith et al. 1993; Smith & H aw ksw orth 1994), the extended options o f con­ serving and rejecting nam es (Greuter & N icolson 1993), and the registration o f all new plant names (Borgen et al. 1997; G reuter & Von Raab-Straube 1998). This paper deals with the last-nam ed issue, namely the proposed sys­ tem by m eans o f which new nam es o f plants and fungi would have to be dealt with from 1 January 2(XX) onwards. A rticle 32.1 o f the International Code o f Botanical N omenclature adopted at the 1993 IBC (G reuter et al. 1994) c o n c lu d e s w ith ‘In addition, subject to the approval o f the XVI International B otanical C ongress, nam es (autonym s excepted) published on or after 1 January 2000 m ust be registered.’ Furtherm ore, A rticle 32.2 states that ‘Registration is effected by sending the printed m atter that includes the protologue(s) w ith the nam e(s) to be registered clearly identified, to any reg is­ tering office designated by the International A ssociation for Plant Taxonom y.' To dem onstrate the feasibility o f a registration system , the International A ssociation of Plant Taxonom y (IA PT) undertakes a non-m andatory tn al o f registration for a tw o-year period, starting 1 January 1998 (Borgen et al. 1997). D etails of the proposed im plem entation o f the registration requirem ent (Borgen et al. 1998) will be considered by the St Louis C ongress in July 1999. Since M ay 1998, a searchable dem onstration database co n tain ­ ing all nam es trial-registered after January 1998 can be freely consulted on the Internet (http://w w w .bgbm .fuberlin.de/registration/Q ueryForm .htm ). B othalia 29,1 (1999) 208 R EGISTRATION CENTR ES A s part o f the registration procedure o f new plant nam es it is im portant to establish a national Registration O ffice (RO ) in as m any countries as possible. This w ould enable editors and authors to register new plant nam es and com binations in their ow n or a neighbouring coun­ try. It is possible that the registration o f all new plant nam es w ould becom e m andatory in future, w ith effect from (but not before) 1 January 2000. The N ational Botanical Institute (NBI) was recently approached by the IA PT to serve as RO for South Africa. The Institute was contacted in view o f its permanence, national role, reliable com m unication system (telephone, fax, e-m ail, postal services) and perm anent filing and library facilities. The N BI provisionally accepted this invi­ tation on behalf o f all South A frican botanists. In the case o f new nam es and com binations for fungi, however, the authors have liaised w ith M s Alice Baxter o f the National Fungal C ollection, and after consultation with her staff she has agreed that the N BI act as RO for those nam es as well. A C C R E D IT E D JO U R N A L S/SE R IE S An enum eration o f 119 journals or series accredited with the IA P T appeared in Taxon 47: 498, 499 (1998). T his list is being regularly updated as new journals are added and is available on the W eb (http://w w w .bgbm .fuberlin.de/iapt/registration/journals.htm ). It will be pub­ lished annually in the journal Taxon. A fter consultation w ith som e o f the editors, it is pro­ posed that the follow ing jo u rn als should receive prefer­ ence w hen botanists w orking on the local flora establish new nam es for plant taxa: 1. Aloe (journal o f the S ucculent Society of South Africa). 2. Bothalia (biannual house journal o f the N ational Botanical Institute). 3. Contributions from the Bolus Herbarium (occasional publication o f the B olus H erbarium , U niversity of Cape Town). 4. Flora o f southern Africa (occasional publication o f the N ational B otanical Institute). 5. Flowering Plants o f Africa (biennial publication o f the N ational Botanical Institute). 6. Palaeoflora o f southern Africa (occasional publication p u b lish ed in the S trelitzia series o f the N ational Botanical Institute). 7. South African Journal o f Botany (tw o-m onthly journal jo in tly published by the N ational B otanical Institute and the South A frican A ssociation o f B otanists). 8. Strelitzia (occasional publication o f the N ational B otanical Institute). Editors o f other South A frican journals who w ould like their publication to be added to this list should co n ­ tact G.F. Sm ith as a m atter o f priority. It should furtherm ore be noted that proposals to restrict publication o f nam es for plant novelties in nonscientific publications will be voted on before or during the 1999 N om enclature Section o f the M issouri IBC (L aferriere 1998). REGISTRATION PR O C ED U R ES Steps to be taken by authors A uthors m ust ensure that nam es o f all new taxa and new com binations are registered and they can do it in one of the follow ing ways: 1. By sending their m anuscripts to an accredited journal (the issue o f accreditation is discussed below ). The ed i­ tor o f that journal will then see that registration takes place. 2. By subm itting a request w ith the relevant printed m at­ ter (in duplicate) and specifying w hich nam es are to be registered to a recording centre (the nearest RO). T his applies only to nam es not published in an accredited journal or serial, for exam ple in a book, m onograph or thesis. Use o f the apposite form s for registration requests is strongly recom m ended (although not m andatory). Registration form s are available from all Registration Offices. Steps are as follows: • R egistration form s will be m ailed upon request. A uthors who wish to use the RO in South A frica run by the NBI should send their requests to the address of the senior author o f this paper. • U pon receipt at the RO, each com pleted registration form (to be subm itted in triplicate) and the accom pa­ nying two copies or reprints o f the relevant article will be date-stam ped. By this act, the subm itted nam es will, technically speaking, becom e registered and dated (irrespective o f their subsequent processing). • Each form will be provided w ith the acronym o f the institution running the RO (PRE for the N ational H erbarium , Pretoria, in the case o f South A frica), and an appropriate NBI filing system num ber will be allo ­ cated. At NBI in Pretoria, the filing system will be m aintained in the M ary G unn Library under supervi­ sion o f the D irector: Research. • A copy o f the form will be returned to the subm itting author. • The R O ’s archival copy o f the registration form will be faxed to the office of the IA PT w ithout delay. • The original form along with one copy or reprint of the article will be sent by registered mail to the IAPT office w ithout delay. The second copy will rem ain in the RO archive perm anently, initially as an insurance against the risk o f postal loss. Steps to be taken by editors of accredited journals • For a journal to be accredited, its publishers must com m it them selves by signing an agreem ent w ith the IAPT. • To make the accredited status o f these journals known to authors and readers, it is suggested by the IA PT that the follow ing wording be used, appropriately placed on the cover and/or in the imprint: ‘A ccredited with the International A ssociation for Plant Taxonom y for the purpose o f registration of new nam es o f vascular plants (excluding fossils)’ or ‘A ccredited w ith the In­ ternational A ssociation for Plant Taxonom y for the purpose o f registration o f new non-fungal plant nam es’ or ‘A ccredited with the International A sso­ ciation for Plant Taxonom y for the purpose o f regis­ tration of all new plant nam es’. The last statem ent w ould also provide for new fungal and fossil names. Bothalia 29,1 (1 9 9 9 ) 209 • The new nam es and new com binations to be regis­ tered should be identified prim arily by being listed in a separate index in each issue or by being explicitly enum erated in the sum m ary/abstract or otherw ise specified by the editor of the journal. • Each individual issue featuring nam es of new taxa or new com binations m ust be subm itted to a pre-defined registration office or centre as soon as it is published and as speedily as possible. This will be Pretoria (PR E), for South A frica, and may be PRE for neigh­ bouring countries not having their own RO. ACKNOW LEDGEM ENTS The authors w ould like to thank M rs Em sie du Plessis and D r O tto Leistner, colleagues at the NBI, for their assistance in com piling this article. REFER EN C ES BO R G E N . L., G R E U TE R , W., HAW KSW O RTH, D L„ N IC O LSO N, D.H. & ZIM M ER , B 1997. A nnouncing a test and tnal phase for the registration o f new plant nam es (1 9 9 8 -1 9 9 9 ). Taxon 46: 8 1 1 -8 1 4 . BO R G EN. L., G R E U TE R , W„ HAW KSW O RTH, D .L .. N IC O L SO N , D.H. & ZIM M ER. B 1998. Proposals to im plem ent mandatory registration o f n ew nam es. Taxon 47: 8 9 9 -9 0 4 . GREUTER, W., BA RR IE. F.R., BU RD ET. H.M ., C H A L O N ER , W .G., D E M O U L IN , V., HAW KSW O RTH, D .L ., J 0R G E N S E N , P.M., N IC O L SO N , D .H ., SILVA, P C . & T R E H A N E , P. 1994. International Code o f Botanical Nomenclature (T okyo C ode). K oeltz, K oenigstein. GREUTER, W. & N IC O L SO N , D .H . 1993. On the threshold to a new nom enclature? Taxon 42: 9 2 5 -9 2 7 . GREU TER . W. & V O N R A A B -ST R A U B E , E. 1998. R egistration progress report, 1. Taxon 47: 4 9 7 -5 0 2 . L A F E R R Ê R E , J.E. 1998. (1 6 - 1 7 ) T w o proposals to restrict publica­ tion in non -scientific publications. Taxon 47: 181. SM ITH, G .F & HAW KSW O RTH, D .L. 1994. Stability o f scientific plant nam es— an attainable goal? South African Journal o f Science 90: 59, 60. SM ITH, G.F., ROURKE. J.P & OLIV ER . E G H 1993. Stability in botanical nom enclature. South African Journal o f Science 89: 313. G.F. SM ITH * and G. G E R M ISH U IZ EN * * National B otanical Institute, Private Bag X I 01, 0001 Pretoria. MS received: 1998-09-25.