Phylum Mollusca • soft bodied; many with protective shell • more species of molluscs in ocean than any other phylum, over 200,000 sp. • the mantle is a thin tissue layer covering body that secretes shell • the foot allows locomotion or attachment • most have a head with eyes • the radula is a tooth like structure used for feeding • Ancestral larvae is a trocophore; some produce a veliger • Open circulatory system • gastropods, bivalves, cephalopods, and chitons Larval Forms Trocophore Veliger Class Gastropoda - snails • “stomach foot” • most have coiled, spiral, shells (dextral or sinistral) • many use radula to scrape algae for food • predatory snails’ radula is modified to capture prey; spear or drill • also include nudibranchs and sea hares • Consist of the Subclasses Prosobranchia (aquatic, gilled snails), Opisthobranchia (non-shelled), Pulmonata (land, lung snails) Figure 7.19 Coiling vs. tortion Tulip Snail Figure 7.20 Nudibranchs (of the Subclass Opisthobranchia) Figure 7.21d • “naked gills” • no shell • often prey on cnidarian polyps and sponges • many are noxious tasting or use nematocysts from cnidarians for protection Class Bivalvia • • • • • • laterally compressed hinged shell no radula, no head; filter feed by siphoning water strong muscles close shell; scallops clams burrow in mud or sand Mussels (byssal threads) and oysters (cement) attach to surfaces • scallops live free; may move by flapping valves • largest is giant clam Figure 7.22a Figure 7.22c Figure 7.22d A pearl is produced when a grain of sand or other particle gets between delicate mantle membrane and shell. The sand grain is calcified and forms a pearl Class Cephalopoda • • • • • • • • “head foot” predatory, highly intelligent, very good vision shell internal or absent suckers on tentacles beak like radula use jet propulsion; siphon and funnel can change color (chromatophores) and/or release ink squid, octopus, cuttlefish, nautilus Figure 7.24a Figure 7.24b Figure 7.24c Figure 7.25a A squid measuring about 60 feet (18 meters) washed up on a Tasmanian beach over the weekend. (CNN.COM) Class Polyplacophora - chitons • Attach tightly to rocks using foot as suction cup • Feed on algae • Shell is made of hard plates