Halimeda is responsible for up to 90% of the total sand deposition on selected shores, that translates to roughly 1 meter height of sand every 500 yrs. After reproduction, the carbonate exoskeleton of Halimeda is gradually eroded either via wave action or bioerosion by species such as the irregular sea urchin Clypeaster rosaceus. Specimens from shallow turtle grass meadows in the Belize Barrier Reef have been found to forage primarily on Halimeda debris. Due to its preference for large particles which are ground to grain sizes <0.5 mm C. rosaceus plays an important role in bioerosion. Approximately 5.5 kg of coarse sediment is converted to silty fine sand by one individual per year A disease which caused a Caribbean-wide mortality in the reef dwelling sea urchin Diadema antillarum was also shown to indirectly affect coral reefs. The sea urchin functions as both a grazer on algae that can otherwise smother coral and as a bioeroder of corals as it feeds on them Dictyosphaeria Hypnea musciformis Cladophora One of the greatest threats to coral reefs is human expansion and development. As development continues to alter the landscape, the amount of freshwater runoff increases. This terriginous runoff may carry large amounts of sediment from land-clearing areas, high levels of nutrients from agricultural areas or septic systems, as well as many pollutants such as petroleum products or insecticides. silting Whether it is direct sedimentation onto the reef or an increase in the turbidity of the water due to eutrophication, decreases in the amounts of light reaching corals may cause bleaching. In addition, increases in the amounts of nutrients enhance the growth of other reef organisms such as sponges which may outcompete the corals for space on crowded reefs Coral bleaching is the whitening of coral colonies due to the loss of symbiotic zooxanthellae from the tissues of polyps. This loss exposes the white calcium carbonate skeletons of the coral colony. Corals naturally lose less than 0.1% of their zooxanthellae during processes of regulation and replacement. However, adverse changes in a coral's environment can cause an increase in the number of zooxanthellae lost. There are a number of stresses or environmental changes that may cause bleaching including disease, excess shade, increased levels of ultraviolet radiation, sedimentation, pollution, salinity changes, and increased temperatures Coral Diseases Several coral conditions have been identified as diseases: white band disease (WBD), black band disease (BBD), bacterial infection, and shut down reaction. They are also susceptible to tumors and parasitic worms. These maladies are all stress related, and anthropogenic stresses can increase a coral's susceptibility to these diseases. Diseases such as BBD and WBD actually kill coral tissue while advancing in a band around the coral and leaving the white coral skeleton behind. BBD, caused by the cyanophyte Phormidium corallyticum, may have a role in maintaining coral diversity because it is most prevalent in coral species that form large colonies and provide a structural framework for the reef. When BBD kills part of these colonies, the skeleton is available to be colonized by other coral species recruits. However, after 25 months, there were not coral recruits among corals infected by BBD. CLOD cyanide dynamite