Modified dorsal fins Caudal fin shapes A - rounded B - truncate C - emarginate D - forked E - lunate F - heterocercal G - abbreviate heterocercal H - homocercal I - isocercal Diphycercal Pectoral fin variation Pelvic fin variation Thunniform Compressiform Depressiform Anguilliform Filiform Taeniform Sagittiform Gl bif Globiform Placoid scales are covered with enamel and are found in chondrichthyans - vertebrate teeth are derived from placoid scales Ganoid scales have an enamel-like covering called ganoine i - found f d in i gars andd relatives Cosmoid scales are covered with cosmoine and are found in coelocanths Cyloid and ctenoid scales are also called elasmoid - They are thin thi andd flexible fl ibl andd are found in most bony fishes Fin rays have a left and right h lf andd usually half ll have h a segmental structure “True” True fin spines originate as a single bony element Fin rays and spines articulate with pterygiophores at their bases Fish skeleton can be divided in axial (skull and vertebral column) and appendicular pp ((fins)) Vertebral structure varies b between abdominal bd i l andd caudal region - has centrum and neural arch - has hemal arch in the tail and ribs (pleural, epipleural, epineural, epicentral) in the thorax Teleost skulls are complex - combination of bones that form the neurocranium (protection of the brain) and branchiocranium (the jaws, gills and supporting structures) Shark skull is entirely g cartilage neurocranium and branchiocranium Teleost neurocranium - a combination of bone types chondrocranium ((endochondral)) and dermatocranium ((dermal)) dermatocranium consists mostly of bones on the outer surfaces Endochondral bone originates as cartilage that is replaced by bone dermal bone does not originate as cartilage branchiocranium – consist of 5 arches – mandibular, palatine, hyoid, y , opercular p and branchial mandibular – form upper jaw original palatoquadrate – the original upper jaw of sharks is entirely inside the mouth of the teleost, teleost on the roof as the palatine the original lower jaw (Meckel’s cartilage) is small, and lies within the dentary new dermal bones form the functional upper jaws – premaxilla, maxilla, and supramaxilla - the th first fi t two t may bear teeth, but in more advanced fishes, only the premaxilla bears teeth Branchiocranium mandibular arch - Meckel Meckel’ss cartilage lies within the dentary palatine arch – consists of palatine bone on roof of mouth hyoid arch – (suspensorium) – attaches lower jaw and opercular apparatus to the skull - consists of the hyomandibula, the symplectic, and quadrate opercular apparatusbones that form the operculum – opercle, l subopercle, b l preopercle, interopercle, p hyoid complex – part of hyoid arch 5 bones that lie inside th lower the l jaw j andd opercular bones glossohyal supports the tongue branchial arch – four pairs of gill arches that support the gills basibranchials hypobranchials basibranchials, hypobranchials, ceratobranchials ceratobranchials, epibranchials epibranchials, pharyngobranchials, upper pharyngeal bones Found in most chondrichthyans and all actinopterygians Found in primitive sharks Found in sarcopterygians and tetrapods Appendicular skeleton Adaptive radiation in Lake Malawi cichlidae telencephalon – olfactory (I) diencephalon – pineal gland – sensitive to light, regulates diurnal cycling and homeostasis mesencephalon – vision (II) metencephalon t h l – muscular l movementt - with ith cerebellum b ll (III – IV) myelencephalon – integration of somatic and visceral functions –homeostasis (V-X) Cranial Nerves I - olfactory II - optic - retina III - oculomotor III, IV, VI - eye muscles IV - trochlear VII & VIII - lateral line on head VI - abduscens V - trigeminal - jaw and face VII - facial f i l VIII - acoustico IX - glossopharyngeal - gill region X - vagus - lateral line of body & viscera