INFORMATIONS DE LA SOC. BELGE DE MALACOLOGIE. SER. 1?. N°1. Janvier 19&L : 5 - 8
54401
FAVARTI A ( F A V A R T I A ) P A U L B O S C H I
(MURICIDAE
A
KATHLEEN
NEW
: MURICOPSINAE)
MUR I C I D
FROM
OMAN.
ROLAND
R. S M Y T H E
HOUART
St. Jobsstraat,
8
B-3330 Landen (Ezemaal)
Be I g i um.
6 B l o n d e l I Ori ve
Bognor Regis, Sussex,
P021 4BQ, U.K.
ABSTRACT.
Th is paper de s cr ib es a new species o f m u r i c i d , FavaAtui paulboschi, from Oman.
The s h e l l , operculum and ra du la are i l l u s t r a t e d . Th is spe cies was f i r s t n o t ic e d
in December 1980 on Mas i rah, Oman and s in c e then 19 l i v i n g and 28 dead examples
have been recorded from Mas i rah and t h e mai land near Muscat, Oman.
Note.
F o l lo w in g a b b r e v i a t i o n s are used in th e t e x t :
IRSNB
MNHN
AMNH
USNM
NMW
NHM
:
:
:
:
:
:
I n s t i t u t Royal des Sciences N a t u r e l l e s de B e lg i q u e , B r u x e l l e s .
Muséum Na ti ona l d ’ H i s t o i r e N a t u r e l l e , P a r i s .
American Museum o f Natural H i s t o r y , New York.
N a tio na l
Museum o f Natural H i s t o r y (Smithsonian I n s t i t u t i o n ) , Washington D.C
N a ti on al
Museum o f Wales.
Natur al H i s t o r y Museum, Oman.
Fava/utla [FavoAtla} p a u lb o sch i
Smythe & Hou ar t, sp. nov.
DESCRIPTION.
Shell medium size d f o r th e genus, t h e l a r g e s t specimen examined being 18,5 mm in
le ng ht , s o l i d and heavy.
covered w i t h a very t h i n
E x te rio r d ull white,
ivory,
l i g h t brown o r b r o w n is h - g r e y ,
in trita c a lx .
A p e rt u r e o v at e, the edges g lo s s y w h i t e , th e i n t e r i o r w h i t e , y e l l o w i s h o r brownish.
Co lu m el la r l i p smooth, com pl e te ly adherent t o t h e s h e l l .
Outer l i p b r o a d ly c r e n u l a t e , r e f l e c t i n g t h e s p i r a l
Anal notch not ap pa re nt .
sculptu re of the s h e l l .
smooth.
S¡phonal canal about t h e same length as t h e a p e r t u r e , n a r r o w l y open in a d u l t
In te rio r
specimens, more w i d e l y so in younger ani ma ls, s l i g h + l y
i n c l i n e d t o th e l e f t and
recurved d o r s a l l y a t t h e t i p .
S p ir e h i g h , c o n s i s t i n g o f two s t r o n g l y c a r í n a t e ,
heavely s c u l p t u r e d p o s t n u c l e a r w h o r l s .
f l a t n u c l e a r wh orl s and f i v e
The e a r l y wh orl s have s i x t o seven v a r ic e s
w h i l e th e body whorl has f i v e , o f which th e a p e r t u r a l one i s narrow and expanded.
There i s a small curved, s p i n e - l i k e ex te n s io n on t h e c a r i n a l
worn in some specimens, but no o t h e r a x i a l
The s p i r a l
part of the va rice s,
sculpture.
s c u l p t u r e on t h e f i r s t t h r e e wh orls c o n s i s t s o f two heavy r o p e - l i k e
t h r e a d s ; t h e r e are f i v e cords on th e body w h o r l , th e upper t h r e e being very s t r p n g
w h i l e t h e a n t e r i o r ones are weak.
These cords become c o a r s e r towards the v a r i c e s
g i v i n g the impression t h a t t h e r e are deep r e c t a n g u l a r p i t s between them.
The operculum is t h i n ,
pale h o r n - c o l o u r e d , w i t h a t e r m in a l nucl eus .
The r a d u la i s muri cop sin e w i t h a s h o r t c e n t r a l
cusp on th e r a c h i d i a n t o o t h which
is s t r o n g ; t h e l a t e r a l s are s le n d e r and hooked.
TYPE MATERIAL.
Holotype : ( P l a t e I I ,
Lenght
fig .
1) B r i t i s h Museum (Nat ur al H i s t o r y ) Reg.N° BM(NH)
13,25 mm, maximum w i d t h 6,5 mm,
1983062.
le nght o f a p e r t u r e i n c l u d i n g canal
6,75 mm. Operculum 2,95 mm x 1,55 mm. Leg. K.R. Smythe 16 December
1980
a t Dhuwwah, Mas i rah, Oman.
Paratype : ( P l a t e I I ,
fig .
2) Smythe c o l l e c t i o n .
6,25 mm. Leg. K.R.Smythe,
1 paratype I . R . S . N.B.
Length 12,45 mm. maximum w id th
16 December 1980, Dhuwwah.
n° : IG 26656/403.
1 paratype M.N.H.N., type c o l l e c t i o n .
1
paratype A.M.N.H. n° : AMNH 186117.
1
paratype N.M.W. n° : NMW:Z: 1983:0 52 .
1
par atype U.S.N.M. n° : USNM 792424.
1
par atype N.H.M., Oman.
23 paratypes K.R. Smythe c o l l e c t i o n .
2 paratypes
19 paratypes
R. Houart coi l e c t i o n .
Donald T. Bosch c o l l e c t i o n .
1 paratype T. Pain c o l l e c t i o n .
TYPE LOCALITY.
Beach near f r e s h water d i s t i l l a t i o n
58°52’ E.
Sludgey s i l t w i t h
in th e s i l t .
p l a n t a t Dhuwwah, Masirah, Oman : 20°39'N,
loose limestone rocks of v a r y i n g s iz e s p a r t i a l l y b u r ie d
Undersides o f th e rocks w i t h some algae and tun i c a t e s .
pauZboAcht i s uncommon, u s u a l l y s i n g l y under a r o c k .
FavcU itia
Other m ollus cs are UeKZta.
a Z b tc lZ Z a , PZanaxÁA AuZcatuA, Eu.che.ZuA aApeti, C h ito n ZamyZ and two small dor id s .
Al so l i v e were marine worms, b r i t t l e s t a r f i s h , crabs and s co rp io n f i s h .
DISTRIBUTION.
VoivaXbLa. pauZboóchl has been found by K.R. Smythe and by Dr. DonaldT. Bosch a l i v e
at Dhuwwah and Rass i e r . Mas i rah, under stones and a t Al Bastan and Qurm near Muscat
on th e mainland o f Oman.
I t has been found dead, u s u a l l y crabbed, a t most l o c a l i ­
t i e s w i t h rocks around the coast o f Mas i rah.
it
is rare.
It
It
i s not common,.and on t h e mainland
is not recorded under any name by Me!v i 11 o r Me Iv i 1 I and Standen
f o r t h e Arabian G u l f , Gu lf o f Oman and North Arabian Sea, nor by any o t h e r au t h o r
f o r th e area t h a i we have s t u d ie d (see Smythe 1976 and 1979 f o r r e f e r e n c e s ) , nor
are t h e r e specimens in the B r i t i s h Museum (Nat ur al H i s t o r y )
(Townsend, Biggs,
C h a ll en ge r and Sykes c o l l e c t i o n s e s p e c i a l l y ) , th e N a ti o n a l Museum o f Wales (Tomlin
and Mel v i l !
c o l l e c t i o n s ) , th e Royal S c o t t i s h Museum ( v a r i o u s c o l l e c t i o n s from the
Arabian Gu lf and Gu lf o f Oman), t h e Ferdowsi U n i v e r s i t y c o l l e c t i o n , Meshhed,
I ra n ,
nor t h e c o l l e c t i o n s of t h e l a t e P r o f e s s o r M. TadjaI I i -Pour in P a r i s and Iran
(K.R.Smythe personal communications and s ea rc he s ).
To d a t e i t appears t o be r e s t r i c ­
ted t o Masirah and the nearby mainland - one o f t h e a ut ho r s (K.R.Smythe) d i d not
fin d
it
in Dho fa r, Southern Oman nor in th e Un it ed Arab Emirates b o r d e r i n g on th e
G u lf of Oman.
ETYMOLOGY.
We have pl e a s u r e in naming t h i s species f o r Paul Bosch, son o f Donald and E l o l s e
Bosch, who has helped us a l l w i t h h i s e n t h u s i a s t i c a s s i s t a n c e in c o l l e c t i n g
mollusca and h i s g e o l o g i c a l
knowledge.
DISCUSSION.
Th is spe cies i s not describ ed nor i l l u s t r a t e d
D 'A ttilio
(1976).
in F a i r (1976) nor Radwin and
Two r e l a t e d s pec ie s are F a v a rtia . pe.cu>e¿ (Tryo n,
F a v a r tia ¿ y k t e i (P r e s to n ,
1904).
F. p&a&eÀ. has a s m a l l e r , rounded a p e r t u r e ,
h e a v ie r s p i r a l cords and rounded v a r i c e s .
r i g h t margin o v e r la p s th e i e f t ,
1880) and
The siphona! canal
i s s h o r t e r and t h e
which does not happen in F. p a u tb o A c k i; t h e l a t t e r
is a l s o a more s le n d e r sp ec ie s .
From F. b y k u i i t d i f f e r s by i t s much more s le n d e r form; a l s o F. ¿ ykt& i. has more
squamose and rounded v a r i c e s and a ve r y d i f f e r e n t pr ot oco nch , c o n s i s t i n g o f 3
rounded, g l o s s y w h i t e w h o rl s .
ACKNOWLEDGEMENTS.
We wish t o thank p a r t i c u l a r l y Dr. S le u r s and Dr. Wouters (IRSNB, Belgium)
mounting t h e r a d u la and f o r t h e scanning m ic ro gr aph s.
for
Our s p e c i a l thanks are a l s o
due t o .Dr. E.H. Vokes f o r her ad v i c e , Dr. P. Graham O l i v e r o f NMW and Dr. D, Heppell
of RSM, Edinburgh f o r checking t h e i r c o l l e c t i o n s f o r us, t o Dr. J.D. T a y l o r f o r
help and access t o th e B r i s t i s h Museum c o l l e c t i o n s and l i b r a r y , t o Mr. Tom Pain
f o r help and t o Dr. Donald T. Bosch and h i s f a m i l y f o r t a k i n g one o f th e autho rs
(K.R.Smythe) t o Masirah and o t h e r places and f o r h e l p i n g in the search f o r t h i s
m ol lu s c and donation of specimens.
BIBLIOGRAPHY.
F a i r , Ruth H.,
1976. The MuA£X Book : an i l l u s t r a t e d cat al o gu e o f Recent M u ri c id ae
( M u r ic i n a e , Muri cop si nae , Ocenebrinae).
Radwin, G.E. and D ' A t t i l i o ,
A.,
138 pages, 23 p l a t e s . Ho no lu lu .
1976. Mutex S h e l l s o f t h e World.
An I l l u s t r a t e d
Guide t o the M u r i c id a e . 284 pages, 32 p l a t e s . S t a n f o r d , C a l i f o r n i a .
Smythe, K.R.,
1975. On th e Occurence of
S a Z in a tO fi fatULQ-iJLii, (Lamarck) in th e
Arabian G u l f . J. Conch. Lond. 28: 335-338.
Smythe, K.R.,
1979. The Marine Mollusca o f t h e Un it ed Arab Emir at es, Arabian G u l f .
J. Conch. Lond. 30: 57-80.
RESUME.
Un nouveau M u ri c id a e (Muricopsinae)
: TavafctLa. [Favasutia.) pauZ boichi Smythe &
Houart s p . n . e s t d é c r i t e t comparé à deux espèces
ptLO&QÂ. (Tryon, 1880) e t Fa vcw tla
La l o c a l i t é t y pe e s t s i t u é e à Dhuwwah,
pl us ou moins proches
; FavaAtia,
( P r e s to n , 1904).
Me de Mas irah, Oman, par 20°30' N e t
58°52’ E.
L'espèce e s t nommée en l' h o n n e u r de Paul Bosch, f i l s du Dr. Don Bosch e t E l o i s e
Bosch.
PL. Il
1. FavoJvLLa. (F.) p a u lb o s c h i Smythe 8 Houart sp, nov. ho lo ty pe BM(NH). 13.25 mm
2. Fa va /u tia (F.) p a u lb o s c h i Smythe 8 Houart sp. nov. paratype Smythe c o l l .
3 - 3A.
Operculum
4. Protoconch
5. Scanning micrograph o f the r ad ul a (x 1250)
(12.45 mm)