Plant Association Group and Special Habitat Descriptions Deschutes and Ochoco National Forests, Crooked River National Grassland Prepared by Katie Grenier, Mike Simpson, Reid Schuller, and Robin Vora September 2010 Background Trees, shrubs and other plant life can be classified in many different ways and for many different purposes. Two classification systems that are commonly used by forest managers to assess vegetation patterns and their changes over time are “Plant Association Groups” (hereafter PAGs) and “Special Habitats.” PAGs are mapped on the Deschutes and Ochoco National Forest in the corporate Geographic Information System (GIS) database. Each PAG is comprised of five to ten closely related plant associations, or groupings of plants that occur together in similar environments. These environments are typically defined by their climates (temperature and moisture), soils, and history of natural disturbances, such as wildfires, diseases and insect outbreaks. Plant associations vary from tens to hundreds of acres in size. PAGs typically occur over hundreds to thousands of acres. Thus, PAGS provide a general picture of major vegetation patterns across a forest. PAGS may also be characterized by features other than vegetation, such as cinder, glacier, lava, meadow, rock and water. Together, the PAGs provide a picture of both the vegetation and the non-vegetative features within a large area like a National Forest. Special habitats represent features which occupy less than an acre up to tens of acres. These habitats are often overlooked when examining PAGs, but are important because they support unusual plant life or contain high biological diversity. For example, non-forest and riparian habitats are often treated as special habitats because they have a disproportionate value in conserving biological diversity (USDA Forest Service 1996), although they occupy only a small proportion of the land base. Similarly, special habitats often support rare plants, mosses, lichens or fungi. A brief description of the PAGs occurring on the Deschutes National Forest and the Ochoco National Forest (including the Crooked River National Grassland) is provided below. This is followed by a description of the Special Habitats occurring on both National Forests. Contents Deschutes National Forest - PAGs .................................................................................................. 2 Ochoco National Forest - PAGs ...................................................................................................... 6 Crooked River National Grassland – PAGs .................................................................................... 8 Special Habitats – Deschutes NF, Ochoco NF ................................................................................ 9 Acknowledgement ........................................................................................................................ 10 References ..................................................................................................................................... 10 Appendix 1. Common and scientific names ..................................................................................11 Page 1 of 11 Deschutes National Forest - PAGs The PAGs mapped on the Deschutes National Forest reflect a broad range of biophysical environments, from the alpine peaks of the Three Sisters which receive 145 inches of annual precipitation, to the juniper woodlands near Bend which averages 12 inches of annual precipitation. Vegetation on the Forests and Grassland varies with elevation and moisture gradients from west to east. On the Deschutes National Forest, elevation, precipitation, and distance from the crest of the Cascade Mountains greatly influence vegetation patterns. These patterns are also influenced by soils from relatively recent volcanic ash and pumice that are excessively well drained and cooler than adjacent areas (Simpson 2007). Table 1 lists the PAGs that are currently mapped on the Deschutes National Forest, and provides the acres and percentage of the total for each PAG. Many of the plant associations grouped to form PAGS are those described in Volland (1985). Since this mapping occurred, a new plant association guide has been completed (Simpson 2007); however, this information is not yet available in the corporate GIS (Geographic Information System) of the Forest Service. This report will be updated when that spatial data becomes available. Table 1. Total acres of mapped Plant Association Groups (PAGs) and their relative percentages occurring on the Deschutes National Forest. PAG Name Alpine Dry Alpine Meadow Alpine shrub Cinder Glacier Hardwood Juniper Woodlands Lava Lodgepole Pine Dry Lodgepole Pine Wet Meadow Mesic Shrub Mixed Conifer Dry Mixed Conifer Wet Mountain Hemlock Dry Ponderosa Pine Dry Ponderosa Pine Moist Riparian Rock Water Whitebark Pine Dry Xeric Shrublands Deschutes NF Acres 506 3,732 184 6,972 937 494 2,911 27,514 496,991 70,352 7,678 10,430 304,180 94,565 191,496 574,852 75,787 8,890 12,886 35,695 145 28,227 Deschutes NF % of Total 0.03 0.2 0.01 0.4 0.05 0.03 0.1 1.4 25 3.6 0.4 0.5 16 5 10 29 4 0.5 1 2 0.01 1 Page 2 of 11 Alpine Dry PAG –Comprising only 0.03% of the mapped PAGS, this type is characterized by high elevation lodgepole pine and whitebark pine. These stands occur on deep ash/pumice deposits that are excessively well-drained. Understory vegetation cover and species diversity are typically low. Herbaceous and shrub fuels are scattered, and litter fuels are scarce. This scarcity results in a natural fire barrier except under extreme fire weather conditions. Alpine Meadow PAG – These non-forest plant associations are dominated by sedges, grasses and forbs occurring at higher elevations in the Cascade Mountains. Soils are imperfectly drained early in the growing season, often remaining moist well into summer. Alpine Shrub PAG – These shrub-dominated plant associations occur at high elevations in the Cascade Mountains. Soils are either imperfectly-drained early in growing season or welldrained. Pink mountain heath (Phyllodoce empetriformis) is a characteristic species. Hardwood PAG – The hardwoods are dominated by broadleaf, deciduous trees and tall shrubs including quaking aspen (Populus tremuloides), cottonwood (Populus trichocarpa), mountain alder (Alnus incana), red osier dogwood (Cornus sericeus), and Engelmann spruce (Picea engelmanii) which occurs in cold air basins. Common snowberry (Symphoricarpos albus) is a conspicuous mid-sized shrub. Juniper Woodlands PAG – Western juniper (Juniperus occidentalis) PAGs occur at low elevations in the warmest and driest areas within the Forest. Antelope bitterbrush (Purshia tridentata), Idaho fescue (Festuca idahoensis) and bluebunch wheatgrass (Pseudoroegneria spicata), and Sandberg bluegrass (Poa secunda) are typical understory species. On the Deschutes National Forest, the Juniper Woodland PAG is only 0.1% of the total mapped PAG acres (Table 1). Downed, dead woody fuel loads are often light. Large downed fuels commonly accounts for 70 percent or more of the total downed woody fuel load. Stands are characterized by discontinuous patches of needle mat layers. The most abundant surface fuel is curing grass or shrubs such as bitterbrush, or sagebrush on more productive sites (Simpson et al. 1994). Occurrence of fire has been reported to be reduced since about 1900 by fire suppression, reduction of understory fuels by domestic grazing, and construction of roads which act as fire breaks (Burkhardt 1969), except for some areas that have never sustained frequent fires and have been occupied by junipers for centuries (e.g., Horse Ridge). Lodgepole Pine forests – In central Oregon, distribution of lodgepole Pine (Pinus contorta) is linked directly to ash/pumice deposits, mostly from Mt. Mazama (Simpson 2007). Lodgepole pine has the widest ecologic variation of all the conifers that occur in central Oregon. It often occupies areas that are either too wet or too dry for other conifers such as ponderosa pine (Pinus ponderosa), white fir (Abies concolor), grand fir (Abies grandis), Shasta red fir (Abies magnifica var. shastensis), or mountain hemlock (Tsuga mertensiana). These PAGs have a mixed severity fire regime, with a combination of low to high severity fires occurring across the Forest (Simpson 2007). Average fire return intervals have been reported from 47 to 63 years in the southern Oregon Cascades, and 60 to 350 years on the Fremont National Forest (Simpson 2007). The Deschutes National Forest has two lodgepole pine PAGs: Lodgepole Pine Dry PAG – Typical understory species may include pinemat manzanita (Arctostaphylos nevadensis), long-stolon sedge (Carex inops), Idaho fescue, and western needlegrass (Achnatherum occidentale). The scarcity of fine surface fuel limits the spread of fire except under extreme fire conditions (Agee 1993). Page 3 of 11 Lodgepole Pine Wetland PAG – Typical species may include western bog blueberry (Vaccinium occidentale), Douglas’ spirea (Spiraea douglasii), widefruit sedge (Carex eurycarpa), kinnikinnick (Arctostaphylos uva-ursi), and few-flowered spikerush (Eleocharis pauciflora). These sites are located within forested floodplains, margins of wet meadows, and shallow concave sub-irrigated drainages. Water tables are at, or above, the soil surface through the early and mid-season parts of the growing season (Simpson 2007). Fires are not limited by surface fuels in this PAG. Meadow PAG – Meadows are dominated by grasses, sedges, rushes, or forbs (USDA Forest Service 1988). Soils are either imperfectly-drained or saturated through most of the growing season. They are often intermixed with riparian areas. Mesic Shrub PAG – This area is dominated by moisture-loving plants. Willows (Salix spp.), mountain alder (Alnus incana), western bog blueberry, and Douglas spirea are typical shrubs. Aquatic sedge (Carex aquatilis), Sitka sedge (Carex sitchensis), beaked sedge (Carex utriculata), and tufted hairgrass (Deschampsia cespitosa) typically occur. Mixed conifer forests -These are comprised of a mix of forest species that may include white fir, Douglas-fir (Pseudotsuga menziesii), mountain hemlock and lodgepole pine. Western white pine, western larch (Larix occidentalis), and incense cedar (Calocedrus decurrens) may also occur. Ponderosa pine occurs throughout the mixed conifer PAG, and often is the dominant species at lower elevations along with white fir, sugar pine (Pinus lambertiana), and lodgepole pine. Two PAGs are included: Mixed Conifer Dry PAG – These PAGs have moderate to high productivity and a mean annual precipitation of 20 to 35 inches. Tree species consist of firs, ponderosa pine, Douglas-fir, larch, incense cedar, and lodgepole pine. This PAG occupies warm or cold dry environments. Soil moisture is low due to the occurrence of excessively drained soils. This PAG is associated with historically very frequent, low-intensity fires. Evidence from the Klamath-Siskiyou Mountains suggests that small, stand-replacing fires were not uncommon, but the most common fire type is a low severity underburn (Simpson 2007). Mixed Conifer Wet PAG – Plant associations in this PAG have a higher productivity than the Mixed Conifer Dry PAG and a mean annual precipitation of 35 to 75 inches. This PAG typically has herb-rich understory vegetation. Shrubs are highly variable but may include Cascade Oregongrape (Berberis nervosa), golden chinquapin (Chrysolepis chrysophylla), common prince’s pine (Chimaphila umbellate), dwarf rose (Rosa gymnocarpa), California blackberry (Rubus ursinus), and creeping snowberry (Symphoricarpos mollis). Douglas-fir is typically a significant component of overstory canopies. This PAG has the longest fire return interval and greater incidence of stand replacing fire within the mixed conifer forest. It is uncertain how fire exclusion has affected this PAG (Simpson 2007). Mountain Hemlock Dry PAG –This type, which occurs between 6000-7500’ elevation, ranges from dense, closed canopy stands of pure mountain hemlock to stands with several dominant tree species including Pacific silver fir (Abies amabilis) and lodgepole pine along with mountain hemlock. This PAG represents the cold, dry environments where effective soil moisture is low Page 4 of 11 and understory species diversity is low. Typical species include smooth woodrush (Luzula hitchcockii), grouse huckleberry (Vaccinium scoparium), big huckleberry (Vaccinium membranaceum), and beargrass (Xerophyllum tenax). Lodgepole pine is common after disturbance. Tree ages are usually old, suggesting that fire frequency is generally low. Wind and snow loading may be more important disturbance agents than fire. Upper slope and ridgetop locations favor lightning strikes, but fire spread is limited by moist fuels and the high incidence of precipitation along with fire strikes (Simpson 2007) Ponderosa pine forests are widely distributed on the lower east slope of the Oregon Cascades. Since ponderosa pine occupies drier sites than any other forest type except western juniper or occasionally lodgepole pine, its distribution is tied closely to available soil moisture. These PAGs represent the classic low-intensity, high-frequency fire regime that has been the most heavily impacted by fire suppression. This has resulted in in-filling by young trees and increased dominance of shrubs and concurrent loss of herbaceous species, as well as environmental changes such as increased duff depths, buildup of bark flakes and needles at tree bases. Fire suppression has contributed to development of more homogeneous stands which are more prone to stand-replacing fires (Hessburg et al. 2004). Where site restoration is the management goal, prescribed burning, used in combination with thinning and slash removal can be effective management techniques (Simpson 2007). Two PAGs are included: Ponderosa Pine Dry PAG – Twenty-nine percent of the Deschutes NF is mapped as belonging to this PAG. Typical species include antelope bitterbrush, Idaho fescue, western needlegrass, greenleaf manzanita (Arctostaphylos patula), and snowbrush (Ceanothus velutinous). This PAG occurs in zones that typically receive less than 20 inches of precipitation. Ponderosa Pine Moist PAG – Four percent of the PAGS on the Deschutes National Forest are mapped as this type. Average tree cover of 40-50% occurs in various mixtures of ponderosa pine and lodgepole pine. Quaking aspen may also be locally abundant. Shrub cover averages 20% and shrub layers lack diversity. Typical shrubs include antelope bitterbrush, greenleaf manzanita, kinnikinnick, bearberry (Arctostaphylos nevadensis), and wax currant (Ribes cereum). Riparian PAG – Riparian zones occur along the interface between aquatic and terrestrial ecosystems (Kovalichik 1987). Although great variation exists from high to low elevations, typical species include: willows, mountain alder, western bog blueberry, and Douglas spirea. Whitebark Pine PAG – Whitebark pine grows at the highest elevations in Oregon of any western tree species. Found on exposed ridges, temperatures are cool and the growing season is short. Winters are long, snowy and frequently involve severe windstorms. While the probability of increased lightning strikes increases on upper slopes and exposed ridges, late-lying snowpack, late persistence of moist fuels, and co-occurrence of rain along with lightning combine to limit the spread of many ignitions, except in prolonged drought periods. Xeric Shrublands PAG – These plant associations are dominated by big sagebrush (Artemisia tridentata) and antelope bitterbrush. Soils are well-drained throughout the growing season (USDA Forest Service 1988). Other typical species include Idaho fescue, bluebunch wheatgrass, and bottlebrush squirreltail (Elymus elymoides). Trees are absent or sparse. Surface fuels comprise the majority of the fuel bed. Page 5 of 11 Ochoco National Forest - PAGs Table 2 provides data on the 9 Plant Association Groups (PAGs) found on the Ochoco NF. The following general descriptions of each PAG are from Simpson et al. (1994) and his subsequent work. Table 2. Total acres of mapped Plant Association Groups (PAGs) and their relative percentages occurring on the Ochoco National Forest. PAG Douglas Fir Dry Grand Fir Dry Ponderosa Pine Juniper Steppe Juniper Woodland Mesic Ponderosa Pine Moist Grand Fir Shrubland Subalpine Fir Ochoco Acres 95,339 193,801 42,604 22,358 29,261 67,417 63,412 93,377 3,862 Ochoco % of Total 16 32 7 4 5 11 10 15 0.6 Douglas-Fir PAG – The Douglas-Fir PAG occurs at an average of 4,800 ft. elevation and aspect is generally south. It includes warm, dry Douglas-fir habitats types where ponderosa pine occurs as a major seral or climax associate. Many stands have a shrub-dominated undergrowth that may include common snowberry (Symphoricarpos albus), chokecherry (Prunus virginiana), antelope bitterbrush (Purshia tridentata), tall Oregongrape (Berberis repens), serviceberry (Amelanchier alnifolia), willow (Salix sp.), and/or mountain snowberry (Symphoricarpos oreophilus). Elk sedge (Carex geyeri) and pinegrass (Calamagrostis rubescens) dominate the herbaceous understory in many areas. Fuel loads are intermediate between the Ponderosa pine PAG and the Dry Grand fir PAG. Fuel loads tend to increase with stand age as a result of accumulated downfall from insect and disease damage, blowdown, and natural thinning. Live fuels can be a significant factor, especially where dense thickets of Douglas-fir develop during fire-free periods (Simpson et al. 1994). Dry Grand Fir PAG – The Dry Grand Fir PAG includes the most widely distributed and abundant plant associations on the Ochoco National Forest. Two plant associations make up this group: grand fir/pinegrass (Abies grandis/Calamagrostis rubescens) and grand fir/elk sedge (Abies grandis/Carex geyeri). These typically occur on south-facing slopes between 4,000-6,500 ft. elevations. Live fuels in the form of understory Douglas-fir and ponderosa pine can add greatly to the crown fire hazard, especially under severe burning conditions. Historically, relatively frequent surface fires maintained open stands dominated by fire-tolerant ponderosa pine and some Douglas-fir. Dry Ponderosa Pine PAG – This PAG is found mostly at lower elevations. As moisture increases at higher elevations in the Forest, the PAG occurs on shallow soils. Typical understory species are bluebunch wheatgrass (Pseudoroegneria spicata), Idaho fescue (Festuca idahoensis), big sagebrush (Artemisia tridentata), antelope bitterbrush, curlleaf mountain mahogany (Cercocarpus ledifolius), western needlegrass (Achnatherum occidentale), and arrowleaf balsamroot (Balsamorhiza sagittata). Downed, dead woody fuel loads are often light, but still contribute 70 percent or more to the woody fuel load. Stands are characterized by discontinuous patches of deep to very deep needle mat layers. The most abundant surface fuel is dead, or curing grass. Dwarf mistletoe adds considerably to fuel loadings in infested stands (Simpson et al. 2001). Before 1900, fire was a frequent event in ponderosa pine stands adjacent to western grasslands, occurring at intervals between 5 and 25 years. The natural role of fire in this PAG is threefold: (1) to maintain grasslands without western juniper or ponderosa pine; (2) to Page 6 of 11 maintain open stands (parklands) and remove dense understories of saplings, pole-sized trees, or (3) to thin overstory trees (Simpson et al. 1994). Juniper Steppe and Juniper Woodland PAGs – Western juniper (Juniperus occidentalis) PAGs occur at low elevations in the warmest and driest parts of the Forest (Simpson et al. 1994). Big sagebrush (Artemesia tridentata), Antelope bitterbrush, Idaho fescue and bluebunch wheatgrass are typical understory species. Downed, dead woody fuel loads are often light. Large downed fuels commonly accounts for 70 percent or more of the total downed woody fuel load. Stands are characterized by discontinuous patches of needle mat layers. The most abundant surface fuel is curing grass or shrubs such as bitterbrush, or sagebrush on more productive sites (Simpson et al. 1994). Occurrence of fire has been reported to be reduced since about 1900 by fire suppression, reduction of understory fuels by domestic grazing, and construction of roads which act as fire breaks (Burkhardt 1969), except for some areas that have never sustained frequent fires and have been occupied by junipers for centuries. The two PAGS differ primarily in the abundance of western juniper trees present and in soils. The Juniper Steppe PAG occurs on shallow soils. These sites are much less productive than the Juniper Woodland PAG sites. Typical shrub associates are low sagebrush (Artemesia arbuscula) or rigid sagebrush (Artemesia rigida). The lower productivity results in sparse vegetation cover and consequently Juniper Steppe PAG sites have much longer fire return intervals than Juniper Woodland PAG sites (80120 years versus 40-50 years). Mesic (moist) Ponderosa Pine PAG – The Mesic Ponderosa Pine PAG is more productive and has a different fire regime than the Dry Ponderosa PAG. Fuel conditions are variable, but are generally intermediate between the Dry Ponderosa PAG and the Dry Grand Fir PAG. Ponderosa pine is the principle tree species within the PAG, and pinegrass (Calamagrostis rubescens), elk sedge (Carex geyeri), and snowberry are the most prominent understory species present (Simpson et al. 1994). Moist Grand Fir PAG – In Moist Grand Fir PAG, typical understory species are queencup beadlily (Clintonia uniflora), twinflower (Linnaea borealis), huckleberries (Vaccinium spp.) and Columbia brome (Bromus vulgaris). The Moist Grand Fir PAG includes the most productive sites on the Forest, and occurs at all but the highest elevations. The fire regime within this PAG consists of two types: (1) a high frequency, low intensity regime which creates small openings in the canopy; and (2) a low frequency, high intensity regimes which results in stand replacement burns over thousands of acres. Fire return intervals are at least 40 to 50 years, and increase to 200+ years on the wettest sites. Live fuels in the form of understory trees can add greatly to the crown fire hazard, especially under severe burning conditions (Simpson et al. 1994). Subalpine Fir PAG – This PAG occupies only a small percentage of the total acreage on the Forest. It occurs at upper elevations with the coldest temperatures and most moisture on the Forest. In addition to subalpine fir (Abies lasiocarpa), Engelmann spruce (Picea engelmanii), lodgepole pine (Pinus contorta), and western larch (Larix occidentalis) occur. In addition, grand fir and Douglas-fir may occur in minor amounts. Typical understory species are grouse huckleberry (Vaccinium scoparium), big huckleberry (Vaccinium membranaceum), currants (Ribes spp.), heartleaf arnica (Arnica cordifolia) and elk sedge. Shrubland PAG – Fifteen percent of the Ochoco NF is mapped as non-forested. These areas are basalt tablelands or dry shrub-dominated areas that are dominated by one or more shrubs, including: low sagebrush, mountain sagebrush (A. tridentata var. vaseyana), and rigid sagebrush along with bluebunch wheatgrass, Sandberg bluegrass (Poa secunda), or Idaho fescue. Trees are absent or sparse. Surface fuels comprise the majority of the fuel bed. Page 7 of 11 Crooked River National Grassland – PAGs Table 3 provides data on the Plant Association Groups (PAGs) found on the Crooked River National Grassland (CRNG). A general description of each PAG follows Table 3; information is taken from Simpson et al. (1994), and subsequent work by Michael Simpson. Table 3. Total acres of mapped Plant Association Groups (PAGs) and relative percentages on the CRNG. PAG Dry Ponderosa Pine Juniper Steppe Juniper Woodland Shrubland CRNG Acres 1,739 8,735 97,156 4,765 CRNG % of Total 2 8 86 4 Dry Ponderosa Pine PAG – This PAG is found along the southwest fringe of the CRNG about 20 miles southwest of Madras. Typical understory species are bluebunch wheatgrass (Pseudoroegneria spicata), Idaho fescue (Festuca idahoensis), big sagebrush (Artemisia tridentata), antelope bitterbrush, curl-leaf mountain mahogany (Cercocarpus ledifolius), western needlegrass (Achnatherum occidentale), and arrowleaf balsamroot (Balsamorhiza sagittata). Downed, dead woody fuel loads are often light, but still contribute 70 percent or more to the woody fuel load. Stands are characterized by discontinuous patches of deep to very deep needle mat layers. The most abundant surface fuel is dead, or curing grass. Dwarf mistletoe adds considerably to fuel loadings in infested stands (Simpson et al. 1994). Before 1900, fire was a frequent event in ponderosa pine stands adjacent to western grasslands, occurring at intervals between 5 and 25 years. The natural role of fire in this PAG is threefold: (1) to maintain grasslands without western juniper or ponderosa pine; (2) to maintain open stands (parklands) and remove dense understories of saplings, pole-sized trees, or (3) to thin overstory trees (Simpson et al. 1994). Juniper Steppe and Juniper Woodland PAGs – Western juniper (Juniperus occidentalis) PAGs occur at low elevations in the warmest and driest parts of the Forest (Simpson et al. 1994). Big sagebrush (Artemesia tridentata), Antelope bitterbrush, Idaho fescue and bluebunch wheatgrass are typical understory species. Downed, dead woody fuel loads are often light. Large downed fuels commonly accounts for 70 percent or more of the total downed woody fuel load. Stands are characterized by discontinuous patches of needle mat layers. The most abundant surface fuel is curing grass or shrubs such as bitterbrush, or sagebrush on more productive sites (Simpson et al. 1994). Fire frequencies have declined since 1900. Occurrence of fire has been reported to be reduced since this time by fire suppression, reduction of understory fuels by domestic grazing, and construction of roads which act as fire breaks (Burkhardt 1969). Two PAGs occur on the CRNG. They differ primarily in the abundance of western juniper trees present and in soils. The Juniper Steppe PAG occurs on shallow soils. These sites are much less productive than the Juniper Woodland PAG sites. Typical shrub associates are low sage (Artemesia arbuscula) or rigid sage (Artemesia rigida). The lower productivity results in sparse vegetation cover and consequently Juniper Steppe PAG sites have much longer fire return intervals than Juniper Woodland PAG sites (80120 years versus 40-50 years). Shrublands PAG –These areas are basalt tablelands or dry shrub-dominated areas that are dominated by one or more shrubs, including: low sagebrush (Artemisia arbuscula), mountain sagebrush (A. tridentata var. vaseyana), and rigid sagebrush (A. rigida) along with bluebunch wheatgrass, Sandberg bluegrass (Poa secunda), or Idaho fescue. Fuel loads are comprised of surface fuels. Fire return intervals and fire behavior vary with site characteristics, but are generally comparable to that of the Juniper Steppe and Juniper Woodland PAGs, especially where juniper occurs at low density. Page 8 of 11 Special Habitats – Deschutes NF, Ochoco NF The following descriptions focus on the following special habitats: wetlands, riparian, meadows (wet, moist, and dry), hardwoods (aspen stands and cottonwood bottomlands), and scablands. Hardwoods – This type includes quaking aspen stands and black cottonwood (Populus trichocarpa) bottomlands. These deciduous hardwood trees provide a variety of important ecological functions, diversification of landscape patterns, and habitat for many other species (USDA Forest Service 1997). Riparian communities composed of these species are typically very rich in species and provide important structural diversity. Hydrologically, these habitats provide important ecosystem services including: stream bank and soil stabilization, flood resistance, and water temperature regulation. Soil fertility may be enhanced through accumulation of leaf-litter. Stream courses up to approximately 4,500 feet in elevation are often dominated by black cottonwood (Populus trichocharpa), a short-lived species (USDA Forest Service 1997). Black cottonwood stands often develop following flooding and deposition of sediment within floodplains and valleys. Regeneration may be by seed, stump sprouting, and rooted cutting. Quaking aspen (Populus tremuloides) grows in self-perpetuating clones in areas of locally high moisture such as meadows, seeps, and stream sides. Aspen and cottonwood are short-lived, early seral species that rely on disturbance for establishment. Across a broader geographic area, black cottonwood has been declining due to conversion of floodplain vegetation to agricultural uses, changes in stream systems (confining streams into a single channel), over browsing by livestock, elk and deer, and conifer encroachment (USDA Forest Service 1997). Quaking aspen is declining as a result of conifer encroachment and increasing competition, from over-browsing by livestock, deer and elk, and loss of habitat where the water table has dropped due to stream downcutting. Meadows (dry, moist and wet) – Meadow communities occur in areas where groundwater is higher than on adjacent upland sites, such as around a spring. Meadows often occur in the vicinity of a water table next to streams. Moist meadows have moisture available in the rootingzone during the growing season, which drops 1-3 feet in the fall. Wet meadows are supported by a water table at or near the surface throughout the growing season (Hall 1998). Dry meadows have no standing water and dry out completely in the summer. Riparian – Riparian areas include stream sides and adjacent upland areas. Plant communities are influenced by stream shape, steepness, and flooding. Plant life is typically diverse, and is characterized by a narrow band of moisture-loving plants directly adjacent to a stream, which includes sedges (Carex species), rushes (Juncus species), and bulrushes (Scirpus species). Riparian areas often receive moderate to heavy grazing use by wildlife and domestic livestock (where it occurs). Roads and trails passing through or parallel to riparian areas can affect the vegetation. Many roads are located along streams, contributing to direct loss or decline of habitat. Streamside environments are desirable locations for camping and water recreation, which can result in trampling of vegetation. Page 9 of 11 Scablands – These are open, rocky areas with shallow soils having a high stone and rock content. Trees are absent or sparse. Shrubs often include: low sagebrush (Artemisia arbuscula), or rigid sagebrush (Artemisia rigida). Low-growing grasses such as Sandberg bluegrass (Poa secunda) and herbs often predominate. Scablands are exposed to summer heat and drying of soils, winter snow, and soil saturation in the spring. Livestock grazing is a predominant land use within scabland habitats, although forage values are low. Wetlands – Wetlands have at least one of the following features: 1) presence of water loving plants (hydrophytes); 2) soils are wet throughout most or all of the summer (hydric soils); or 3) the rooting substrate is non-soil and is either saturated or covered by shallow water at some time during the growing season (Kovalchik 1987). Wetlands include: Ponds – this habitat is a body of standing water and its associated edge of water-tolerant vegetation. The pond may be permanent or seasonal. Bogs and fens – A bog is a permanently wet area of rain and snow-deposited moisture that is often dominated by sphagnum and other acid-tolerant plants. There is no outlet for bog water, which often saturates the soil and slows decomposition of plant material. A fen is fed by rainwater, snow melt, and underground springs which flow through a site, providing oxygen to the soil and enhancing the decomposition of living and dead plant materials. Both bogs and fens support a high number of plants, mosses, and lichens. Swamps – A swamp is a permanently wet area, often shaded by a canopy of tall shrubs or trees. Acknowledgement: Thanks to Emily Heyerdahl for review. References Burkhardt, J.W.; Tisdale, E.W. 1969. Nature and successional status of western juniper vegetation in Idaho. J. Range Mgmt. 22:264-270. Hall, F.C. 1998. Pacific Northwest Ecoclass Codes for Seral and Potential Natural Communities. USDA Forest Service Pacific Northwest Research Station, General Technical Report PNW-GTR-418. Kovalchik, B.L. 1987. Riparian zone associations, Deschutes, Ochoco, Fremont, and Winema National Forests. USDA Forest Service, Pacific Northwest Region, Region 6 Ecology Technical Paper 279-87. Simpson, M. 2007. Forested plant associations of the Oregon East Cascades. Tech. Pap. R6NR-ECOL-TP-03-2007. USDA Forest Service, Pacific Northwest Region. Simpson, M.; Zalunardo, D.; Eglitis, A.; Wood, D.C.; Roy, D.; Johnson, S. 1994. Viable Ecosystems Management Guide (draft). Ochoco National Forest, Prineville, Oregon. 108 p. + appendices. USDA Forest Service. 1996. Metolius Watershed Analysis. Sisters Ranger District, Deschutes National Forest, Sisters, Oregon. USDA Forest Service. 1997. Mill Creek Watershed Analysis. Ochoco National Forest, Prineville, Oregon. Volland, L. A. 1985. Plant associations of the central Oregon pumice zone. U.S. Forest Service Publication R6-ECOL-104-1985. Page 10 of 11 Appendix 1. Common and scientific names Common name antelope bitterbrush aquatic sedge arrowleaf balsamroot beaked sedge bearberry beargrass big huckleberry big sagebrush black cottonwood bluebunch wheatgrass bottlebrush squirreltail California blackberry Cascade Oregongrape chokecherry Columbia brome common prince’s pine common snowberry creeping snowberry curl-leaf mountain mahogany currants Douglas’ spirea Douglas-fir dwarf rose Engelmann spruce few-flowered spikerush Geyer's sedge golden chinquapin grand fir greenleaf manzanita grouse huckleberry heartleaf arnica huckleberries Idaho fescue incense cedar kinnikinnick Scientific name Purshia tridentata Carex aquatilis Balsamorhiza sagittata Carex utriculata Arctostaphylos nevadensis Xerophyllum tenax Vaccinium membranaceum Artemisia tridentata Populus trichocarpa Pseudoroegneria spicata Elymus elymoides Rubus ursinus Berberis nervosa Prunus virginiana Bromus vulgaris Chimaphila umbellata Symphoricarpos albus Symphoricarpos mollis Cercocarpus ledifolius Ribes spp. Common name lodgepole Pine long-stolon sedge low sagebrush mountain alder mountain alder Scientific name Pinus contorta Carex inops Artemisia arbuscula Alnus incana Alnus incana mountain hemlock mountain sagebrush Tsuga mertensiana Artemisia tridentata var. vaseyana mountain snowberry Pacific silver fir pinegrass Symphoricarpos oreophilus Abies amabilis Calamagrostis rubescens pinemat manzanita pink mountain heath ponderosa pine quaking aspen queencup beadlily red osier dogwood rigid sagebrush Sandberg bluegrass serviceberry Shasta red fir Spiraea douglasii Pseudotsuga menziesii Rosa gymnocarpa Picea engelmanii Eleocharis pauciflora Carex geyeri Chrysolepis chrysophylla Abies grandis Arctostaphylos patula Vaccinium scoparium Arnica cordifolia Vaccinium spp. Festuca idahoensis Calocedrus decurrens Arctostaphylos uvaursi Sitka sedge Smooth woodrush Arctostaphylos nevadensis Phyllodoce empetriformis Pinus ponderosa Populus tremuloides Clintonia uniflora Cornus sericeus Artemisia rigida Poa secunda Amelanchier alnifolia Abies magnifica var. shastensis Carex sitchensis Luzula hitchcockii snowbrush sugar pine tall Oregongrape tufted hairgrass twinflower Ceanothus velutinous Pinus lambertiana Berberis repens Deschampsia cespitosa Linnaea borealis wax currant western bog blueberry Western juniper western larch western needlegrass white fir widefruit sedge willows Ribes cereum Vaccinium occidentale Juniperus occidentalis Larix occidentalis Achnatherum occidentale Abies concolor Carex eurycarpa Salix spp. Page 11 of 11