2409.26f,900 Page 1 of 16 FSH 2409.26f - SEED HANDBOOK R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 CHAPTER 900 - TERMINOLOGY Angiosperm: Member of a group of plants having seed which develop in an ovary. Most common broad-leaved trees are angiosperms. Asexual Reproduction: Reproduction by purely vegetative means accomplished in woody plants usually by rooting stem or root cuttings, air layering, grafting or budding. Certified Seed: Seed attested by a recognized certifying agency to be from trees, shrubs or other plants of proven genetic superiority, as defined by that agency, and produced so as to insure genetic identity. Clone: Group of genetically identical plants produced by vegetatively propagating a single plant over one or more vegetative generations; accomplished in woody plants by rooting of cuttings, budding, grafting, or air layering. Cone: One of the reproductive structures of conifers. A female cone consists of a central axis supporting overlapping bracts (modified leaves) each of which underlies a scale bearing naked seeds. A male cone consists of a central axis supporting spirally arranged microsporophylls each of which bares pollen sacs containing pollen grains. Syn. strobilus. Conelet: Immature female cone of gymnosperms, sometimes described as a flower. Cotyledons: First leaves developed in the embryo of a seed. They may contain stored food for initial growth of the seedling or they may become functional leaves after germination. Also known as seed-leaves. Dioecious (two households): Having staminate (male) flowers and pistillate (female) flowers borne on different individual plants. Most Juniperus and many deciduous trees such as cottonwoods, willows, ashes, and maples are dioecious. Embryo: A small rudimentary plant contained within a seed resulting from the union of male and female gametes (fertilization). Embryo Cavity: The hollow portion in the endosperm which holds the embryo. Generally, a mature embryo completely fills the full length of the embryo cavity. Endosperm: Storage tissue surrounding the embryo in a seed and consisting of thin-walled cells rich in carbohydrates. Fertilization: Union of nucleus and other cellular constituents of a male gamete (sperm) with those of a female gamete (egg) to form a zygote (fertilized egg). In the pines fertilization occurs about 12-13 months after pollination, while in the R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 2 of 16 spruces, Douglas-fir, and true firs fertilization occurs within 1-2 months after pollination. Flower: Angiosperm reproductive structure bearing pistils, stamens, or both, and usually also sepals and petals. In conifers, the so-called "flowers" are the male and female strobili before and during pollination. Gametes: Mature sex cells, either sperm or eggs. Gene: Hereditary unit occupying a fixed position on a chromosome which through transcription has a specific effect on the phenotype. Genetic Gain: Average improvement among progeny over the means for the parents with respect to the characteristics used in selecting the parents. Genotype: 1. An individual's hereditary constitution, with or without phenotypic expression of the one or more characteristics it underlies. The genotype is determined chiefly from performance of progeny and other relatives. It interacts with the environment to produce the phenotype. 2. Individual(s) characterized by a certain genic constitution. Germination: Resumption of active growth in the embryo of a seed as demonstrated by the protrusion of the radicle. Geographic Variation: Phenotypic differences among native trees growing in different portions of a species range. Gymnosperm (naked seed): Member of a group of plants having naked seed (seeds not enclosed in an ovary) that are usually borne on the scales of a cone. Monoecious (single household): Having staminate (male) flowers and pistillate (female) flowers borne, usually in different locations, on the same plant. Most conifers and many deciduous trees such as walnuts, birches, and oaks are monoecious. Ovary: In angiosperms, the basal portion of a pistil within which seeds develop. Ovule (egg): A rounded outgrowth of the ovary in seed plants that develop into a seed usually after fertilization. Peduncle: The stalk or stem that bears a flower or a flower cluster. In conifers it is the stem which connects a female cone to the branch. Phenology: Study of relationships between seasonal climatic changes and periodic biological phenomena as the flowering and fruiting of plants, leafing, growth flushing and dormancy. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 3 of 16 Phenotype: All characteristics of a plant; morphological, anatomical and physiological; as we see it. It is the product of the interaction of the genetic makeup (genotype) of an organism and the environment. Pollen Grains: The microspores and young male gametophytes of seed plants. Pollination: The transfer of pollen grains from a stamen or microsporophyll to a stigma or ovule. In conifers, pollen is transported from the male reproductive structures to the female reproductive structures by the wind (wind pollination). Many angiosperms are pollinated by insects (insect pollination). Progeny Test: Evaluation of selected individuals (parents) by comparing performance of their offspring (progeny). Seed Collection Zone: Area having defined boundaries and altitudinal limits within which soil and climate are sufficiently uniform to indicate high probability for maintaining a single subdivision of plants that are adapted to a specific set of environmental conditions. Seed Lot: An indefinite quantity of seed having uniform quality, produced at a specific location, or within a single seed collection zone, and collected from a single annual crop. Seed Orchard: A plantation consisting of clones or seedlings from phenotypically superior parents established and managed for early and abundant seed production. They are usually isolated to reduce pollination from outside sources and rogues of undesirable individuals. Rogueing may proceed after testing of individuals and/or on the basis of phenotypic appearance. They are protected against insect and disease and may be treated with chemicals such as fertilizers and gibberellins to increase seed production. Seed Production Area: Stand of better-than-average phenotypes upgraded further by periodic removal of less desirable trees and cultured to increase seed production. Seed Source: Geographic location where a seed lot was collected; also the seed itself. Serotinous: Pertaining to cones that remain closed on the tree for several months or a year or more and are, therefore, late in dispensing seed. Source-Identified Seed: Seed attested by a recognized certifying agency as being from the specified seed source. Strobilus: The "cone" of conifers and certain other plants. The strobili contain the reproductive organs of one or both sexes. Before and during pollination time, they are often called "flowers". Between the time of pollination and fertilization (about 12 months) they are referred to as conelets. Male reproductive structures (pollen producers) are called male strobili (strobilus) and female reproductive structures (develop into cones) are called female strobili. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 4 of 16 Superior: A nontechnical term referring to selections which appear or have been proved to be outstanding. Variety: Subdivision of a species, usually separated geographically from the typical, having one or more heritable, morphological characteristics which differ from the typical even when grown under the same environmental conditions; a morphological variant. Viable: Capacity of a seed to germinate. EXHIBIT 1-12 (all are hard copy exhibits) SEED ZONES 1-1 Bighorn NF 1-2 Black Hills NF 1-3 Grand Mesa, Uncompahgre and Gunnison NF's 1-4 Medicine Bow NF 1-5 Nebraska NF 1-6 Rio Grande NF 1-7 Arapaho and Roosevelt NF's 1-8 Routt NF 1-9 Pike and San Isabel NF's 1-10 San Juan NF 1-11 Shoshone NF 1-12 White River NF R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 5 of 16 900 - Exhibit 01 BIGHORN NF SEED ZONES SEE THE PAPER COPY OF THE MASTER SET FOR SECTION 900 - EXHIBIT 01. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 6 of 16 900 - Exhibit 02 BLACK HILLS NF SEED ZONES SEE THE PAPER COPY OF THE MASTER SET FOR SECTION 900 - EXHIBIT 02. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 7 of 16 900 - Exhibit 03 GRAND MESA, UNCOMPAHGRE AND GUNNISON NF'S SEED ZONES SEE THE PAPER COPY OF THE MASTER SET FOR SECTION 900 - EXHIBIT 03. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 8 of 16 900 - Exhibit 04 MEDICINE BOW NF SEED ZONES SEE THE PAPER COPY OF THE MASTER SET FOR SECTION 900 - EXHIBIT 04. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 9 of 16 900 - Exhibit 05 NEBRASKA NF SEED ZONES SEE THE PAPER COPY OF THE MASTER SET FOR SECTION 900 - EXHIBIT 05. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 10 of 16 900 - Exhibit 06 RIO GRANDE NF SEED ZONES SEE THE PAPER COPY OF THE MASTER SET FOR SECTION 900 - EXHIBIT 06. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 11 of 16 900 - Exhibit 07 ARAPAHO AND ROOSEVELT NF'S SEED ZONES SEE THE PAPER COPY OF THE MASTER SET FOR SECTION 900 - EXHIBIT 07. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 12 of 16 900 - Exhibit 08 ROUTT NF SEED ZONES SEE THE PAPER COPY OF THE MASTER SET FOR SECTION 900 - EXHIBIT 08. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 13 of 16 900 - Exhibit 09 PIKE AND SAN ISABLE NF'S SEED ZONES SEE THE PAPER COPY OF THE MASTER SET FOR SECTION 900 - EXHIBIT 09. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 14 of 16 900 - Exhibit 10 SAN JUAN NF SEED ZONES SEE THE PAPER COPY OF THE MASTER SET FOR SECTION 900 - EXHIBIT 10. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 15 of 16 900 - Exhibit 11 SHOSHONE NF SEED ZONES SEE THE PAPER COPY OF THE MASTER SET FOR SECTION 900 - EXHIBIT 11. R2 AMENDMENT 2409.26f-93-1 EFFECTIVE 6/15/93 2409.26f,900 Page 16 of 16 900 - Exhibit 12 WHITE RIVER NF SEED ZONES SEE THE PAPER COPY OF THE MASTER SET FOR SECTION 900 - EXHIBIT 12.