This PDF describes Biophysical Setting (BpS) models in the mapping zone labeled below. BpS descriptions are listed alphabetically on the left-hand margin. You can use the Adobe “find” function (ctrl+f) to locate a BpS by numerical code or alpha string. LANDFIRE Mapping Zones of the United States Mapping Zone 73 LANDFIRE Biophysical Setting Model Biophysical Setting 7316011 Western North American Boreal Treeline White Spruce Woodland - Boreal This BPS is lumped with: WNA Boreal Spruce-Lichen Woodland (in part) This BPS is split into multiple models: Western North American Boreal Treeline White Spruce Woodland was split into a boreal and sub-boreal variant for BpS modeling so that regional differences could be represented. Boreal Spruce Lichen Woodland may occur as a seral stage or variant of Boreal Treeline White Spruce Woodland, Boreal Mesic Black Spruce Forest, or, less commonly, in these same systems in the sub-boreal region. General Information Contributors (also see the Comments field Modeler 1 Tricia Wurtz twurtz@fs.fed.us 4/8/2008 Reviewer Reviewer Reviewer Modeler 2 Modeler 3 Map Zone 73 Vegetation Type Forest and Woodland Dominant Species* PIGL PIMA BENA VAUL Date General Model Sources LEDUM ALVI5 SAPU15 BEPA Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This type is found in boreal AK from the south slopes of the Brooks Range to the north slopes of the Alaska Range and west to the limit of tree growth. Boreal Spruce-Lichen Woodland, which is included as a variant of a later seral stage within this type, occurs primarily in the northern and western portions of the boreal zone. Biophysical Site Description Boreal White Spruce Woodland occurs primarily near the elevational and latitudinal limit of tree growth. It occurs above Boreal White Spruce to 900m (Boggs et al. 2001) and below the subalpine shrub and tundra systems and can be seen as the forested transition zone between boreal white spruce forest and nonforested subalpine vegetation. Depending on the topography, this system can occupy a narrow band just below non-forested subalpine or a broad expanse across gentle slopes and benches. Soils are cold, but peatforming mosses are not common in the ground layer. Vegetation Description This type is dominated by Picea glauca although Picea mariana may be codominant (NatureServe 2008). Canopy cover is generally between 10-25%. Hardwoods, if present, can include Betula papyrifera and Populs tremuloides. The shrub layer typically features Betula nana, but other shrubs such as Vaccinium uliginosum, Ledum groenlandicum and Salix pulchra may be common or dominant (NatureServe 2008). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 1 of 204 In some locations Alnus viridis is the dominant understory shrub (NatureServe 2008). Feathermoss may be common in the in the ground layer (NatureServe 2008). On drier or more exposed sites, Cladina spp.replace feathermosses as the dominant ground cover (Viereck 1979). Disturbance Description This information was taken from the draft Boreal Ecological Systems description (NatureServe 2008), which notes that fire information is lacking: Fire return interval is in the vicinity of 100yrs shorter than for Western North American Boreal Treeline White Spruce Woodland - Alaska Sub-boreal. A possible scenario for post-fire succession in this type is the resprouting of low shrubs from underground propagules followed by Picea glauca invading by seed from adjacent stands or surviving trees. Betula papyrifera may invade the site if a seed source is available and site conditions are favorable. The typical succession sequence for this type does not include a hardwood sere. The rate of succession depends on severity of fire and seed source, and some sites may be shrub-dominated for long periods without spruce invasion. The successional relationships of the Boreal Spruce-Lichen Woodland included here are poorly understood. This system has been included in this model as a variant of the late successional stage (class C). Adjacency or Identification Concerns This system is similar to Western North American Boreal Treeline White Spruce Woodland - Alaska Subboreal but does not include an insects and disease probability and has a more frequent MFRI. Native Uncharacteristic Conditions Scale Description Large patch. Issues/Problems Experts at the LANDFIRE Fairbanks modeling meeting (Nov. 07) indicated the need for a late seral woodland white spruce-lichen class that could develop after approximately 200yrs in the absence of disturbance. This class was not included as a separate class because it was felt that LANDFIRE would not be able to distinguish it for mapping. The woodland white spruce-lichen concept is included in class C in the current model. Comments This model was based on input from the experts who attended the LANDFIRE Fairbanks modeling meeting (Nov. 07) and refined by Tricia Wurtz. Vegetation Classes *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 2 of 204 Class A Structure Data (for upper layer lifeform) Min Max 10 % Early Development 1 All Structures Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position BENA VAUL LEGR SAPU15 Upper Upper Upper Upper Dwarf Shrub (< 20 cm) Tree Size Class Tall Shrub (>1.5 m) Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Description 0-24yrs This class is characterized by herbaceous and shrub vegetation. Shrubs resprout from underground propagules and then Picea glauca invades by seed from adjacent stands or surviving trees. The shrub layer typically features Betula nana, but other shrubs such as Vaccinium uliginosum, Ledum groenlandicum and Salix pulchra may be common or dominant. In some locations Alnus viridis is the dominant understory shrub. Feathermoss may be common in the in the ground layer. On drier or more exposed sites, Cladina spp. replace feathermosses as the dominant ground cover (Viereck 1979). The rate of succession depends on severity of fire and seed source, and some sites may be shrub-dominated for long periods without spruce invasion. The main successional trajectory is to class C (white spruce woodland) but an alternate succession pathway to class B (probability = 0.04) represents sites where hardwoods may invade the site if a seed source is available and site conditions are favorable. Replacement MFRI = 150yrs. Class B Structure Data (for upper layer lifeform) Min Max 15 % Mid Development 1 Open Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position BEPA POTR5 PIGL PIMA Upper Upper Upper Upper Woodland (10-24% tree cover) Closed (60-100% tree cover) Dwarf Tree (< 3 m) Tree Size Class Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. Presence of hardwoods distinguishes class B from class C. Description 25-69yrs This class is characterized by hardwood or white spruce-hardwood mixed forest. Betula papyrifera and/or Populs tremuloides invade with or without spruce and gain canopy dominance over the shrubs. Forest canopy cover is generally 10-25%. Eventually hardwoods senesce and white spruce gains canopy dominance (class C). Succession to class C. Replacement MFRI = 150yrs. Mixed fire (MFRI = 300yrs) maintains this class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 3 of 204 Class C 75 % Late Development 1 Open Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position PIGL PIMA BENA CLADI3 Cover Height Structure Data (for upper layer lifeform) Min Max Woodland (10-24% tree cover) Closed (60-100% tree cover) Dwarf Tree (< 3 m) Tree Size Class Upper Upper Low-Mid Lower Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Absence of hardwoods distinguishes class C from class B. Description 25yrs+ This class is characterized by open white spruce woodland. Forest canopy cover is generally 10-25%. Hardwoods, if previously present in the stand, lose dominance in overstory during this phase. The understory may include various combinations of low shrubs, herbs and mosses. Lichens grow as the stand ages. This class includes the Spruce-Lichen Woodland system on some sites. The dominant lichen genus is typically Cladina; species include C. arbuscula, C. mitis, C. rangiferina and C. stellaris. Other lichens may include Cetraria cucullata, C. islandica, C. nivalis, Bryoria spp., Alectoria nigricans and Alectoria ochroleuca. Beetles affect this class but because the rate of outbreaks is uncertain and generally not severe enough to cause a state transition, they were not included in the VDDT model. The frequency and severity of beetle outbreaks will likely increase under a warmer climate. This class persists in the absence of disturbance. Replacement MFRI = 150yrs. Mixed fire (MFRI = 300yrs) maintains this class. Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 4 of 204 Disturbances Fire Regime Group**: Fire Intervals IV Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Avg FI Min FI Max FI Probability 150.2 333.0 0.00666 0.00300 104 0.00967 Percent of All Fires 69 31 Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Boggs, K., A. Garibaldi, J. Stevens, J. Grunblatt and T. Helt. 2001. Denali National Park and Preserve Landcover mapping project. Volume 2: Landcover classes and plant associations. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage, 707 A Street, Anchorage, AK. 164 pp. Foote, J.M. 1983. Classification, description, and dynamics of plant communities after fire in the taiga of interior Alaska. Res. Pap. PNW-307. Portland, OR. USDA Forest Service. Pacific Northwest Research Station. 108 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck, L.A. 1979. Characteristics of treeline plant communities in Alaska. Holarctic Ecology. 2: 228-238. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 5 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316012 Western North American Boreal Treeline White Spruce Woodland - Alaska Sub-boreal This BPS is lumped with: WNA Boreal Spruce-Lichen Woodland (in part) This BPS is split into multiple models: Western North American Boreal Treeline White Spruce Woodland was split into a boreal and sub-boreal variant for BpS modeling so that regional differences could be represented. Boreal Spruce Lichen Woodland may occur as a seral stage or variant of Boreal Treeline White Spruce Woodland, Boreal Mesic Black Spruce Forest, or, less commonly, in these same systems in the sub-boreal region. General Information Contributors (also see the Comments field Modeler 1 Tina Boucher Modeler 2 Kori Blankenship Modeler 3 antvb@uaa.alaska.edu kblankenship@tnc.org 3/26/2008 Reviewer Beth Schulz Reviewer Map Zone 73 Forest and Woodland General Model Sources PIGL ALVI5 BENA SAPU15 VAUL LEDUM bschulz@fs.fed.us Reviewer Vegetation Type Dominant Species* Date Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system occurs throughout the boreal transition region of AK and west to the limit of tree growth. It is not common in the Kenai Mountains, where mountain hemlock dominates treeline forest types, but may be found in western Kenai highlands between Skilak and Tustemena lakes and in higher elevations of Caribou Hills north of Homer. Boreal Spruce-Lichen Woodland, which is included as a variant of a later seral stage within this type, occurs commonly in the western and southwestern boreal transition region (Nulato Hills and Ahklun Mts). Biophysical Site Description This system occurs above White Spruce zone to 900m (Boggs et al. 2001) and below the subalpine shrub and tundra systems and can be seen as the forested transition zone between boreal white spruce forest and non-forested subalpine vegetation. Depending on the topography, this system can occupy a narrow band just below non-forested subalpine or a broad expanse across gently slopes and benches. This system also occurs at lower elevations at the western limit of white spruce. Soils are cold, but peat-forming mosses are not common in the ground layer (NatureServe 2008). Vegetation Description The Boreal Transition White Spruce Woodland occurs primarily near the elevational limit and the *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 6 of 204 western limit of tree growth. Forest canopy cover is dominated by white spruce and is generally between 10-25% (NatureServe 2008). Picea mariana may be codominant in the overstory. Common shrub species include Betula nana, but other shrubs such as Vaccinium uliginosum, Ledum groenlandicum and Salix pulchra may be common or dominant. In some locations Alnus viridis is the dominant understory shrub. Feathermoss may be common in the in the ground layer (NatureServe 2008). On drier or more exposed sites, Cladina spp. replace feathermosses as the dominant ground cover (Viereck 1979). Lichen species may include Cladina arbuscula, C. mitis, C. rangiferina and C. stellaris, as well as Cetraria cucullata, Cetraria islandica, Cetraria nivalis, Bryoria spp., Alectoria nigricans and Alectoria ochroleuca. Disturbance Description There is little information about the fire return interval for this type. Experts at the LANDFIRE Anchorage workshop indicated that White Spruce Woodlands in the AK sub-boreal region burn less frequently than those in the interior boreal region. They estimated a fire return interval of about 300yrs. Spruce bark beetle (Dendroctonus rufipennis) infestations are a major natural disturbance affecting this BpS. Beetle outbreaks in spruce forest occurred at an estimated return interval of every 52yrs on the Kenai Peninsula over the past 250yrs (Berg and Scott 2006). Outbreaks that thin stands and produce a growth release in surviving trees occur on average every 50yrs in white and Lutz spruce forests on the Kenai Peninsula (Berg 2004). Spruce bark beetle outbreaks that produce a more substantial thinning occur at longer intervals, with the last two severe infestations occurring in the 1870s-1880s and 1987-present (Berg 2004). The bark beetle outbreak that began in 1987 on the Kenai Peninsula has killed over 1.3 million acres of spruce (USDA Forest Service 2002). This recent outbreak is associated with warmer than average growing season temperatures that allowed beetles to mature in one year rather than two (Werner and Holsten 1985, Barber et al. 2000 as cited in Werner et al. 2006). Berg (2004) and Berg and Scott (2006) found no association between spruce bark beetle mortality and fire in the past. When the canopy of these forests is thinned by spruce bark beetle-mortality, bluejoint grass (Calamagrostis canadensis) may proliferate rapidly from its pre-disturbance low level network of rhizomatous roots and may develop into a thick, seedling-excluding sod within a few years (Berg 2004). Boucher (2003) found that rapid spread of Calamagrostis occurs primarily on sites with deep, loamy soils. Boucher and Mead (2006) found that vegetation response varied following the recent outbreak in different geographic locations on the Kenai Peninsula. Some areas exhibited an increase in early seral species (e.g. Calamagrostis canadensis and Chamerion angustifolium); other areas exhibited an increase in late seral mountain hemlock, while in other areas vegetation composition did not shift substantially (Boucher and Mead 2006). The following information was taken from the Ecological Systems draft description for Boreal Transition White Spruce Woodland (NatureServe 2008), which notes that fire information is lacking: A possible scenario for post-fire succession in this type is the resprouting of low shrubs from underground propagules, followed by invasion of Picea glauca by seed from adjacent stands or surviving trees. Betula papyrifera may invade the site if a seed source is available and site conditions are favorable, but the hardwood phase only occurs on a small fraction of the landscape (may be more common in southwest AK). The typical succession sequence for this type does not include a hardwood sere. The rate of succession depends on severity of fire and seed source, and some sites may be shrub-dominated for long periods without spruce invasion. Boreal Spruce-Lichen Woodland is included in this model as a possible variant of the late successional stage (class C). This is a best guess as to where this system fits, as its successional relationships are poorly *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 7 of 204 understood. Adjacency or Identification Concerns This BpS is similar to the interior Boreal Treeline White Spruce Woodland BpS but it includes an insects and disease probability and a less frequent MFRI. Adjacent systems may include Alaska Sub-boreal White Spruce-Hardwood Forest or Western North American Boreal Mesic Scrub Birch-Willow Shrubland Alaska Sub-boreal. Native Uncharacteristic Conditions Scale Description Large patch Issues/Problems Very little information is available on the fire return interval for this type. Comments This model was based on input from the experts who attended the LANDFIRE Anchorage modeling meeting (Dec. 07) with additional refinement by Tina Boucher and Kori Blankenship. It is similar to Western North American Boreal Treeline White Spruce Woodland - Boreal but includes insects and disease probability and a less frequent MFRI. Beth Schulz reviewed an initial draft of this model. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 5% Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Indicator Species* and Canopy Position BENA VAUL LEGR SAPU15 Upper Upper Upper Upper Height Dwarf Shrub (< 20 cm) Tree Size Class Low Shrub (20 cm to 1.5 m) Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. This class may uncommonly include tall shrubs. Description 0-24yrs Post disturbance regeneration: herbaceous to low shrub. Shrubs resprout from underground propagules and then Picea glauca invades by seed from adjacent stands or surviving trees. The shrub layer typically features Betula nana, but other shrubs such as Vaccinium uliginosum, Ledum groenlandicum and Salix pulchra may be common or dominant. In some locations Alnus viridis is the dominant understory shrub. Feathermoss may be present in the in the ground layer. The rate of succession depends on severity of fire and seed source, and some sites may be shrub-dominated for long periods without spruce invasion. The typical succession sequence for this type does not include a hardwood sere (class B). Succession to class C. Replacement MFRI = 400yrs. Alternate succession (probability = 0.04) causes a transition to class B and represents sites where Betula papyrifera may invade if a seed source is available and *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 8 of 204 site conditions are favorable. Class B Structure Data (for upper layer lifeform) Min Max 5% Mid Development 1 Open Cover Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position PIGL BEPA BENA Shrub Tree Woodland (10-24% tree cover) Open (25-59% tree cover) Tree (> 3 m) Tree Size Class Upper Upper Lower Lower Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. Presence of hardwoods distinguishes class B from C. Description 25-69yrs Woodland to open hardwood or white spruce-hardwood mix. Tree saplings gain canopy dominance over shrubs. Forest canopy cover is generally 10-25%. Betula papyrifera invades and stands may be either hardwoods or a spruce-hardwood mix. Succession to class C. Replacement MFRI = 400yrs. Mixed fire (MFRI = 1000yrs) maintains this class. Class C Structure Data (for upper layer lifeform) Min Max 90 % Late Development 1 Open Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position PIGL BENA VAUL CLADI3 Upper Low-Mid Low-Mid Lower Woodland (10-24% tree cover) Tree Size Class Open (25-59% tree cover) Tree (> 3 m) Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Absence of hardwoods distinguishes class C from B. Description 25yrs+ Open white spruce woodland. Site is dominated by mature conifers. Forest canopy cover is generally 1025%. Hardwoods, if previously present in the stand, lose dominance in overstory during this phase. The understory may include various combinations of low shrubs, herbs and mosses. Feathermoss are often common in the ground layer. On drier or more exposed sites, Cladina spp. replace feathermosses as the dominant ground cover (Viereck 1979). Lichens may become more common in older stands, with some sites developing into Spruce-Lichen Woodland. Clidina species include C. arbuscula, C. mitis, C. rangiferina, and C. stellaris. Other lichens include Cetraria cucullata, C. islandica, C. nivalis, Bryoria spp., Alectoria nigricans and Alectoria ochroleuca. This class will persist in the absence of disturbance. Replacement MFRI = 400yrs. Mixed fire (MFRI = 1000yrs) maintains this class. Insects and disease (probability = 0.02) will thin large overstory trees but will not cause a state transition. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 9 of 204 Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Min Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals V Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 398 1036 0.00251 0.00097 287 0.00349 Percent of All Fires 72 28 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Barber, V.A., Juday, P.A., Finney, B., 2000. Reduced growth of Alaskan white spruce in the twentieth century from temperature induced drought stress. Nature 405: 668–673. Berg, E. 2004. White spruce fire history on the Kenai Peninsula, Alaska, based on radiocarbon-dated soil charcoal. Unpublished manuscript. Kenai National Wildlife Refuge, Alaska. Berg, E.E and R.S. Anderson. 2006. Fire history of white and Lutz spruce forests on the Kenai Peninsula, Alaska, over the last two millennia as determined from soil charcoal. Forest Ecology and Management 227(3): 275-283. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 10 of 204 Berg, E.E, J.D. Henry, C.L. Fastie, A.D. De Volder and S.M. Matsuoka. 2006. Spruce beetle outbreaks on the Kenai Peninsula, Alaska, and Kluane National Park and Reserve, Yukon Territory: Relationship to summer temperatures and regional differences in disturbance regimes. Forest Ecology and Management 227(3): 219-232. Boggs, K., A. Garibaldi, J. Stevens, J. Grunblatt and T. Helt. 2001. Denali National Park and Preserve Landcover mapping project. Volume 2: Landcover classes and plant associations. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage, 707 A Street, Anchorage, AK. 164 pp. Boucher, T.V. 2003. Vegetation response to prescribed fire in the Kenai Mountains, Alaska. Research Paper PNW-RP-554. Portland, OR: USDA Forest Service, Pacific Northwest Research Station. 59 pp. Boucher, T.V. and B.R. Mead. 2006. Vegetation change and forest regeneration on the Kenai Peninsula, Alaska following a spruce beetle outbreak, 1987–2000. Forest Ecology and Management 227(3): 233-246. Foote, J.M. 1983. Classification, description, and dynamics of plant communities after fire in the taiga of interior Alaska. Res. Pap. PNW-307. Portland, OR. USDA Forest Service. Pacific Northwest Research Station. 108 pp. Matsuoka, S.M., E.H. Holsten, M.E. Shephard, R.A. Werner and R.E. Burnside. 2006. Spruce beetles and forest ecosystems of south-central Alaska. Forest Ecology and Management 227(3): 193-194. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. USDA Forest Service. 2002. Revised land and resource management plan, final environmental impact statement. Alaska Region Chugach National Forest. R10-MB-480d, May 2002. Viereck, L.A. 1979. Characteristics of treeline plant communities in Alaska. Holarctic Ecology. 2: 228-238. Werner, R.A. and Holsten, E.H., 1985. Effect of phloem temperature on development of spruce beetles in Alaska. In: Safrankik, L. (Ed.), The Role of the Host in the Population Dynamics of Forest Insects. For. Can., Pac. For. Cent. Victoria, British Columbia, pp. 155–163. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 11 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316030 Western North American Boreal White Spruce-Hardwood Forest This BPS is lumped with: Western North American Boreal White Spruce Forest This BPS is split into multiple models: Boreal White Spruce Forest was lumped with this type because it was felt that although it is a separate existing vegetation type, a separate biophysical environment could not be described for it. General Information Contributors (also see the Comments field Modeler 1 Kori Blankenship Modeler 2 Robert Lambrecht 3/12/2008 Reviewer Tina Boucher kblankenship@tnc.org Robert_Lambrecht@fws Reviewer .gov antvb@uaa.alaska.edu Reviewer Modeler 3 Map Zone 73 Vegetation Type Forest and Woodland Dominant Species* PIGL PIMA BEPA POTR5 Date General Model Sources ARUV VAVI BENA CHANA Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range Boreal White Spruce-Hardwood Forest occurs from the southern slopes of the Brooks Range to southcentral AK and west to the limit of tree growth. Biophysical Site Description These systems occur on rolling hills, inactive terraces, and mountain side slopes up to 750m. Soils are typically derived from glacial or other depositional processes and include moraines, drumlins, eskers, kettle-kame, colluvium and loess deposits (NatureServe 2008). These systems commonly occur on welldrained upland terrain on west, east and south aspects, but are possible on all aspects (NatureServe 2008). Permafrost is commonly absent. Boreal Spruce-Lichen Woodland occurs most commonly on cool, well-drained sites with thin soils (NatureServe 2008). Vegetation Description Canopy cover in mature stands is dominated by Picea glauca and typically ranges from 25-70%, except in the case of Boreal Spruce-Lichen Woodland, which has a more open canopy (10-25% cover). Other trees such as Picea mariana, Betula papyrifera and Populus tremuloides may be subdominant in the overstory, but Picea glauca contributes at least 75% of the total forest canopy in the forested type (NatureServe 2008). Mature stands are often open-canopied with a well-developed shrub layer (NatureServe 2008). The woodland type may be dominated by Picea mariana (NatureServe 2008). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 12 of 204 Common understory shrubs include Arctostaphylos uva-ursi, Vaccinium vitis-idaea, V. uliginosum, Betula nana, Empetrum nigrum, Ledum palustre ssp. decumbens, L. groenlandicum, Rosa acicularis and Viburnum edule. Arctostaphylos rubra and Shepherdia canadensis are typically found on dryer sites (Viereck et al. 1992). Common herbaceous species include Chamerion angustifolium ssp. angustifolium and Calamagrostis canadensis. Other herbaceous species can include Equisetum sylvaticum, E. arvense, Geocaulon lividum, Mertensia paniculata, Pyrola ssp., Linnaea borealis and Goodyera repens (Viereck et al. 1992). Feather mosses such as Hylocomium splendens and Pleurozium schreberi are common in the ground layer of the forested type (Boggs and Sturdy 2005). In the mature woodland type, feather mosses are less important, while lichens, primarily Cladina spp, form a very important component of the understory (NatureServe 2008). Disturbance Description The disturbance regime is characterized by large crown fires with estimates of mean fire return intervals ranging from 50-238yrs (Rowe 1972; Heinsleman 1981; Yarie 1981, 1983; Foote 1983; Duchesne and Hawkes 2000). Except in the case of severe fires, post-fire succession tends to return to the pre-disturbance forest type (Foote 1983). Pre-burn species colonize the site via rhizomes, root sprouts and trunk sprouts (NatureServe 2008). A variety of herbaceous communities dominate: primarily Chamerion angustifolium ssp. angustifolium and Calamagrostis canadensis (NatureServe 2008). Betula papyrifera, Populus tremuloides or Picea glauca may individually invade and dominate sites, but eventually Picea glauca gains dominance over hardwoods (NatureServe 2008). In severe fires, the organic layer is consumed killing the underground propagules, and revegetation of the site is by seed. A typical successional sequence progresses from herbaceous to shrub to hardwood/hardwood-spruce to spruce (NatureServe 2008). In upland spruce stands post-burn succession occasionally skips the hardwood/hardwood-spruce stage (class C in this model) and proceeds directly to a spruce dominated stage (Viereck et al. 1992). The former successional scenario is typical of the Fairbanks area while the later is more common around Yukon Charley, Noatak, NE boreal and at higher elevations (NatureServe 2008). Boreal Spruce-Lichen Woodland may occur as a very late successional stage of this system. Post-fire regeneration of white spruce appears to be more successful when fires occur in mast years (Peters et al. 2005). This interaction between fire, masting and subsequent tree regeneration could have implications for historical stand structure and successional dynamics over time (Peters et al. 2005). Adjacency or Identification Concerns This system may be found alongside any boreal black spruce or hardwood systems. Native Uncharacteristic Conditions Recent warmer and drier conditions, along with human activities including fire suppression and some logging practices have likely increased the current frequency and severity of spruce bark beetle outbreaks compared with presettlement conditions. Scale Description This BpS occurs in a matrix with other vegetation types. Crown fires were typically large patch occurrences. Issues/Problems It should be noted that there is a considerable variation across the range of this community type. This description and the associated model attempt to capture the most common attributes of the system across *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 13 of 204 its range. Comments This model was based on input from the experts who attended the LANDFIRE Fairbanks modeling meeting (Nov. 07) and refined by Robert Lambrecht. Experts from this workshop indicated the potential need for a self-replacement spruce model and another for the short-term mix of spruce and hardwood. This model represents both concepts using a deterministic pathway to represent the more common sprucehardwood pathway and an alternate succession pathway to represent the less common spruce-spruce pathway. This model may need to be split into two regional variants: this model to cover most of the boreal region and a variant with a lower fire return interval and no birch, to apply to the Copper River Basin and the Wrangell Mountains. Though Robert Lambrect originally lumped Western North American Boreal Spruce-Lichen Woodland into this model as a late seral stage, a reviewer suggested that the Spruce Lichen Woodland would be a better fit as a seral stage of the Boreal and Sub-boreal Treeline White Spruce Woodland and the Mesic Black Spruce Forest models. Colleen Ryan removed the reference to the lump in this model but did not alter the description or VDDT model. The model still includes a seral stage with lichens. Tina Boucher reviewed a draft of this model. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 5% Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position CHANA2 CACA4 EQSY EQAR Upper Upper Upper Upper Herbaceous Height Herbaceous Tree Size Class Seedling/Sapling <5" Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description 0-4yrs Post disturbance regeneration. A variety of herbaceous communities dominate; primarily Chamerion angustifolium ssp. angustifolium and Calamagrostis canadensis. Other herbaceous species can include Equisetum sylvaticum, E. arvense, Geocaulon lividum, Mertensia paniculata and Pyrola ssp. (Viereck et al. 1992). Shrubs and trees resprout from root stocks, but woody cover is low. Succession to class B. Replacement MFRI = 150yrs. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 14 of 204 Class B Structure Data (for upper layer lifeform) Min Max 15 % Early Development 2 All Structures Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position ARUV VAVI BENA LEPAD Shrub Tree Dwarf Shrub (< 20 cm) Tree Size Class Upper Upper Upper Upper Tall Shrub (>1.5 m) Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Description 5-29yrs This stage is dominated by shrubs and saplings. Common shrub species include Rosa acicularis, Viburnum edule, Betula nana, Ledum palustre ssp. Decumbens, L. groenlandicum, Vaccinium vitis-idaea, V. uliginosum and Empetrum nigrum. Arctostaphylos uva-ursi, Arctostaphylos rubra and Shepherdia canadensis are typically found on dryer sites (Viereck et al. 1992). Betula papyrifera and Populus tremuloides saplings are common on some sites. Succession to class C. The alternate succession pathway to D (probability = 0.01) represents self-replacing white spruce stands in areas where there is no adjacent hardwood seed source (e.g., Noatak, northeast boreal and higher elevation). Replacement MFRI = 150yrs. Class C Structure Data (for upper layer lifeform) Min Max 30 % Mid Development 1 Open Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position BEPA POTR5 PIGL ROAC Upper Upper Upper Upper Open (25-59% tree cover) Closed (60-100% tree cover) Dwarf Tree (< 3 m) Tree Size Class Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. Presence of hardwoods distinguishes class C from D. Description 30-129yrs This is predominantly a hardwood forest although conifers may be present and mixed with the hardwoods. Trees begin to shade out the shrub understory. The overstory dominants include Betula papyrifera and Populus tremuloides. Picea glauca and P. mariana may be present. Common understory species include Rosa acicularis, Viburnum edule, Arctostaphylos spp., Linnaea borealis, Chamerion angustifolium and Geocaulon lividum. Succession to class E. Replacement MFRI = 150yrs. Mixed fire (MFRI = 300yrs) maintains this class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 15 of 204 Structure Data (for upper layer lifeform) Class D 10 % Mid Development 2 Open Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position PIGL BENA ARRU VAVI Upper Lower Lower Lower Cover Height Min Woodland (10-24% tree cover) Max Closed (60-100% tree cover) Tree (> 3 m) Tree Size Class Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. Absence of hardwoods distinguishes class D from C. There are not mapping criteria to distinguish class D from E but trees are likely shorter and the stand likely more dense in D than in E. Description 30-129yrs This class represents mid-seral, self-replacing white spruce stands in areas where there is no adjacent hardwood seed source or geographic areas that tend to lack the hardwood component such as the Noatak, NE boreal region and higher elevations. Picea glauca dominates the overstory but P. mariana may be present. Succession to class E. Replacement MFRI = 150yrs. Mixed fire (MFRI = 300yrs) maintains this class. Class E Structure Data (for upper layer lifeform) 40 % Min Late Development 1 Open Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position PIGL ROAC VIED BENA Upper Lower Lower Lower Woodland (10-24% tree cover) Max Closed (60-100% tree cover) Tree (> 3 m) Tree Size Class Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Absence of hardwoods distinguishes class E from C. There are not mapping criteria to distinguish class D from E but trees are likely shorter and the stand likely more dense in D than in E. Description 130yrs+ Mature spruce forest. Hardwoods senesce. Accumulation of evergreen litter begins to change soil characteristics. Picea glauca dominates the overstory but P. mariana may be present. Common understory species include Rosa acicularis, Viburnum edule, Shepherdia canadensis, Vaccinium vitis-idaea, Arctostaphylos spp.,Linnaea borealis, Chamerion angustifolium and Geocaulon lividum. This stage incorporates the concept of Boreal Spruce-Lichen Woodland where lichens, primarily Cladina spp., are a very important component of the understory. Feather mosses are not as important as in other white spruce systems. This class persists in the absence of disturbance. Replacement MFRI = 150yrs. Mixed fire (MFRI = 300yrs) *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 16 of 204 maintains this class. Disturbances Fire Regime Group**: Fire Intervals IV Replacement Mixed Historical Fire Size (acres) Avg FI Min FI Max FI Probability 149.7 364.2 0.00668 0.00275 106 0.00943 Percent of All Fires 71 29 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Boggs, K. and Sturdy, M. 2005. Plant associations and post-fire vegetation succession in Yukon-Charley Rivers National Preserve. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage. Prepared For: National Park Service, Landcover Mapping Program, National Park Service-Alaska Support Office, Anchorage, AK 99501. Duchesne L.C. and B.C. Hawkes. 2000. Fire in northern ecosystems. In: Brown, J.K. and J.K. Smith (eds.) Wildland fire in ecosystems: effects of fire on flora. Gen. Tech. Rep. RMRS-GTR-42-vol 2. Ogden, UT: USDA Forest Service, Rocky Mountain Research Station. 257 pp. Foote, J.M. 1983. Classification, description, and dynamics of plant communities after fire in the taiga of interior Alaska. Res. Pap. PNW-307. Portland, OR. USDA Forest Service. Pacific Northwest Research Station. 108 pp. Heinselman, M.L. 1981. Fire and succession in the conifer forests of northern North America. In: West, D.C., H.H. Shugart, and D.B. Botkin. Forest succession: concepts and application. Springer-Verlag, New York. Chapter 23. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Peters, V.S., S.E. Macdonald and M.R.T. Dale. 2005. The interaction between masting and fire is key to white spruce regeneration. Ecology 86(7): 1744-1750. Rowe, J.S. 1972. Forest Regions of Canada. Canadian Forest Service, Department of Environment. Ottawa. Inform. Can. Catalogue #FO 47-1300. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 17 of 204 Yarie J. 1981. Forest fire cycles and life tables – a case study from interior Alaska. Can. J Forest Res. 11:554-562. Ritter, D. F. 1986. Process geomorphology. Wm. C. Brown Publishers, Dubuque, Iowa. 579 pp. Yarie J. 1983. Forest community classification of the Porcupine River drainage, interior Alaska, and its application to forest management. USDA Forest Service GTR PNW-154. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 18 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316041 Western North American Boreal Mesic Black Spruce Forest - Boreal This BPS is lumped with: WNA Boreal Spruce-Lichen Woodland (in part) This BPS is split into multiple models: Western North American Boreal Mesic Black Spruce Forest was split into a boreal and sub-boreal variant for BpS modeling so that regional differences could be represented. Boreal Spruce Lichen Woodland may occur as a seral stage or variant of Boreal Treeline White Spruce Woodland, Boreal Mesic Black Spruce Forest, or, less commonly, in these same systems in the sub-boreal region. General Information Contributors (also see the Comments field Modeler 1 Jill Johnstone Date jill.johnstone@usask.ca Reviewer Will Putnam wputman@tananachie fs.org Reviewer Reviewer Modeler 2 Modeler 3 Map Zone 73 Vegetation Type Forest and Woodland Dominant Species* PIMA BEPA POTR5 PIGL 3/12/2008 General Model Sources BENA LEDUM SALIX CACA4 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This type occurs throughout the boreal region of AK. Biophysical Site Description This type is typically found on upland slopes and inactive alluvial deposits (NatureServe 2008). Soils are well-drained, and permafrost is generally absent (NatureServe 2008). There is little to no peat development; but there may be an organic layer derived from non-sphagnum mosses (NatureServe 2008). Vegetation Description Picea mariana is the dominant overstory species, but Picea glauca may be codominant on some sites (NatureServe 2008). Early successional stands may be dominated by Betula papyrifera or Populus tremuloides (NatureServe 2008). Populus tremuloides replaces Betula papyrifera on drier sites (Foote 1983, Chapin et al. 2006). Common understory shrubs include Rosa acicularis, Betula nana, Ledum spp., V. uliginosum, Vaccinium vitis-idaea and Empetrum nigrum (NatureServe 2008). Herbaceous species include Calamagrostis canadensis, Chamerion angustifolium (Epilobium angustifolium) and Equisetum spp. Common mosses include Hylocomium splendens and Pleurozium schreberi (NatureServe 2008). Lichens, such as Cladina spp., may be abundant, especially in later seral stages (NatureServe 2008). Total tree cover in mature stands typically ranges from 40-70% (NatureServe 2008). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 19 of 204 Disturbance Description Crown fires or ground fires of enough intensity to kill overstory trees are the dominant disturbance influencing Boreal Mesic Black Spruce. Mean fire return interval estimates for Boreal Black Spruce in interior Alaska range from 25-130yrs (Yarie 1983, Heinselman1981, Viereck 1983, Viereck 1986). The post-fire successional trajectory may be self-replacement, with black spruce following the early seral herb and shrub stages; alternatively, black spruce-hardwood may follow the early seral stages before returning to black spruce (Chapin et al. 2006). The pre-burn stand composition will influence the likely successional trajectory with pre-burn spruce stands more likely to succeed to spruce and pre-burn hardwood stands more likely to succeed to hardwood after the fire. If white spruce is present in the conifer initiation, then white and black spruce can be codominant in the conifer canopy throughout the successional stages. Wind and insect damage affect this type, but very little research exists to help describe or model that effect. These disturbances are also much smaller in their impacts than the dominant, stand-replacement disturbances caused by fire. Adjacency or Identification Concerns Native Uncharacteristic Conditions Scale Description Matrix Issues/Problems The probability of fire in the mixed spruce-hardwood stage (class C) relative to the spruce dominated stages (class D and E) is unclear. It was assumed that the herb and shrub dominated stages (class A and B) are less fire prone, and therefore have a lower fire probability, than the later seral stages. The probability and effects of insects, disease and wind/weather events in this system are unclear and are included in the old-growth stage (class E) of the model as a place holder. Comments This model was based on input from the experts who attended the LANDFIRE Fairbanks (Nov. 07) and Anchorage (Dec. 07) modeling meetings and refined by Jill Johnstone. Will Putnam expressed concern that the distinction between this BpS and Western North American Boreal White Spruce-Hardwood Forest is ambiguous. They can occur on similar sites, and in many cases the relative dominance of P. mariana vs. P. glauca is mostly a function of succession and/or disturbance history. Vegetation Classes *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 20 of 204 Class A Structure Data (for upper layer lifeform) Min Max 5% Early Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Tree Size Class None Herbaceous Shrub Tree Indicator Species* and Canopy Position CACA4 CHAN9 EQUIS Upper Upper Upper Herbaceous Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description 0-4yrs Post-disturbance, herbaceous vegetation. Common herbaceous species include Calamagrostis canadensis, Chamerion angustifolium (Epilobium angustifolium) and Equisetum spp (NatureServe 2008). Succession to class B. Replacement MFRI = 250yrs. Class B Structure Data (for upper layer lifeform) Min Max 15 % Early Development 2 Open Cover Open Shrub (25-74% shrub cover) Open Shrub (25-74% shrub cover) Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position SALIX Upper BENA Upper LEDUM Upper ROAC Low Shrub (20 cm to 1.5 m) Seedling/Sapling <5" Tall Shrub (>1.5 m) Tree Size Class Upper layer lifeform differs from dominant lifeform. Description 5-29yrs This class is shrub dominated. Common species include willow, Betula nana, Ledum spp, Rosa acicularis, Vaccinium uliginosum, V. vitis-idaea and Empetrum nigrum. Both hardwoods and spruce regeneration may be present. The successional trajectory of the shrub stage will depend on the pre-burn stand composition. If hardwoods dominated the site before the burn, they are likely to vigorously resprout and gain dominance after the fire. If spruce dominated the site before a burn, they are likely to regain dominance in the post-fire stand. Succession to class C or D. One possible successional trajectory (represented by deterministic succession in the model) would include a hardwood phase (class C). But this class can also take an alternative successional pathway (probability = 0.02) directly to a spruce dominated phase (class D). Replacement MFRI = 250yrs. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 21 of 204 Class C 30 % Mid Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position BEPA POTR5 PIMA PIGL Cover Height Structure Data (for upper layer lifeform) Min Max Open (25-59% tree cover) Closed (60-100% tree cover) Tree (> 3 m) Tree Size Class Upper Upper Upper Upper Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. Presence of hardwoods distinguishes C from D and E. Description 30-119yrs Mixed hardwood-spruce forest. Hardwoods and spruce overtop shrubs and gain dominance. Early in this age class trees are at least 2.5cm DBH and 4-8m tall (Foote 1983). Populus tremuloides replaces Betula papyrifera on drier sites (Foote 1983, Chapin et al. 2006). Spruce may occur as an understory, subdominant and/or co-dominant component. Tree density may be < or > 60% depending on site conditions. Beneath trees, shrubs, herbs and mosses exist. As the stage advances, spruce and moss become more important. Succession to class E. Replacement MFRI = 150yrs. Mixed fire (MFRI = 250yrs) maintains this class. Structure Data (for upper layer lifeform) Class D 30 % Mid Development 2 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position PIMA PIGL BENA LEDUM Upper Upper Lower Lower Height Min Max Open (25-59% tree cover) Closed (60-100% tree cover) Dwarf Tree (< 3 m) Tree Size Class Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping purposes, if tree size class can't distinguish class D and E, class D can be considered closed and E woodland and open. Description 30-119yrs Mid-seral black spruce/feathermoss. Picea mariana dominates but Picea glauca may be codominant on some sites. Spruce overtops the shrubs. Spruce canopy cover is commonly 50-70%. Succession to class E. Replacement MFRI = 80yrs. Mixed fire (MFRI = 133yrs) maintains this class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 22 of 204 Class E Structure Data (for upper layer lifeform) 20 % Late Development 1 Open Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree PIMA PIGL BENA LEDUM Upper Upper Lower Lower Cover Height Min Woodland (10-24% tree cover) Max Open (25-59% tree cover) Tree (> 3 m) Tree Size Class Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping purposes, if tree size class can't distinguish class D and E, class D can be considered closed and E woodland and open. Description 120yrs+ Open, old-growth black spruce. Picea mariana dominates but Picea glauca may be codominant on some sites. Spruce gains dominance over hardwoods (if previously present). Tree canopy cover is generally <60% and may be <25% (woodland) depending on site conditions. Occasional hardwoods may remain. The understory may include various combinations of tall shrubs, low shrubs, herbs, mosses and lichens. If fire is absent for long periods paludification may occur, resulting in an opening of the tree canopy to woodland conditions. Black Spruce-Lichen Woodland may occur in this class. This class persists in the absence of disturbance. Replacement MFRI = 80yrs. Mixed fire (MFRI = 133yrs) maintains this class. Insects/disease (probability = 0.001) and wind/weather/stress (probability = 0.01) can cause a transition to class D. Disturbances Fire Regime Group**: Fire Intervals III Replacement Historical Fire Size (acres) Mixed Surface All Fires Avg 0 Min 0 Max 0 Avg FI Min FI Max FI Probability 111.9 206.3 0.00893 0.00485 73 0.01379 Percent of All Fires 65 35 Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Chapin, F.S., Oswood, M.W., Van Cleve, K., Viereck, L.A. and Verblya, D.L. (eds.) 2006. Alaska’s Changing Boreal Forest. Oxford University Press, NY. 354 pp. Heinselman, M.L. 1981. Fire and succession in the conifer forests of northern North America. In: West, D.C., H.H. Shugart, and D.B. Botkin. Forest succession: concepts and application. Springer-Verlag, New *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 23 of 204 York. Chapter 23. Foote, J.M. 1983. Classification, description, and dynamics of plant communities after fire in the taiga of interior Alaska. Res. Pap. PNW-307. Portland, OR. USDA Forest Service. Pacific Northwest Research Station. 108 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck, L.A. 1983. The effects of fire in black spruce ecosystems of Alaska and northern Canada. In: Wein, Ross W.; MacLean, David A., eds. The role of fire in northern circumpolar ecosystems. New York: John Wiley & Sons Ltd.: 201-220. Chapter 11. Viereck, L.A. Van Cleve, K. and Dyrness, C.T. 1986 Forest Ecosystems in the Alaska Taiga. In: Van Cleve, K., Chapin F.S. III, Flanagan, P.W. (and others), eds. Forest Ecosystems in the Alaskan taiga: a synthesis of structure and function. New York: Springer-Verlag; 1986: 22-43. Yarie J. 1983. Forest community classification of the Porcupine River drainage, interior Alaska, and its application to forest management. USDA Forest Service GTR PNW-154. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 24 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316042 Western North American Boreal Mesic Black Spruce Forest - Alaska Sub-boreal This BPS is lumped with: WNA Boreal Spruce-Lichen Woodland (in part) This BPS is split into multiple models: Western North American Boreal Mesic Black Spruce Forest was split into a boreal and sub-boreal variant for BpS modeling so that regional differences could be represented. Boreal Spruce Lichen Woodland may occur as a seral stage or variant of Boreal Treeline White Spruce Woodland, Boreal Mesic Black Spruce Forest, or, less commonly, in these same systems in the sub-boreal region. General Information Contributors (also see the Comments field Modeler 1 Michelle Schuman 4/7/2008 michelle.schuman@ak. usda.gov Reviewer Reviewer Reviewer Modeler 2 Modeler 3 Map Zone 73 Vegetation Type Forest and Woodland Dominant Species* PIMA PIGL BENA LEDUM Date General Model Sources VAUL VAVI EMNI HYSP70 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system occurs in the boreal transition region of AK, south of the Alaska Range, including the Susitna and Matanuska Valleys and the Kenai Peninsula (NatureServe 2008). Biophysical Site Description This information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): Sub-boreal Mesic Black Spruce Forest occurs on well-drained to moderately well-drained sites including old alluvial fans, abandoned floodplains and inactive terraces. Soils are gravelly and feature shallow to moderately deep organic horizons. Permafrost is absent. Vegetation Description This information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): Picea mariana and P. glauca are the dominant overstory species in an open forest canopy. Common understory shrubs include Betula nana, Ledum spp., V. uliginosum, Vaccinium vitis-idaea and Empetrum nigrum. Common mosses include Hylocomium splendens and Pleurozium schreberi. Total tree cover typically ranges from 40-70%. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 25 of 204 Disturbance Description This information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): The disturbance regime is characterized by crown fires or ground fires of enough intensity to kill overstory trees. Mean fire return interval estimates for this type have not been defined, but estimates for similar sites from interior Alaska range from 25-100yrs (Rowe et al. 1974, DeVolder 1999 [Kenai Lowland including human-caused], Yarie 1983, Heinselman 1978, Heinselman 1981, Viereck 1983, Viereck 1986). It is likely that the natural fire return interval is longer than those estimated for boreal sites due to less frequent lightning strikes. A “best guess” for this system without human disturbance has been estimated at 170yrs (FRCC expert’s consultation, 2004). Succession after fire can return to black spruce or can pass through a hardwood sere before returning to black spruce. Seasonality affects burn severity. An early season burn can kill the overstory without affecting the ground layer, but a late-season burn can reduce the duff layer and kill the understory plants. Calamagrostis is not a major factor in sub-boreal black spruce succession. Adjacency or Identification Concerns In some locations, this BpS can be confused with the White Spruce BpS because black and white spruces often mix, especially on sites with transitional moisture and thermal conditions (Murphy and Witten 2006). Native Uncharacteristic Conditions The following information was taken from the Black Spruce Southcentral PNVG model description (Murphy and Witten 2006): In recent decades black spruce began encroaching into drying sphagnum bogs, creating “islands” of spruce where spruce were not previously present (Ed Berg, personal communication, March 4, 2004). This drying and resultant encroachment is attributed to the warming climate. Scale Description This Bps is typically found in large patches (NatureServe 2008). The following information was taken from the Black Spruce Southcentral PNVG (Murphy and Witten 2006) model description: Fires tend to be large – 50,000ha or larger. During most fire years a small number of large fires account for most of the total area burned (Gabriel and Tande 1983). Ecologically significant fires usually occur during the exceptional fire years and cover 200,000ha+ (Viereck 1983). Issues/Problems Comments This model was based on the FRCC Guidebook PNVG model for Black Spruce Southcentral (BSSC; Murphy and Witten 2006) and input from the experts who attended the LANDFIRE Fairbanks (Nov. 07) and Anchorage (Dec. 07) modeling meetings. It was refined by Michelle Schuman. The class definitions and age ranges were taken from experts at the Anchorage and Fairbanks meetings with input from Michelle Schuman and the disturbance probabilities are similar to those in the BSSC model. Vegetation Classes *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 26 of 204 Class A Structure Data (for upper layer lifeform) Min Max 10 % Early Development 1 All Structures Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position BENA LEDUM VAUL VAVI Upper Middle Low-Mi Lower Dwarf Shrub (< 20 cm) Tree Size Class Tall Shrub (>1.5 m) Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Description 0-14yrs Moss, herbs, seedlings of trees and shrubs establish three months to three years post fire (Foote 1983). Shrubs and saplings 1.4-7m tall typically begin capturing sites 4-5yrs post fire. The tall shrub and sapling layer is characterized by 60-100% canopy closure. Tree saplings may include spruce, hardwoods or both. Common understory shrubs include Betula nana, Ledum spp., Vaccinium uliginosum, V. vitis-idaea and Empetrum nigrum. Common mosses include Hylocomium splendens and Pleurozium schreberi. Succession to class B. Alternate succession (probability = 0.011) causes a transition to class C and represents the probability that some stands will go through a hardwood (with spruce understory) or spruce-hardwood stage rather than following the main successional pathway to a black spruce dominated stage (class B). Replacement MFRI = 167yrs. Class B 10 % Mid Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position PIMA PIGL BENA LEDUM Upper Upper Lower Lower Cover Height Structure Data (for upper layer lifeform) Min Max Open (25-59% tree cover) Closed (60-100% tree cover) Dwarf Tree (< 3 m) Tree Size Class Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. B should be distinguished from D and E based on tree size class but if that is not mapped, B can be considered a dwarf tree class for mapping. Description 15-74yrs Black or white spruce overtops shrubs and gains dominance. Tree density may be < or > 60% depending on site conditions. The alternate succession pathway to E represents an alternative to the main successional trajectory to class D. Succession to class D. Alternate succession (probability = 0.015) causes a transition to class E and represents the probability that some stands go directly to a closed spruce stage rather than following the main successional pathway to an open spruce stage (class D). Mixed (MFRI = 830yrs) maintains the class. Replacement MFRI = 450yrs. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 27 of 204 Class C 5% Mid Development 2 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position BEPA POTR5 PIMA PIGL Cover Height Structure Data (for upper layer lifeform) Min Max Open (25-59% tree cover) Closed (60-100% tree cover) Tree (> 3 m) Tree Size Class Upper Upper Mid-Upper Mid-Upper Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. Presence of hardwoods distinguishes C from B, D and E. Description 15-74yrs Hardwoods (with spruce in the understory) or hardwoods and spruce overtop shrubs and gain dominance. Early in this age class trees are at least 2.5cm DBH and 4-8m tall (Foote 1983). Populus tremuloides replaces Betula papyrifera on drier sites (Foote 1983, Chapin et al. 2006). Spruce may occur as an understory, subdominant and/or co-dominant component. Tree density may be < or > 60% depending on site conditions. Beneath trees shrubs, herbs and mosses exist. As the stage advances spruce and moss become more important. Succession to class D. Replacement MFRI = 250yrs. Structure Data (for upper layer lifeform) Class D 50 % Late Development 1 Open Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position PIMA PIGL BENA LEDUM Upper Upper Lower Lower Min Woodland (10-24% tree cover) Height Tree Size Class Max Open (25-59% tree cover) Tree (> 3 m) Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Description 75yrs+ This class is characterized by open spruce lichen forest or woodland. Spruce gains dominance over hardwoods (if previously present). Tree canopy cover is <60% and maybe <25% (woodland) depending on site conditions. Occasional hardwoods may remain. The understory may include various combinations of tall shrubs, low shrubs, herbs, mosses and lichens. For spruce lichen woodland, the dominant lichen genus is Cladina; species include C. arbuscula, C. mitis, C. rangiferina and C. stellaris. Other lichens include Cetraria cucullata, C. islandica, C. nivalis, Bryoria spp., Alectoria nigricans and Alectoria ochroleuca. If fire is absent for long periods paludification may occur. This class persists in the absence of disturbance. Mixed (MFRI = 1400yrs) maintains the class. Replacement MFRI = 200yrs. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 28 of 204 Class E Structure Data (for upper layer lifeform) 25 % Late Development 1 Closed Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree PIMA PIGL BENA LEDUM Upper Upper Lower Lower Cover Height Min Closed (60-100% tree cover) Max Closed (60-100% tree cover) Tree (> 3 m) Tree Size Class Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Description 75yrs+ This class is characterized by closed spruce forest. Site is dominated by mature black or white spruce with >60% canopy closure although cover generally does not exceed 70%. The understory may include various combinations of tall shrubs, low shrubs, herbs, mosses and lichens. This class persists in the absence of disturbance. Mixed (MFRI = 1400yrs) causes a transition to class D. Replacement MFRI = 200yrs. Insects and disease (probability = 0.001) and wind/weather/stress (probability = 0.001) cause a transition to class D. Disturbances Fire Regime Group**: Fire Intervals IV Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 216.9 1430 0.00461 0.0007 188 0.00532 Percent of All Fires 87 13 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Chapin, F.S., Oswood, M.W., Van Cleve, K., Viereck, L.A. and Verblya, D.L. (eds.) 2006. Alaska’s Changing Boreal Forest. Oxford University Press, NY. 354 pp. De Volder, A. 1999. Fire and climate history of lowland black spruce forests, Kenai National Wildlife Refuge, Alaska. Flagstaff, AZ. Northern Arizona University. 128 pp. MS Thesis. Foote, J.M. 1983. Classification, description, and dynamics of plant communities after fire in the taiga of interior Alaska. Res. Pap. PNW-307. Portland, OR. USDA Forest Service. Pacific Northwest Research Station. 108 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 29 of 204 Gabriel, H.W. and G.F. Tande. 1983. A regional approach to fire history in Alaska. BLM Alaska TR-83-9. Heinselman, M.L. 1978. Fire in wilderness ecosystems. In J.C.Hendee, G.H. Stankey & R.C. Lucas, eds. Wilderness Management. USDA Forest Service, Misc. Pub. 1365. Heinselman, M.L. 1981. Fire and succession in the conifer forests of northern North America. In: West, D.C., H.H. Shugart, and D.B. Botkin. Forest succession: concepts and application. Springer-Verlag, New York. Chapter 23. Jorgenson, M.T. et al. 2003. An ecological land survey for Fort Richardson, Alaska. Cold Regions Research and Engineering Laboratory, Hanover, NH, ERDC/CRREL TR-03019. Murphy, K.A. and E. Witten. 2006. Black Spruce Southcentral. In Fire Regime Condition Class (FRCC) Interagency Guidebook Reference Conditions. Available at www.frcc.gov. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Rowe, J.S., J.L Bergsteinsson, G.A. Padbury and R. Hermesh. 1974. Fire Studies in the Mackenzie Valley. ALUR Report 73-74-61. Arctic Land Use Research Program, Department of Indian Affairs and Human Development, Ottawa, Canada. 123 pp. Viereck, L.A. 1983. The effects of fire in black spruce ecosystems of Alaska and northern Canada. In: Wein, Ross W.; MacLean, David A., eds. The role of fire in northern circumpolar ecosystems. New York: John Wiley & Sons Ltd.: 201-220. Chapter 11. Viereck, L.A. Van Cleve, K. and Dyrness, C.T. 1986 Forest Ecosystems in the Alaska Taiga. In: Van Cleve, K., Chapin F.S. III, Flanagan, P.W. (and others), eds. Forest Ecosystems in the Alaskan taiga: a synthesis of structure and function. New York: Springer-Verlag; 1986: 22-43. Yarie J. 1983. Forest community classification of the Porcupine River drainage, interior Alaska, and its application to forest management. USDA Forest Service GTR PNW-154. Yarie J. 1981. Forest fire cycles and life tables – a case study from interior Alaska. Can. J Forest Res. 11:554-562. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 30 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316050 Western North American Boreal Mesic BirchAspen Forest This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Michelle Schuman Modeler 3 Kori Blankenship Reviewer Will Putnam Forest and Woodland General Model Sources VIED SHCA ALNUS LEDUM Literature Local Data Expert Estimate wputman@tananachie fs.org Reviewer Reviewer Map Zone 73 Vegetation Type BEPA POTR5 POBA2 ROAC 4/16/2008 michelle.schuman@ak. usda.gov mitch.michaud@ak.usd a.gov kblankenship@tnc.org Modeler 2 Mitch Michaud Dominant Species* Date Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range Found throughout boreal AK. Biophysical Site Description This system occurs on rolling hills and mountain sideslopes on west, east, and south aspects up to 750m (NatureServe 2008). Soils are well-drained and develop on residual material or retransported deposits including glacial till, loess, and colluvium (NatureServe 2008). Hardwood-dominated sites often persist on slopes that are warmer and drier than white spruce or mixed white spruce hardwood sites, with aspen dominating the driest, warmest sites (Viereck et al. 1992, Chapin et al. 2006). Vegetation Description Canopy cover is dominated by Betula papyrifera or Populus tremuloides and typically ranges from 2590%. P. balsamifera may be a common associate. Stands are often closed-canopied with an open shrub or herbaceous understory. Common understory species include Alnus spp., Ledum spp., Vaccinium vitisidaea, Betula nana, Rosa acicularis, Viburnum edule and Equisetum spp. (NatureServe 2008). Shepherdia canadensis is common on drier sites, especially well-drained riparian gravel bars. Feathermosses such as Hylocomium splendens and Pleurozium schreberi are common in the ground layer (Jorgenson et al. 1999; Boggs and Sturdy, 2005). Disturbance Description Little research exists on the fire ecology of this type. The system often acts as a fire break. It is estimated that the MFRI is longer than that of white and black spruce sites and maybe comparable to Boreal White *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 31 of 204 Spruce-Hardwood Forest system. Adjacency or Identification Concerns This system can be easily confused with seral stages of two other ecological systems in boreal AK: Western North American Boreal White Spruce-Hardwood Forest and Western North American Boreal White Spruce Forest. Adjacent systems include Boreal White Spruce-Hardwood, Boreal White Spruce Forest or Boreal Mesic Black Spruce Forest. Native Uncharacteristic Conditions Recent ongoing leaf miner activity has been observed in birch and aspen, but no long-term information on its impact is available. Scale Description Large patch Issues/Problems There is uncertainty about whether Boreal Mesic Birch-Aspen Forest is a separate BpS from Boreal White Spruce-Hardwood Forest and Boreal White Spruce Forest. This system may occur only where spruce seed sources are lacking. Comments This model was based on input from the experts who attended the LANDFIRE Fairbanks modeling meeting (Nov. 07) and refined by Michelle Schuman, Mitch Michaud and Kori Blankenship with input from Tina Boucher. Boreal Mesic Birch-Aspen Forest is treated as a separate BpS within the boreal region because experts felt it could be distinguished as occupying different biophysical settings from the Boreal White Spruce Forest and Boreal White Spruce - Hardwood Forest systems. In contrast, the Boreal Mesic Birch-Aspen Forest system was lumped with the Sub-boreal White Spruce-Hardwood Forest system within the sub-boreal region because experts there felt that they could not distinguish the biophysical settings that these types occur on. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 5% Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position CHAN9 CACA4 EQUIS MEPA Upper Herbaceous Height Herbaceous Tree Size Class None Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description 0-4yrs Herbaceous species dominate, including Chamerion angustifolium ssp angustifolium, Calamagrostis canadensis, Equisetum sylvaticum, E. arvense, Mertensia paniculata and Geocaulon lividum. Shrubs are present but not dominant. Following fire, aspen resprouts and birch appears to invade by seed (Viereck and Schandelmeier 1980) *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 32 of 204 Succession to class B. Replacement MFRI = 200yrs. Class B Structure Data (for upper layer lifeform) Min Max 5% Early Development 2 All Structures Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position ROAC VIED LEDUM ALNUS Shrub Tree Dwarf Shrub (< 20 cm) Tree Size Class Upper Upper Upper Upper Tall Shrub (>1.5 m) Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Description 4-14yrs Shrubs gain dominance over the herbs. Hardwood seedlings are present. Common shrubs include Alnus spp., Ledum spp., Vaccinium vitis-idaea, Betula nana, Rosa acicularis, Shepherdia canadensis and Viburnum edule. Succession to class C. Replacement MFRI = 200yrs. Class C Structure Data (for upper layer lifeform) Min Max 15 % Mid Development 1 Closed Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position BEPA POTR5 ROAC VIED Closed (60-100% tree cover) Tree Size Class Upper Upper Lower Lower Closed (60-100% tree cover) Dwarf Tree (< 3 m) Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping purposes if classes C and D can't be distinguished based on structural characteristics, split them based on height (C = trees <3m; D = trees >3m). Description 15-49yrs Hardwoods gain dominance over shrubs. This class is characterized by dense stands of sapling and pole sized trees. Betula papyrifera or Populus tremuloides typically dominate but P. balsamifera may be a common associate. Common understory species include Alnus spp., Ledum spp., Vaccinium vitis-idaea, Betula nana, Rosa acicularis, Shepherdia canadensis, and Viburnum edule. This stage tends to be more flammable than the others (personal communication, Joan Foote). Succession to class D. Replacement MFRI = 150yrs. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 33 of 204 Structure Data (for upper layer lifeform) Class D 15 % Mid Development 2 Closed Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position BEPA POTR5 ROAC VIED Upper Upper Lower Lower Cover Height Min Closed (60-100% tree cover) Max Closed (60-100% tree cover) Tree (> 3 m) Tree Size Class Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Description 50-99yrs This stand is characterized by mature hardwood trees with more dead and downed fuel. Betula papyrifera or Populus tremuloides typically dominate but P. balsamifera may be a common associate. Common understory species include Ledum spp., Vaccinium vitis-idaea, Betula nana, Rosa acicularis, Shepherdia canadensis and Viburnum edule. Feathermosses such as Hylocomium splendens and Pleurozium schreberi are common in the ground layer (Boggs and Sturdy 2005). Succession to class E. Replacement MFRI = 200yrs. Class E Structure Data (for upper layer lifeform) 60 % Min Late Development 1 Open Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position BEPA POTR5 ALNUS LEDUM Upper Upper Lower Lower Open (25-59% tree cover) Tree Size Class Max Open (25-59% tree cover) Tree (> 3 m) Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Description 100yrs+ Late seral stands are characterized by large hardwood trees. This class captures the old, open birch calamagrostis stands. A mixed-age stand can develop as aspen clones resprout when individual trees die. Betula papyrifera or Populus tremuloides typically dominate but P. balsamifera may be a common associate. Spruce may be present in the canopy, and in the absence of fire, could potentially occupy the site. Common understory species include Alnus spp., Ledum spp., Vaccinium vitis-idaea, Betula nana, Rosa acicularis, Shepherdia canadensis and Viburnum edule. Feathermosses such as Hylocomium splendens and Pleurozium schreberi are common in the ground layer (Boggs and Sturdy 2005). This class persists in the absence of disturbance. Replacement MFRI = 200yrs. Mixed fire (MFRI = 300yrs) maintains this class. Disturbances *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 34 of 204 Fire Regime Group**: Fire Intervals IV Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 190.7 503.5 0.00524 0.00199 138 0.00724 Percent of All Fires 72 27 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Boggs, K. and Sturdy, M. 2005. Plant associations and post-fire vegetation succession in Yukon-Charley Rivers National Preserve. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage. Prepared For: National Park Service, Landcover Mapping Program, National Park Service-Alaska Support Office, Anchorage, AK 99501. Chapin, F.S., Oswood, M.W., Van Cleve, K., Viereck, L.A. and Verblya, D.L. (eds.) 2006. Alaska’s Changing Boreal Forest. Oxford University Press, NY. 354 pp. Jorgenson, M.T., J.E. Roth, M. Raynolds, M.D. Smith, W. Lentz, A. Zusi-Cobb, and C.H. Racine. 1999. An ecological land survey for Fort Wainwright, Alaska. U.S. Army Cold Regions Research and Engineering Laboratory, Hanover, NH. U.S. Army Cold Regions Research Engineering Laboratory, Hanover, NH CRREL Report 99-9. 83 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck, L.A. and L.A. Schandelmeier. 1980. Effects of fire in Alaska and adjacent Canada: a literature review. USDI BLM. GLM-Alaska Technical Report 6. Alaska State Office. Anchorage, AK. Viereck, L.A., Dyrness, C.T., Batten, A.R. and Wenzlick, K.J. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 35 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316061 Western North American Boreal Dry AspenSteppe Bluff - Lower Elevations This BPS is lumped with: This BPS is split into multiple models: Western North American Boreal Dry Aspen-Steppe Bluff was split into a lower elevation and a higher elevation model. The lower elevation model occur below treeline and trees are present. The higher elevation model generally occurs above treeline although sparse aspen (<10% cover) may be present. General Information Contributors (also see the Comments field Modeler 1 Mitch Michaud Modeler 2 Michelle Schuman 3/8/2008 mitch.michaud@ak.usd a.gov michelle.schuman@ak. usda.gov Reviewer Reviewer Reviewer Modeler 3 Map Zone 73 Vegetation Type Forest and Woodland Dominant Species* POTR5 PIGL ROAC JUCO6 Date General Model Sources ARAL5 CAPU PSSP6 ARFR4 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This type is found in AK from the southern slopes of the Brooks Range to southcentral AK and west to the limit of tree growth (NatureServe 2008). Biophysical Site Description This system occurs commonly on moderately steep to very steep, dry, south-facing slopes and on windswept bluffs above major rivers (NatureServe 2008). Soils are typically well-drained to excessively welldrained and develop on glacial, loess, or fluvial deposits or residual material (NatureServe 2008). Soils are often unstable and rocky outcrops are common (NatureServe 2008). Vegetation Description Disturbance Description Fire and insects are the dominant disturbance factors influencing this vegetation type. White Spruce (Picea glauca), which can be present in later seral stages, tends to take longer to regenerate after fire than aspen which quickly resprouts. As a result, fire tends to favor aspen but spruce will eventually come back in (Viereck et al. 1992). In the VDDT model the replacement fire probability is the same in all classes for lack of evidence to suggest otherwise. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 36 of 204 Large aspen tortrix (Choristoneura conflictana) is an aspen defoliator which can have a severe effect on mature and overmature aspen stands. Although aspen tortrix would have been present under the reference condition, its presence and affect have increased in fire suppressed areas today. Other disturbance agents include aspen leaf miners (Phyllocinistis populiella) and ground rot. Adjacency or Identification Concerns Native Uncharacteristic Conditions Scale Description Large patch Issues/Problems Need more information on the fire return intervals for this type. Comments This model was based on input from the experts who attended the LANDFIRE Fairbanks (Nov. 07) and Anchorage modeling meetings (Dec. 08) and refined by Mitch Michaud and Michelle Schuman. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 5% Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Indicator Species* and Canopy Position CAPU PSSP6 ARFR4 JUCO6 Upper Upper Upper Upper Height Dwarf Shrub (< 20 cm) Tree Size Class Low Shrub (20 cm to 1.5 m) Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Herbs or shrubs can dominate this class. Description 0-2yrs This stage is characterized by dry herbaceous and low shrub vegetation coming in after a major disturbance. Common shrub species include Vaccinium vitis-idaea, Rosa acicularis, Shepherdia canadensis and Chamerion angustifolium (Boggs and Sturdy, 2005). Very dry sites on bluffs typically have steppe vegetation including Juniperus communis, Arctostaphylos uva-ursi, Artemisia frigida, A. alaskana, Calamagrostis purpurascens, Pseudoroegneria spicata (= Agropyron spicatum), Festuca altaica and Poa spp (Viereck et al. 1992, Chapin et al. 2006 p.89). This stage was modeled with a duration of two years. This represents a typical post-fire scenario where aspen quickly resprout and overtop the shrubs in 1-2yrs. It is possible that this stage may last much longer. For example, some sites severely affected by the tortrix moth (Choristoneura conflictana) can become heavily dominated by Calamagrostis canadensis which can impede aspen regeneration, possibly for up to 10yrs, and increase the ability of the site to carry fire. Succession to class B. Replacement MFRI = 100yrs. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 37 of 204 Class B Structure Data (for upper layer lifeform) Min Max 15 % Mid Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position POTR5 ROAC JUCO6 ARAL5 Shrub Tree Open (25-59% tree cover) Closed (60-100% tree cover) Dwarf Tree (< 3 m) Tree Size Class Upper Mid-Upper Mid-Upper Mid-Upper Dwarf Tree (< 3 m) Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Description 3-9yrs This stage is characterized by suckering Populus tremuloides and low shrubs. Populus tremuloides tends to overtop the shrub and herbaceous vegetation. The shrubs and herbaceous species listed in class A are still present. For mapping purposes this class was defined as a state where trees are less than three meters tall. The age range at which Populus tremuloids exceeds three meters in height varies based on site conditions and can often happen in less than 10yrs. Heavy ungulate browsing will delay growth. Succession to class C. Replacement MFRI = 100yrs. Class C Structure Data (for upper layer lifeform) Min Max 70 % Late Development 1 Open Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position POTR5 ROAC JUCO6 ARAL5 Upper Lower Lower Lower Woodland (10-24% tree cover) Tree Size Class Closed (60-100% tree cover) Tree (> 3 m) Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping, class C can be distinguished from class D by the absence of spruce in the upper-layer. Description 10yrs+ This stage is characterized by mature and eventually over-mature Populus tremuloides. Populus tremuloides dominates the site but Picea glauca may be present in the mid or understory. The shrubs and herbaceous species listed in class A may still be present. Experts who provided input at the LANDFIRE Fairbanks (Nov. 07) and Anchorage modeling meetings (Dec. 08) described a mature and an over-mature aspen sere. Both concepts are incorporated into this stage because it was felt that LANDFIRE mapping teams would not be able to distinguish them as separate units. This class can persist in the absence of disturbance or follow an alternate succession pathway to class D (probability = 0.004) which represents sites where Picea glauca is present in the overstory. Replacement MFRI = 100yrs. Mixed fire (MFRI = 200yrs) maintains this class. Insects/disease (probability = 0.0125) *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 38 of 204 cause a transition to class A. Structure Data (for upper layer lifeform) Class D 10 % Late Development 2 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position POTR5 PIGL ROAC JUCO6 Upper Upper Lower Lower Min Max Woodland (10-24% tree cover) Closed (60-100% tree cover) Height Tree (> 3 m) Tree Size Class Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping, class D can be distinguished from class C by the presence of spruce in the upper-layer. Description 60yrs+ This stage is characterized by mature Populus tremuloides mixed with Picea glauca. Populus tremuloides typically decline between 60 and 100yrs of age allowing spruce to become more dominant on some sites (Viereck et al. 1992). Aspen are generally even aged initiating from a disturbance event while spruce tend to be uneven aged and come into the stand gradually over time (Viereck et al.1992). Canopy cover can range from woodland to closed but stands tend to open up overtime. The shrubs and herbaceous species listed in class A may still be present. This class can persist in the absence of disturbance. Replacement MFRI = 100yrs. Mixed fire (MFRI = 200yrs) maintains this class. Insects/disease (probability = 0.0125) cause a transition to class A. Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 39 of 204 Fire Regime Group**: Fire Intervals III Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 100.2 241.1 0.00998 0.00415 71 0.01414 Percent of All Fires 71 29 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Boggs, K. and Sturdy, M. 2005. Plant associations and post-fire vegetation succession in Yukon-Charley Rivers National Preserve. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage. Prepared For: National Park Service, Landcover Mapping Program, National Park Service-Alaska Support Office, Anchorage, AK 99501. Chapin, F. S., Oswood, M. W., Van Cleve, K., Viereck, L. A., Verblya, D. L. (eds.) 2006. Alaska’s Changing Boreal Forest. Oxford University Press, NY. 354 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 40 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316062 Western North American Boreal Dry AspenSteppe Bluff - Higher Elevations This BPS is lumped with: This BPS is split into multiple models: Western North American Boreal Dry Aspen-Steppe Bluff was split into a lower elevation and a higher elevation model. The lower elevation model occur below treeline and trees are present. The higher elevation model generally occurs above treeline although sparse aspen (<10% cover) may be present. General Information Contributors (also see the Comments field Modeler 1 Tina Boucher Modeler 2 Colleen Ryan antvb@uaa.alaska.edu colleen_ryan@tnc.org 4/24/2008 Reviewer Reviewer Reviewer Modeler 3 Map Zone 73 Vegetation Type Upland Shrubland Dominant Species* ARFR4 ARAL5 PSSP6 JUCO6 Date General Model Sources ARUV BRINA CAPU Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system occurs at high elevations throughout the boreal and sub-boreal regions of AK. Biophysical Site Description This low open shrub system occurs in the alpine or subalpine on steep south-facing, wind-swept bluffs and ridges. It may be associated with river systems above treeline. The substrate is steep, unstable, dry mineral soil. Rocky outcrops are common. Vegetation Description The vegetation cover in this system is typically open and discontinuous with much exposed mineral soil. Artemisia species are the primary characteristic dominants, but this system supports a unique assemblage of species including Artemisia frigida, Artemisia alaskana, Juniperus communis, Arctostaphylos uva-ursi, Pseudoroegneria spicata (= Agropyron spicatum), Bromus pumpellianus, Calamagrostis purpurascens, Festuca altaica and Poa spp. (Chapin et al. 2006). Populus tremuloides may be present, but canopy cover is <10% (NatureServe 2008). Disturbance Description The fire regime for this system is expected to be similar to that of the adjacent Western North American Boreal Dry Aspen-Steppe Bluff - Lower Elevations. However, this system has less fuel, which would tend to produce fewer and patchier fires. For this model, the FRI for replacement fires was estimated at 140yrs (vs. 100yrs for Boreal Dry Aspen and Steppe Bluff). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 41 of 204 Because this system is often found on unstable slopes with dry mineral soils, shifting slopes are an ongoing disturbance. Grazing is probably an important factor in shaping this system, as this is important Dall sheep habitat. Neither grazing nor erosion was included as disturbances in the model because these are long-term, ongoing phenomena. Adjacency or Identification Concerns This system is similar to Western North American Boreal Dry Aspen-Steppe Bluff - Lower Elevations, but lacks aspen and rose as dominant species. This system occurs above treeline and may be adjacent to Western North American Boreal Dry Aspen-Steppe Bluff - Lower Elevations, which includes a mosaic of aspen and steppe vegetation. Native Uncharacteristic Conditions Scale Description Linear; small patch Issues/Problems Comments Fire frequencies in this model were adapted from the Western North American Boreal Dry Aspen-Steppe Bluff - Lower Elevations model created by Mitch Michaud and Michelle Schuman. Suggested reviewers for this system include: Carl Roland and Dalia Vargis-Kretzinger for information on grazing. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 5% Early Development 1 Open Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position PSSP6 BRINA CAPU feal Upper Upper Upper Upper Herbaceous Height Herbaceous Tree Size Class None Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Typically herb-dominated, but herb cover may be sparse, and sprouting shrubs may be present. Description 0-4yrs Post-disturbance dry herbaceous or sparse shrubs. Grasses typically dominate the site. Shrubs may sprout from rootstock. Herbaceous cover tends to be open or sparse. In some cases, grasses may be absent and the ground will be bare until shrubs resprout. Pseudoroegneria spicata (= Agropyron spicatum), Bromus pumpellianus, Calamagrostis purpurascens and Festuca spp. are typical species. Artemisia frigida and Artemisia alaskana are typically regenerating in the understory. Succession to class B. Replacement MFRI = 140yrs. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 42 of 204 Class B Structure Data (for upper layer lifeform) Min Max 95 % Late Development 1 Open Cover Open Shrub (25-74% shrub cover) Open Shrub (25-74% shrub cover) Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position ARFR4 ARAL5 PSSP6 BRINA Shrub Tree Low Shrub (20 cm to 1.5 m) Tree Size Class Low Shrub (20 cm to 1.5 m) Pole 5-9" DBH Upper Upper Upper Upper Upper layer lifeform differs from dominant lifeform. Description Five years plus Dry, low, open shrubs. Mature shrubs. Shrubs become dominant, but an herbaceous layer usually persists. Shrub cover is 25-75%. Artemisia frigida and Artemisia alaskana dominate. Common grasses include Pseudoroegneria spicata (= Agropyron spicatum), Bromus pumpellianus and Calamagrostis purpurascens. Populus tremuloides may be present, but canopy cover is <10%. This class persists in the absence of disturbance. Replacement MFRI = 140yrs. Mixed (MFRI = 200yrs) maintains this class. Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 43 of 204 Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals III Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Avg FI Min FI Max FI Probability 143.2 206.6 0.00698 0.00484 85 0.01183 Percent of All Fires 59 41 Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Batten, A.R., Murray, D.F. and Dawe, J.C. 1979. Threatened and Endangered Plants in selected areas of the BLM Fortymile Planning Unit.. File Report for Contract No. YA-512-CT8-162. USDI BLM AK State Office 701 C Street, Anchorage, AK 99513. 127 pp. Boggs, K. and Sturdy, M. 2005. Plant associations and post-fire vegetation succession in Yukon-Charley Rivers National Preserve. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage. Prepared For: National Park Service, Landcover Mapping Program, National Park Service-Alaska Support Office, Anchorage, AK 99501. Chapin, F.S., Oswood, M.W., Van Cleve, K., Viereck, L.A. and Verblya, D.L. (eds.) 2006. Alaska’s Changing Boreal Forest. Oxford University Press, NY. 354 pp. Hanson, H.C. 1951. Characteristics of some grassland, marsh, and other plant communities in western Alaska. Ecol. Monogr. 21 (4): 317-378. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 44 of 204 *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 45 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316070 Western North American Boreal Subalpine Balsam Poplar-Aspen Woodland This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Mitch Michaud Modeler 2 Michelle Schuman 3/27/2008 mitch.michaud@ak.usd a.gov michelle.schuman@ak. usda.gov Reviewer Reviewer Reviewer Modeler 3 Map Zone 73 Vegetation Type Forest and Woodland Dominant Species* POBA2 POTR5 VIED ROAC Date General Model Sources ARCTO3 SALIX CACA4 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system occurs commonly throughout the mountain ranges of southcentral AK and also near the northern and western limit of the boreal region (NatureServe 2008). It occurs beyond the coniferous treeline in western and northern AK (NatureServe 2008). Biophysical Site Description The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): Western North American Boreal Subalpine Balsam Poplar-Aspen Woodland occurs on well-drained upland terrain on southerly aspects on upper slopes to treeline. In the upper elevation range of this system, it can occur in the subalpine zone above the coniferous treeline. Soils are generally well-drained, shallow, and develop on colluvial deposits, glacial till or bedrock. Vegetation Description Populus balsamifera ssp. balsamifera and/or P. tremuloides are the dominant overstory species. Trees are often stunted on exposed sties (Jorgenson et al. 2003). Plots from Denali National Park showed an average tree height of 2.4m for Dwarf Poplar Aspen Forest (Clark and Duffy 2006). Common understory shrubs include Viburnum edule, Rosa acicularis, Arctostaphylos spp. and Salix spp. A wide variety of herbaceous species may occur including Calamagrostis canadensis, Pyrola spp. and Aconitum delphinifolium (Viereck 1979, Jorgenson et al. 2003). Disturbance Description The disturbance dynamics of these high elevation forests are unclear but wind/exposure and fire could be *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 46 of 204 driving disturbance process. Balsam poplar and quaking aspen are fire adapted. Balsam poplar may be top killed by moderate intensity fires but can root sprout within several weeks following fire (Harris 1990). Aspen is also easily top killed by fire but fire stimulates vigorous root sprouting (Howard 1996). The probability of fire in the VDDT model is a best guess based on the assumption that this type would have a fire return interval similar to that of tundra types. Adjacency or Identification Concerns Native Uncharacteristic Conditions Scale Description Large patch or small patch Issues/Problems This model was defined at the Fairbanks workshop as having an additional early seral stage dominated by alpine tundra shrubs, herbaceous vegetation or bare ground. This class was described as a persistent state occurring on sites frequently disturbed by avalanches or other weather events (snow, ice, wind, etc). It was later determined that this stage could not be adequately modeled with the two tree dominated stages (class A and B in the current model) and that it was probably best to represent the non-forested sites as a different BpS. Comments This model was based on input from the experts who attended the LANDFIRE Fairbanks (Nov. 07) and Anchorage modeling meetings (Dec. 08) and refined by Mitch Michaud and Michelle Schuman. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 25 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position POBA2 POTR5 VIED ROAC Upper Upper Lower Lower Height Open (25-59% tree cover) Dwarf Tree (< 3 m) Seedling/Sapling <5" Tree Size Class Closed (60-100% tree cover) Tree (> 3 m) Upper layer lifeform differs from dominant lifeform. Description 0-49yrs This class is characterized by young stands of balsam poplar and/or aspen. Trees come in immediately via suckering in the post-disturbance stand along with the shrubs and herbaceous species. Wind desiccation can delay the development of trees. Succession to class B. Replacement MFRI = 200yrs resets age to zero. Wind/weather/stress (probability = 0.0067) maintains this class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 47 of 204 Class B Structure Data (for upper layer lifeform) Min Max 75 % Late Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position POBA2 POTR5 VIED ROAC Shrub Tree Open (25-59% tree cover) Closed (60-100% tree cover) Tree (> 3 m) Tree Size Class Upper Upper Lower Lower Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping purposes, if tree size class or image texture can't be used to distinguish class A and B, split them on height (A = trees <3m; B = trees >3m). Description 50yrs+ This class is characterized by mature balsam poplar and/or aspen. Mature stands can be more open than early seral stands (class A) but not always. Hardwoods begin to senesce after about 150yrs and a mixed age stand develops. This class persists in the absence of disturbance although it becomes even aged as hardwoods senesce. Mixed fire (MFRI = 150yrs) causes a transition to class A. Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 48 of 204 Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals III Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Avg FI Min FI Max FI Probability 715 205 0.0014 0.00488 159 0.00629 Percent of All Fires 22 78 Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Clark, M.H. and M.S. Duffy. 2006.Soil Survey of Denali National Park, Alaska. USDA NRCS. Available online at: http://soildatamart.nrcs.usda.gov/Manuscripts/AK651/0/DenaliPark.pdf. Harris, Holly T. 1990. Populus balsamifera subsp. balsamifera. In: Fire Effects Information System, [Online]. USDA Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/ [2008, March 27]. Howard, Janet L. 1996. Populus tremuloides. In: Fire Effects Information System, [Online]. USDA Forest Service, Rocky Mountain Research Station, Fire Sciences Laboratory (Producer). Available: http://www.fs.fed.us/database/feis/ [2008, March 27]. Jorgenson, M.T. et al. 2003. An ecological land survey for Fort Richardson, Alaska. Cold Regions Research and Engineering Laboratory, Hanover, New Hampshire, ERDC/CRREL TR-03019. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck, L.A. 1979. Characteristics of treeline plant communities in Alaska. Holarctic Ecology. 2: 228-238. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 49 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316080 Alaska Sub-boreal Avalanche Slope Shrubland This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Kori Blankenship kblankenship@tnc.org Modeler 2 Modeler 3 Upland Shrubland ALVIS SARA2 SALIX CACA4 4/15/2008 Reviewer Tina Boucher Reviewer Reviewer Map Zone 73 Vegetation Type Dominant Species* Date General Model Sources CHAN9 ATFI DREX2 Literature Local Data Expert Estimate antvb@uaa.alaska.edu Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system is found on steep mountain slopes in the sub-boreal region and less frequently in the boreal region of AK. Biophysical Site Description Avalanche slopes occur where mountain slopes are steep enough to produce frequent snow slides preventing forest development. Upper avalanche slopes typically have a slope angle of at least 70% but the lower slopes and run-out zones may be much less steep. Soils are shallow and stony, underlain by colluvium, glacial till and residuum (NatureServe 2008). Vegetation Description The dominant shrub species is typically Alnus viridis ssp. sinuata; but other shrubs including Sambucus racemosa, Salix spp. and Spirea stevenii may be common (NatureServe 2008). Herbaceous patches are often dominated by Calamagrostis canadensis and Chamerion angustifolium; other common herbs include Athyrium filix-femina, Dryopteris expansa and Veratrum viride (Viereck et al. 1992). Tree seedlings and saplings may be common on some slopes but do not emerge as an overstory due to frequent snow avalanches (NatureServe 2008). Disturbance Description Avalanche slopes can extend from the alpine into the montane and lower toe slopes; this model, however, applies only to avalanche slopes occurring below treeline. Snow avalanche is the dominant disturbance, but rocks, soil, and debris can also be transported in the slide. This system represents a topoedaphic climax (Viereck et al. 1992). Alnus viridis ssp. sinuata has a growth form that tolerates avalanche disturbance and can maintain dominance on the site (NatureServe 2008). Frequent snow slides prevent tree seedlings and saplings from reaching the upper canopy (NatureServe 2008). Fire is minimal and is *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 50 of 204 expressed only around the edges of this type as fires are introduced from neighboring types. Adjacency or Identification Concerns This system is similar in species composition to Alaska Sub-Boreal Mesic Subalpine Alder Shrubland, but it occurs below the subalpine zone and tree growth is limited by avalanche frequency rather than elevation as in the subalpine system (NatureServe 2008). Adjacent forest systems may include Alaska Sub-boreal Mountain Hemlock-White Spruce Forest, Alaska Sub-boreal Mountain Hemlock Forest or Alaska Subboreal White Spruce-Hardwood Forest. Though avalanche slopes can occur from alpine to lower slopes, this model excludes the avalanche slopes above treeline—these would be included in other systems according to vegetation type: Alaska Sub-Boreal Mesic Subalpine Alder Shrubland, Western North American Boreal Alpine Dwarf-Shrubland, Alaska Subboreal and Maritime Alpine Mesic Herbaceous Meadow, etc. Native Uncharacteristic Conditions Scale Description Small to large patch Issues/Problems No information is available on the MFRI for this type. The MFRI was estimated to be the same as the Alaska Sub-Boreal Mesic Subalpine Alder Shrubland BpS. Comments This model was developed by Kori Blankenship using the class age ranges from the Persistent Shrub North PNVG model (Murphy and Witten 2006) and a similar MFRI to the Alaska Sub-Boreal Mesic Subalpine Alder Shrubland BpS. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 70 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position CACA4 CHAN9 ATFI DREX2 Upper Upper Upper Upper Height Tree Size Class Herbaceous Herbaceous Seedling/Sapling <5" Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description 0-4yrs Forbs, shrubs and deciduous trees resprout immediately following disturbance, but herbaceous vegetation dominates in early succession. Herbaceous patches are often dominated by Calamagrostis canadensis and Chamerion angustifolium; other common herbs include Athyrium filix-femina, Dryopteris expansa and Veratrum viride (Viereck et al. 1992). Succession to class B. Avalanche (probability = 0.25) resets the age of this class. Replacement MFRI = 900yrs. Mixed MFRI = 5000yrs. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 51 of 204 Late Development 1 All Structures Structure Data (for upper layer lifeform) Min Max Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Class B 30 % Herbaceous Indicator Species* and Canopy Position ALVIS SALIX SARA2 SPST3 Shrub Tree Dwarf Shrub (< 20 cm) Tree Size Class Tall Shrub (>1.5 m) Pole 5–9" (swd)/5–11" (hwd) Upper Upper Upper Upper Upper layer lifeform differs from dominant lifeform. Description Five years plus Shrubs can become established at the edges or the bottom of the chutes. The dominant shrub species is typically Alnus viridis ssp. sinuata; but other shrubs including Sambucus racemosa, Salix spp. and Spiraea stevenii may be common. Tree seedlings and saplings may be common on some slopes but typically do not emerge as an overstory due to frequent snow avalanche. On sites where avalanche activity is less frequent trees can occasionally emerge temporarily in the overstory. This class persists in the absence of disturbance. Avalanche (probability = 0.25) resets the age of this class. Replacement MFRI = 900yrs. Mixed MFRI = 5000yrs. Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 52 of 204 Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals V Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Avg FI Min FI Max FI Probability 909.1 5000 0.0011 0.0002 769 0.00131 Percent of All Fires 84 15 Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Avalanche Other (optional 2) References Murphy, K.A. and E. Witten. 2006. Persistent Shrub North. In Fire Regime Condition Class (FRCC) Interagency Guidebook Reference Conditions. Available at www.frcc.gov. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 53 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316090 Alaska Sub-Boreal Mesic Subalpine Alder Shrubland This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Kori Blankenship kblankenship@tnc.org Modeler 2 Modeler 3 Upland Shrubland ALVIS SALIX SARA2 SPST3 4/11/2008 Reviewer Tina Boucher Reviewer Reviewer Map Zone 73 Vegetation Type Dominant Species* Date General Model Sources CACA4 CHAN9 ATFI DREX2 Literature Local Data Expert Estimate antvb@uaa.alaska.edu Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range Alaska Sub-boreal Mesic Subalpine Alder Shrubland is widespread on upper mountain slopes above treeline throughout the boreal transition region of AK (Viereck 1979). A riparian shrub variant of this system is found throughout the boreal and sub-boreal regions of AK in subalpine through alpine valleys. Biophysical Site Description These systems occur on well-drained mesic sites in the subalpine zone and in constrained riparian corridors on slopes in the alpine and subalpine. Seasonal overbank flooding may occur in the riparian areas, but generally it does not result in shifting channels or gravel bar formation (NatureServe 2008). Soils are shallow, stony and well-drained, underlain by colluvium, glacial till and residuum (NatureServe 2008). Vegetation Description This system often appears as a band of alder above treeline and below the alpine systems. Alnus viridis ssp. sinuata is the dominant shrub species, but other shrubs including Salix spp. (sometimes the dominant shrub), Sambucus racemosa and Spiraea stevenii may be common (NatureServe 2008). Herbaceous patches often occur within the shrub zone and may be dominated by Calamagrostis canadensis and Chamerion angustifolium; other common herbs include Athyrium filix-femina, Dryopteris expansa, Veratrum viride, Valeriana sitchensis, Lupinus nootkatensis and Sanguisorba sitchensis (Viereck et al. 1992). Disturbance Description This system represents a topoedaphic climax (Viereck et al. 1992). It occurs above treeline and is not controlled by avalanche activity, although avalanches may occur. Alder will resprout following fire but no studies exist on fire effects in this type. The fire return interval is likely long, possibly 500-1000yrs. Early *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 54 of 204 season fire prior to green-up would be more likely to carry than late season fire. Alder is also affected by insects and diseases (NatureServe 2008). Flooding and herbivory may be important disturbances, especially for the Alpine Riparian system (NatureServe 2008). Adjacency or Identification Concerns This system is similar in species composition to Alaska Sub-boreal Avalanche Slope Shrubland, but it occurs in the subalpine zone and tree growth is limited by elevation not avalanche frequency (NatureServe 2008). Pacific Maritime Tall Shrubland occupies a similar landscape position along the Gulf Coast of AK, but this system may be dominated by Rubus spectabilis, which does not occur in boreal regions. In the boreal transition region, the alder zone is intermixed with mesic herbaceous meadows (Calamagrosits canadensis and Chamerion angustifolium) (NatureServe 2008). This system includes riparian shrub types in the alpine and subalpine zones. Native Uncharacteristic Conditions This system may have been expanding further into the alpine in recent decades (NatureServe 2008). Scale Description Large patch Issues/Problems In the absence of data on MFRI for this system, the MFRI was estimated to be slightly lower than that in the FRCC Guidebook Persistent Shrub North model. Other major disturbances, including flooding, avalanches and herbivory, are not included in the model because it is believed that shrubs would resprout immediately following these disturbances. Comments This model was based on the FRCC Guidebook PNVG model for Persistent Shrub North (Murphy and Witten 2006). The MFRI was increased based on input from the experts who attended the LANDFIRE Anchorage (Dec. 07) modeling meeting. Tina Boucher reviewed an initial draft of this model and recommended eliminating mixed fire and decreasing the MFRI for Replacement Fire slightly so that the AllFire MFRI is slightly lower than that for Persistent Shrub North (these suggestions are reflected in the current draft of the model). Vegetation Classes *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 55 of 204 Class A Structure Data (for upper layer lifeform) Min Max 5% Early Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Tree Size Class None Herbaceous Shrub Tree Indicator Species* and Canopy Position CACA4 CHAN9 ATFI DREX2 Upper Upper Upper Upper Herbaceous Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description 0-4yrs Grasses, sedges and/or forbs dominate the site. Shrubs sprout from rootstock. Herbaceous patches often occur within the shrub zone and may be dominated by Calamagrostis canadensis and Chamerion angustifolium; other common herbs include Athyrium filix-femina, Dryopteris expansa, and Veratrum viride, Valeriana sitchensis, Lupinus nootkatensis and Sanguisorba sitchensis (Viereck et al. 1992). Succession to class B. Replacement MFRI = 2000yrs. Class B Structure Data (for upper layer lifeform) Min Max 95 % Mid Development 1 Closed Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position ALVIS SALIX SARA2 SPST3 Shrub Tree Dwarf Shrub (< 20 cm) None Tall Shrub (>1.5 m) Tree Size Class Upper Upper Upper Upper Upper layer lifeform differs from dominant lifeform. Description Five years plus Shrubs overtop herbaceous layer and become dominant. A low shrub and/or herbaceous layer usually persists. Shrub cover is 25-75%. Alnus viridis ssp. sinuata is the dominant shrub species, but other shrubs including Salix spp. (sometimes the dominant shrub), Sambucus racemosa and Spiraea stevenii may be common. This class persists in the absence of disturbance. Replacement MFRI = 830yrs. Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 56 of 204 Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Min Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals V Replacement Historical Fire Size (acres) Avg FI Min FI Max FI Probability 833 0.00120 832 0.00122 Percent of All Fires 98 Mixed Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References DeVelice, R.L., Hubbard, C.J., Boggs, K., Boudreau, S., Potkin, M., Boucher, T. Wertheim, C. 1999. Plant community types of the Chugach National Forest. Tech. Publ. R10-TP-76. Juneau, AK: USDA Forest Service, Alaska Region. 375 pp. Jorgenson, M.T. et al. 2003. An ecological land survey for Fort Richardson, Alaska. Cold Regions Research and Engineering Laboratory, Hanover, New Hampshire, ERDC/CRREL TR-03019. Murphy, K.A. and E. Witten. 2006. Persistent Shrub North. In Fire Regime Condition Class (FRCC) Interagency Guidebook Reference Conditions. Available at www.frcc.gov. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 57 of 204 NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Talbot, S.S, Talbot, S.L. and Daniels, F.J.A. 2005. Comparative phytosociological investigation of subalpine alder thickets in southwestern Alaska and the North Pacific. Phytocoenologia. 35(4): 727-759. Viereck, L.A. 1979. Characteristics of treeline plant communities in Alaska. Holarctic Ecology. 2: 228-238. Viereck, L.A., Dyrness, C.T. Batten, A.R. and Wenzlick, K.J. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 58 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316101 Western North American Boreal Mesic Scrub Birch-Willow Shrubland - Boreal This BPS is lumped with: This BPS is split into multiple models: Boreal Mesic Scrub Birch-Willow Shrubland was split into Boreal Mesic Scrub Birch-Willow Shrubland - Boreal and Boreal Mesic Scrub Birch-Willow Shrubland - Sub-boreal for BpS modeling so that a longer fire return interval could be applied to the sub-boreal variant. General Information Contributors (also see the Comments field Modeler 1 Jennifer Allen 4/4/2008 Jennifer_Allen@nps.go v Reviewer Lisa Saperstein Lisa_Saperstein@fws. gov Reviewer Reviewer Modeler 2 Modeler 3 Map Zone 73 Vegetation Type Upland Shrubland Dominant Species* BENA VAUL LEPAD SAPU15 Date General Model Sources SABA3 EMNIN VAVI HYSP70 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system is found in the boreal region from low elevations to the subalpine zone. Biophysical Site Description This system occurs on well-drained sites often in the subalpine. It is found on mesic sites on mid to upper slopes, above tree line and on terraces and sideslopes. Soils are mineral with a well-decomposed organic layer of 5-30cm thick (Viereck et al. 1992, NatureServe 2008). Vegetation Description Betula nana (this species now includes B. glandulosa) usually dominates the shrub layer, but Vaccinium uliginosum, Ledum decumbens (Ledum palustre L. ssp. Decumbens), Salix pulchra, S. barclayi, S. glauca or other Salix spp. may also be common or occasionally dominant (Viereck 1979, Viereck et al. 1992, NatureServe 2008). Dwarf shrubs such as Empetrum nigrum and Vaccinium vitis-idaea may be common under the low shrub layer. Herbaceous species are sparse, but may include Festuca altaica, Hierochloe alpina, Calamagrostis canadensis and Chamerion angustifolium. Feathermoss (Hylocomium splendens and Pleurozium schreberi) and lichens are common, but peat-forming mosses and sedges are not common (Viereck et al. 1992, NatureServe 2008). Disturbance Description This system represents a topoedaphic climax in some areas, in other cases it may be seral to shrub-tussock over long time periods (Viereck et al. 1992). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 59 of 204 There is little information available about the fire history of shrub communities in AK. Birch and ericaceous shrub tundra tends to produce more severe burns than sedge-shrub tussock tundra (Racine 1979). After fire, shrubs resprout readily from underground propagules if they have not been burned, and a shrub community re-establishes on the site within five years. After severe fires that remove the organic layer and burn the propagules, herbaceous species that establish by seed may dominate the site for more than five years. Burned-over spruce woodlands near treeline may be converted to low shrub after fire (Pegau 1972) and may slowly regenerate a spruce overstory. The fire return interval is longer in the subboreal region than in boreal AK. Adjacent vegetation influences the fire frequency. If the adjacent vegetation is flammable, then the low shrub type will have a more frequent fire return. Fire return intervals are long, likely greater than 100yrs. Trees may also invade these shrublands but over long time frames (NatureServe 2008). Adjacency or Identification Concerns At treeline, this system occurs above the Boreal Treeline White Spruce Woodland - Boreal (NatureServe 2008). Sites dominated by non-riparian or wetland Salix spp. are included in this type (NatureServe 2008). Low shrub types on peat deposits are included in the wetland types (NatureServe 2008). Native Uncharacteristic Conditions Scale Description Large patch Issues/Problems The probability for fire in the VDDT model is a best guess, not based on literature. Comments This model was based on input from the experts who attended the LANDFIRE Fairbanks (Nov. 07) modeling meeting and refined by Jennifer Allen. Vegetation Classes Class A 5% Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position FEAL HIAL3 CACA4 CHAN9 Upper Upper Upper Upper Cover Height Structure Data (for upper layer lifeform) Min Max Herbaceous Herbaceous Tree Size Class Herbaceous Herbaceous None Upper layer lifeform differs from dominant lifeform. Description 0-4yrs After fire, herbaceous species such as Festuca altaica and Hierochloe alpina may dominate, especially at higher altitudes. Calamagrostis canadensis and Chamerion angustifolium are common. Low shrubs can resprout following fire, quickly regaining dominance of a site. This class may persist for more than five years *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 60 of 204 if fire severity is high enough to remove the organic layer. Succession to class B. Replacement fire (MFRI = 200yrs) resets age to zero. Class B Structure Data (for upper layer lifeform) Min Max 95 % Late Development 1 All Structures Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position BENA VAUL LEPAD SALIX Shrub Tree Dwarf Shrub (< 20 cm) None Tall Shrub (>1.5 m) Tree Size Class Upper Upper Upper Upper Upper layer lifeform differs from dominant lifeform. Description Five years plus This class is dominated by shrubs, often Betula nana. Vaccinium uliginosum, Ledum decumbens, Salix pulchra, S. barclayi, S. glauca or other Salix spp. may also be common (Viereck 1979, Viereck et al. 1992). Dwarf shrubs such as Empetrum nigrum and Vaccinium vitis-idaea may be common under the low shrub layer. Some sites may include very low cover of trees, especially black spruce. This class persists in the absence of disturbance. Replacement MFRI = 200yrs causes a transition to A. Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Indicator Species* and Canopy Position Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 61 of 204 Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals IV Replacement Historical Fire Size (acres) Avg FI Min FI Max FI Probability 200 0.005 200 0.00502 Percent of All Fires 100 Mixed Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Anderson, J.H. 1974. Plants, Soils, Phytocenology and Primary Production of the Eagle Summit Tundra Biome Site. US Tundra Biome Data Report 74-42. Ecosystem Analysis Studies, US International Biological Program, US Arctic Research Program. 142 pp. Batten, A.R. 1977. The vascular floristics, major vegetation units, and phytogeography of the Lake Peters area, northeastern Alaska. M.S. thesis. University of Alaska, Fairbanks, AK. 330 pp. Batten, A.R., Murray, D.F.and Dawe, J.C. 1979. Threatened and Endangered Plants in selected areas of the BLM Fortymile Planning Unit.. File Report for Contract No. YA-512-CT8-162. USDI BLM AK State Office 701 C street, Anchorage, AK 99513. 127 pp. Hanson, H.C. 1951. Characteristics of some grassland, marsh, and other plant communities in western Alaska. Ecol. Monogr. 21 (4): 317-378. Hanson, H.C. 1953. Vegetation types in northwestern Alaska and comparisons with communities in other arctic regions. Ecology 34(1): 111-140. Hettinger, L.R. and A.J. Janz. 1974. Vegetation and soils of northeastern Alaska. Arct. Gas Biol. Rep. Ser. 21, 206 pp. North. Eng. Serv., Co., Ltd., Edmonton, Can. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 62 of 204 Hulten, E. 1966. Contributions to the knowledge of flora and vegetation of the southwestern Alaskan mainland. Sven. Bot. Tidskr. 60(1): 175-189. Jorgenson, M.T. 1984. The response of vegetation to landscape evolution on glacial till near Toolik Lake, Alaska. Pages 134-142 in Inventorying forest and other vegetation of the high latitude and high altitude regions: Proceedings of an International Symposium, Society of American Foresters Regional Technical Conference. Fairbanks, AK. Society of American Foresters, Bethesda, MD. Kessel, B., and G.B. Schaller. 1960. Birds of the upper Sheenjek Valley, northeastern Alaska. Biol. Pap. 4: 59 pp. Univ. Alaska, Fairbanks. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Pegau, R.E. 1968. Reindeer range appraisal in Alaska. M.S. thesis. University of Alaska, Fairbanks, AK. 130 pp. Pegau, R.E. 1972. Caribou investigations-analysis of range. In: Pegau, R.E. and J.E. Hemming (ed.). Caribou report. Volume 12. Progress report. Federal Aid in Wildlife Restoration, Projects W-17-2 and W17-3, Job 3.3R. Alaska Dept. of Fish and Game, Juneau, AK: 1-216. Racine. 1979. Climate of the Chucki-Imuruk area. Pages 32-37 in H. R. Melchior, ed., Biological Survey of the Bering Land Bridge National Monument. Alaska Cooperative Park Studies Unit, University of Alaska Fairbanks, Fairbanks, AK. Steigers, W.D. Jr., D. Helm and J.G. MacCracken. 1983. Alaska Power Authority, Susitna Hydroelectric Project, environmental studies- subtask 7.12: 1982 plant ecology studies. Final Report. University of Alaska, Agricultural Experiment Station, Palmer, AK. 288 pp. Viereck, L.A. 1962. Range survey: sheep and goat investigations. [Place of publication unknown]: [Publisher unknown]; completion report, W-6-R-3, Alaska work plan E, Job 2-A. 21 pp. Viereck, L.A. 1963. Sheep investigations: survey of range ecology. Project W-6-R-4, Work plan E. Job 2-A. Alaska Department of Fish and Game, Juneau, AK. Viereck, L.A. 1979. Characteristics of treeline plant communities in Alaska. Holarctic Ecology. 2: 228-238. Viereck, L.A., and Little, E.L. 1972. Alaska Trees and Shrubs. USDA Forest Service Ag. Handbook 410. University of Alaska Press, Fairbanks, Alaska. 265 pp. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. Webber, P.J., Komarkova, V., Walker, D.A. and E. Werbe. 1978. Vegetation mapping and response to disturbance along the Yukon River-Prudhoe Bay Haul Road. In: Brown, J. (principal investigator). Ecological baseline investigations along the Yukon River-Prudhoe Bay Haul Road, Alaska. Hanover, NH: Corps of Engineers, U.S. Army Cold Region Research and Engineering Laboratory: 25-87. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 63 of 204 Young, S.B. and Racine, C.H. 1978. Ecosystems of the proposed Katmai western extension, Bristol Bay lowlands, Alaska. Final Rep. Contributions from the Center for Northern Studies 15. Wolcott, VT: Center for Northern Studies. 94 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 64 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316102 Western North American Boreal Mesic Scrub Birch-Willow Shrubland - Alaska Sub-boreal This BPS is lumped with: This BPS is split into multiple models: Western North American Boreal Mesic Scrub Birch-Willow Shrubland was split into boreal and sub-boreal variants for BpS modeling so that a longer fire return interval could be applied to the sub-boreal variant. General Information Contributors (also see the Comments field Modeler 1 Kori Blankenship Modeler 2 Tina Boucher Modeler 3 kblankenship@tnc.org antvb@uaa.alaska.edu Reviewer Reviewer Map Zone 73 Upland Shrubland BENA VAUL LEPAD SAPU15 4/24/2008 Reviewer Vegetation Type Dominant Species* Date General Model Sources SABA3 EMNIN VAVI HYSP70 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system is found in the sub-boreal region from low elevations to the subalpine zone. This system is more common in the northern portion of the sub-boreal region (e.g., northern Cook Inlet); the tall shrub system dominated by Alnus viridis spp. sinuata replaces it in the southern portion of the region. Biophysical Site Description This system occurs on well-drained sites often in the subalpine. It is found on mesic sites on mid to upper slopes, above tree line and on terraces and sideslopes. Soils are mineral with a well-decomposed organic layer of 5-30cm thick (Viereck et al. 1992, NatureServe 2008). Vegetation Description Betula nana (this species now includes B. glandulosa) usually dominates the shrub layer; Vaccinium uliginosum, Ledum decumbens (Ledum palustre L. ssp. Decumbens), Salix pulchra, S. barclayi or other Salix spp. may also be common or occasionally dominant (Viereck 1979, Viereck et al. 1992, NatureServe 2008). Dwarf shrubs such as Empetrum nigrum and Vaccinium vitis-idaea may be common under the low shrub layer. Herbaceous species are sparse, but may include Festuca altaica and Hierochloe alpina. Feathermoss (Hylocomium splendens and Pleurozium schreberi) and lichens are common, but peat-forming mosses and sedges are not common (Viereck et al. 1992, NatureServe 2008). Disturbance Description The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): This system represents a topoedaphic climax in some areas, in other cases it may be seral to shrub-tussock *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 65 of 204 over long time periods (Viereck et al. 1992). There is little information available about the fire history of shrub communities in AK. Birch and ericaceous shrub tundra tends to produce more severe burns than sedge-shrub tussock tundra (Racine 1979). After fire, shrubs resprout readily from underground propagules if they have not been burned, and a shrub community re-establishes on the site within five years. After severe fires that remove the organic layer and burn the propagules, herbaceous species that establish by seed may dominate the site for more than five years. Burned-over spruce woodlands near treeline may be converted to low shrub after fire (Pegau 1972) and may slowly regenerate a spruce overstory. The fire return interval is longer in the subboreal region than in boreal AK. Adjacent vegetation influences the fire frequency. If the adjacent vegetation is flammable, then the low shrub type will have a more frequent fire return. Fire return intervals are long, likely greater than 100yrs. Trees may also invade these shrublands but over long time frames. Adjacency or Identification Concerns At treeline, this system occurs above the Western North American Boreal Treeline White Spruce Woodland - Alaska Sub-boreal (NatureServe 2008). Sites dominated by non riparian or wetland Salix spp. are included in this type (NatureServe 2008). Low shrub types on peat deposits are included in the wetland types (NatureServe 2008). Native Uncharacteristic Conditions Scale Description Large patch Issues/Problems The probability for fire in the VDDT model is a best guess, not based on literature. Comments This model was based on the Western North American Boreal Mesic Scrub Birch-Willow Shrubland Boreal model created by Jennifer Allen. Kori Blankenship and Tina Boucher increased the MFRI for the sub-boreal variant of the model and made minor edits to the description. Vegetation Classes Class A 5% Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position FEAL HIAL3 Upper Upper Cover Height Structure Data (for upper layer lifeform) Min Max Herbaceous Herbaceous Tree Size Class Herbaceous Herbaceous None Upper layer lifeform differs from dominant lifeform. Description 0-4yrs After fire, herbaceous species such as Festuca altaica and Hierochloe alpina typically dominate. This class may *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 66 of 204 persist for more than five years if fire severity is high enough to remove the organic layer. Succession to class B. Replacement MFRI = 300yrs resets age to zero. Class B Structure Data (for upper layer lifeform) Min Max 95 % Late Development 1 All Structures Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position BENA VAUL LEPAD SALIX Shrub Tree Dwarf Shrub (< 20 cm) None Tall Shrub (>1.5 m) Tree Size Class Upper Upper Upper Upper Upper layer lifeform differs from dominant lifeform. Description Five years plus This class is dominated by shrubs, often Betula nana. Betula glandulosa, Vaccinium uliginosum, Ledum decumbens, Salix pulchra, S. barclayi or other Salix spp. may also be common (Viereck 1979, Viereck et al. 1992). Dwarf shrubs such as Empetrum nigrum and Vaccinium vitis-idaea may be common under the low shrub layer. Trees may invade the shrubland over long time frames. This class persists in the absence of disturbance. Replacement MFRI = 300yrs cause a transition to A. Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Indicator Species* and Canopy Position Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 67 of 204 Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals V Replacement Historical Fire Size (acres) Avg FI Min FI Max FI Probability 300 0.00333 300 0.00335 Percent of All Fires 99 Mixed Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Hulten, E. 1966. Contributions to the knowledge of flora and vegetation of the southwestern Alaskan mainland. Sven. Bot. Tidskr. 60(1): 175-189. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Pegau, R.E. 1972. Caribou investigations-analysis of range. In: Pegau, R.E. and J.E. Hemming (ed.). Caribou report. Volume 12. Progress report. Federal Aid in Wildlife Restoration, Projects W-17-2 and W17-3, Job 3.3R. Alaska Dept. of Fish and Game, Juneau, AK: 1-216. Racine. 1979. Climate of the Chucki-Imuruk area. Pages 32-37 in H. R. Melchior, ed., Biological Survey of the Bering Land Bridge National Monument. Alaska Cooperative Park Studies Unit, University of Alaska Fairbanks, Fairbanks, AK. Viereck, L.A. 1963. Sheep investigations: survey of range ecology. Project W-6-R-4, Work plan E. Job 2-A. Alaska Department of Fish and Game, Juneau, AK. Viereck, L.A. 1979. Characteristics of treeline plant communities in Alaska. Holarctic Ecology. 2: 228-238. Viereck, L.A., and Little, E.L. 1972. Alaska Trees and Shrubs. USDA Forest Service Ag. Handbook 410. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 68 of 204 University of Alaska Press, Fairbanks, Alaska. 265 pp. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 69 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316110 Western North American Sub-boreal Mesic Bluejoint Meadow This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Tina Boucher antvb@uaa.alaska.edu colleen_ryan@tnc.org Modeler 2 Colleen Ryan Modeler 3 Upland Grassland/Herbaceous Dominant Species* 4/25/2008 Reviewer Reviewer Reviewer Vegetation Type General Model Sources CACA4 CHANA HEMA80 ATFI Date Literature Local Data Expert Estimate Map Zone 73 Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system occurs throughout the boreal and boreal transition regions of AK, though it appears to be less common north of the Alaska Range (NatureServe 2008). Biophysical Site Description Soils are typically fine-textured mineral and may be poorly drained (on flats) to well drained (on sideslopes). In the Sub-boreal region, mesic Calamagrostis canadensis meadows often occur near treeline interspersed with subalpine tall shrub (NatureServe 2008). Its elevational limit is just above the limit of tall shrub (within 100m). This system may also occur as a small patch in drained lake beds within the boreal region. Vegetation Description Calamagrostis canadensis is the species that characterizes this system, though other grasses, forbs and ferns may codominate. Fern- and forb-dominated patches commonly occur in a matrix of bluejoint meadow. Mosses are uncommon, but patchy feathermosses may be present in more open stands. Lichens and woody plants are absent or scarce, though this system is often found in a mosaic with tall shrub (especially alder) communities (Viereck et al. 1992). The vegetation is usually dense, with canopy height of 0.8-1.4m, occasionally reaching two meters (Viereck et al. 1992). Species composition ranges from nearly pure stands of Calamagrostis canadensis to mixtures of C. canadensis with forbs, ferns and other grasses, including Heracleum maximum, Angelica lucida, Chamerion angustifolium, Athyrium filix-femina, Dryopteris expansa, Equisetum arvense and Veratrum viride (Viereck et al 1992). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 70 of 204 Disturbance Description Though Mesic Bluejoint Meadows often occur on disturbed sites, the Sub-boreal Mesic Bluejoint Meadow system, as described here, is stable over long periods. The fire regime is likely to be similar to that of the alder tall shrub vegetation typically found just downslope from this system. However, the Sub-boreal Mesic Bluejoint Meadow system is extremely flammable in early summer prior to green-up. Fires in this time period generally do not burn the duff layer, since the ground is usually still wet or frozen). For this model, overall MFRI was estimated at approximately 800yrs. Avalanches are possible but are unlikely to affect the vegetation. Adjacency or Identification Concerns Mesic Bluejoint Meadow vegetation may also appear as a seral stage in a variety of other systems following fire or other disturbance. This system includes meadows that are long-term landscape features. This system is typically found just downslope from the Alaska Sub-boreal and Maritime Alpine Mesic Herbaceous Meadow or Alpine Dwarf-Shrubland system and upslope or interspersed with Alaska SubBoreal Mesic Subalpine Alder Shrubland. In some cases this system may be found directly adjacent to treeline. Native Uncharacteristic Conditions Scale Description Large patch Issues/Problems In the absence of data, the fire regime for this system was assumed to be similar to that of the Alaska SubBoreal Mesic Subalpine Alder Shrubland system often found adjacent to this system. Comments The fire regime for this system was based on the Alaska Sub-Boreal Mesic Subalpine Alder Shrubland model by Kori Blankenship. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 100 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position CACA4 CHANA2 HEMA80 ATFI Upper Upper Upper Upper Height Tree Size Class Herbaceous Herbaceous Herbaceous Herbaceous None Upper layer lifeform differs from dominant lifeform. Description Zero years plus This class represents the stable Mesic Bluejoint Meadow system. Calamagrostis canadensis and other grasses, forbs and/or ferns dominate the site. Replacement MFRI = 1000yrs. Mixed MFRI = 5000yrs. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 71 of 204 Class B Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Cover Indicator Species* and Canopy Position Height Tree Size Class Shrub Tree Upper layer lifeform differs from dominant lifeform. Description Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 72 of 204 Fire Regime Group**: Fire Intervals V Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 909.1 5000 0.0011 0.0002 769 0.00131 Percent of All Fires 84 15 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References DeVelice, R.L., Hubbard, C.J., Boggs, K. et al. 1999. Plant community types of the Chugach National Forest. Tech. Publ. R10-TP-76. Juneau, AK: USDA Forest Service, Alaska Region. 375 pp. Jorgenson, M.T. et al. 2003. An ecological land survey for Fort Richardson, Alaska. Cold Regions Research and Engineering Laboratory, Hanover, New Hampshire, ERDC/CRREL TR-03019. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 73 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316120 Western North American Boreal Dry Grassland This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Tina Boucher antvb@uaa.alaska.edu colleen_ryan@tnc.org Modeler 2 Colleen Ryan Modeler 3 Upland Grassland/Herbaceous Dominant Species* General Model Sources BRINA ARFR4 ARAL5 4/25/2008 Reviewer Reviewer Reviewer Vegetation Type FEAL FERU2 CAPU PSSP6 Date Literature Local Data Expert Estimate Map Zone 73 Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range Western North American Boreal Dry Grassland occurs across the boreal and sub-boreal regions of AK. Biophysical Site Description This system typically occurs on dry sideslopes or bluffs. Soils are well drained to excessively drained, and permafrost is absent. Some slopes may have steep, unstable soil (NatureServe 2008). Vegetation Description These sites are typically dominated by grasses, though forbs and shrubs may be common, but shrub cover is <25%. Common species include Festuca altaica, F. rubra, Calamagrostis purpurescens, Elymus inovatus, Artemisia frigida, Artemisia alaskana, Pseudoroegneria spicata, Bromus pumpellianus, Poa spp. and Achillea spp.( NatureServe 2008). Disturbance Description The fire regime for this system is expected to be similar to that of the adjacent Western North American Boreal Dry Aspen-Steppe Bluff - Higher Elevations. Because this system is often found on loose, dry mineral soils on unstable slopes, shifting slopes are an ongoing disturbance. Grazing is probably an important factor in shaping this system, as this is important Dall sheep habitat. Grazing and erosion were not included as disturbances in the model because these are long-term, ongoing phenomena. Adjacency or Identification Concerns Adjacent systems may include Western North American Boreal Dry Aspen-Steppe Bluff - Higher Elevations or Western North American Boreal Dry Aspen-Steppe Bluff - Lower Elevations. This system *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 74 of 204 does not encompass the coastal Leymus-forb meadows that occur in Aleutians, southwest, southcentral and southeast Alaska (NatureServe 2008). Native Uncharacteristic Conditions Scale Description Small to large patch. Issues/Problems Comments Fire intervals in this model were adapted from the Boreal Subalpine Steppe Bluff model, also by Boucher and Ryan. Suggested reviewers for this system include Carl Roland and Dalia Vargis-Kretzinger for information on grazing. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 100 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position FEAL FERU2 CAPU BRINA Upper Upper Upper Upper Herbaceous Height Herbaceous Tree Size Class None Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description Zero years plus This class represents the stable dry grassland community. Grasses, sedges and/or forbs dominate the site. Replacement MFRI = 200yrs. Mixed MFRI = 1000yrs. Class B Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 75 of 204 Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Min Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: III Historical Fire Size (acres) Avg 0 Min 0 Max 0 Sources of Fire Regime Data Literature Local Data Expert Estimate Fire Intervals Replacement Mixed Avg FI Min FI Max FI Probability 143.0 200.6 0.00699 0.00499 83 0.01199 Percent of All Fires 58 42 Surface All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 76 of 204 Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 77 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316141 Western North American Boreal Montane Floodplain Forest and Shrubland - Boreal This BPS is lumped with: This BPS is split into multiple models: Western North American Boreal Montane Floodplain Forest and Shrubland was split into a boreal and sub-boreal variant for BpS modeling so that regional differences could be represented. General Information Contributors (also see the Comments field Modeler 1 Michelle Schuman Modeler 2 Karen Murphy Modeler 3 Evie Witten Map Zone 73 Forest and Woodland Dominant Species* 4/7/2008 michelle.schuman@ak. Reviewer Tom Paragi usda.gov karen_a_murphy@fws.g Reviewer ov Reviewer ewitten@tnc.org Vegetation Type POBAB2 PIGL ALNUS SALIX Date General Model Sources SHCA ROAC VIED EQUIS Literature Local Data Expert Estimate tom.paragi@alaska.go v Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This type occurs on floodplains throughout the boreal region of AK. Biophysical Site Description The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): The substrate is typically well-drained sand or cobble, although finer silts and clays can be found on higher terraces, in ponds, on distal floodplains, and in lower energy systems. Permafrost is usually absent. Oxbows and other wet depressions commonly form on the floodplains, and these sites commonly develop into wetlands. Vegetation Description Primary succession on floodplains begins when new alluvial surfaces are colonized by tree, shrub and herbaceous species. Common early seral woody species include Populus balsamifera (seedlings), Picea glauca (seedlings), Alnus viridis ssp. sinuata, Alnus incana ssp. tenuifolia, Salix barclayi and Salix alaxensis (Boggs 2000, Scott 1974, Shephard 1995, Thilenius 1990, Viereck 1966). Common early seral herbaceous species may include Lupinus spp., Hedysarum spp. and Equisetum spp. (NatureServe 2008). The next seral stage includes communities dominated by Populus balsamifera and/or Picea glauca with an understory of Alnus viridis ssp. sinuata, Salix spp. and bryophytes. On dry sites the shrub layer may be dominated by Shepherdia canadensis, Dryas octopetala, D. integrifolia and fruticose lichens *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 78 of 204 (Steroucaulon spp.) (NatureServe 2008). The tall shrub component of the early successional stages diminishes rapidly, probably because of decreased light from the dense tree overstory. Populus balsamifera does not regenerate in the understory and consequently, Picea glauca gains dominance in the overstory within 150yrs (NatureServe 2008). Rosa acicularis and Viburnum edule are common understory shrubs on older surfaces. Disturbance Description The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): Flooding is the primary disturbance in this system. Flooding can be caused by snowmelt, precipitation, ice jams and glacial runoff. Different rivers or portions of rivers may be more prone to certain types of flooding. Frequent flooding and channel migration create a pattern of gravel bars and early successional stages across the valley bottom. Sediment deposition raises the surface of the floodplain over time. As the terrace becomes farther removed from the channel, flooding becomes less frequent. Water availability on terraces plays a major role in community structure and composition. Water inputs are from overbank flow (flooding), ground water and precipitation. Deposits with high permeability become progressively drier as they are vertically and horizontally removed from the active channels. Vegetation succession on gravel bars can be represented by the following seral stages: barren or herbaceous, willow or willow-alder, alder, poplar or spruce poplar, spruce. Oxbows and other wet depressions commonly form on the floodplains, and develop into wetlands. Succession and species composition is, however, variable due to diverse environmental conditions such as water depth, substrate and nutrient input. The following information was taken from the Riparian Spruce Hardwood Potential Natural Vegetation Group (PNVG) model (Murphy and Witten 2006): Estimates of mean fire return intervals include: -200yrs+ (200-300yr range) (Viereck 1973, Barney 1971) -300yrs (Rowe et al 1974) (for alluvial white spruce MacKenzie River Valley) -300yrs (Heinselman 1981) -300yrs (Duchesne and Hawkes 2000) -300yrs (personal communication experts’ workshop March 2004) Small, relatively infrequent, mixed severity fires characterize this PNVG due to the sites’ proximity to rivers, which act as fire breaks (Viereck 1973, Barney 1971, Foote 1983). High moisture content of the vegetation, high percentage of deciduous species, and high relative humidity also contribute to making fires less frequent in the Riparian Spruce Hardwood PNVG than in typically adjacent PNVGs. In interior AK the oldest white spruce stands (350yrs+) are commonly found on islands of floodplains where they are protected from fire (Viereck 1973). Adjacency or Identification Concerns This model applies to forest and shrub systems in the active and inactive portions of the floodplain, but not abandoned floodplains. Oxbows and other wet depressions commonly form on the floodplains. Floodplain Wetlands are a separate ecological system and a separate BpS. Native Uncharacteristic Conditions Scale Description Linear. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 79 of 204 Issues/Problems Wetlands that occur on the floodplain are not considered in this model. Comments This model was based on the FRCC Guidebook PNVG model for Riparian Spruce Hardwood (RSH; Murphy and Witten 2006) and input from the experts who attended the LANDFIRE Fairbanks (Nov. 07) modeling meeting. It was refined by Michelle Schuman. The relative age function used in the RSH model was not used in any class except A to comply with LANDFIRE modeling rules and the 10,000yr replacement fire was removed from class D. These changes did not change the model results. Because changes to the VDDT model were relatively minor Karen Murphy and Evie Witten were retained as modelers and Michelle Schuman's name was added. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 5% Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Open Shrub (25-74% shrub cover) Open Shrub (25-74% shrub cover) Indicator Species* and Canopy Position LUPIN HEDYS EQUIS SALIX Upper Upper Upper Upper Height Dwarf Shrub (< 20 cm) Tree Size Class Tall Shrub (>1.5 m) Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Description 0-4yrs This class is characterized by post disturbance regeneration (herbs, shrub regeneration and seedlings). Silt is deposited on the inside of river meanders following flood events. Flooding deposits seeds which germinate and take root. Lupinus spp., Hedysarum spp., Equisetum spp. and Salix spp. colonize in the first year. Within five years Salix spp and balsam poplar seedlings are abundant. Plant cover is 1-2% first year. Shrub cover increases up to 40% by the fifth year, with a diverse herbaceous layer underneath. Occasionally white spruce will germinate in large numbers on mineral soil after flooding, resulting in a dense, even-aged stand and eventual succession to class E (via class B). Succession to class B. Flooding (probability = 0.05), represented by Option 1 in the model, resets the age of this class. Class B 20 % Mid Development 1 Closed Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position SALIX ALNUS POBAB2 ROAC Upper Upper Upper Lower Structure Data (for upper layer lifeform) Min Max Cover Closed Shrub (> 75% shrub cover) Closed Shrub (> 75% shrub cover) Height Tall Shrub (>1.5 m) Low Shrub (20 cm to 1.5 m) Tree Size Class Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 80 of 204 Description 5-29yrs Tall shrubs (Salix spp., Alnus spp.) and saplings with a closed canopy (>60%). Saplings may consist of balsam poplar with white spruce in the understory (succession to class C), or saplings may consist of pure, even-aged spruce (succession to class E). Saplings overtop shrubs at 20-40yrs, when shade-intolerant pioneer shrub species decline and shade-tolerant shrubs (Rosa acicularis (prickly rose), Viburnum edule (high bushcranberry) become more common and have a canopy cover of 10%. Succession to class C or E. The deterministic succession pathway is to class C. The alternate succession pathway to E (probability = 0.01) represents possibility that white spruce will germinate in large numbers on mineral soil after flooding, resulting in a dense, even-aged stand. Flooding (probability = 0.03), represented by Option 1 in the model, causes a transition to class A. Class C Structure Data (for upper layer lifeform) Min Max 40 % Mid Development 1 Open Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position POBAB2 PIGL ROAC VIED Closed (60-100% tree cover) Tree Size Class Upper Middle Lower Lower Closed (60-100% tree cover) Tree (> 3 m) Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping, absence of spruce in the overstory distinguishes class C from class E. Description 30-149yrs Closed balsam poplar forest. Balsam poplar is the dominant overstory species but white spruce is commonly in the understory. Shade-tolerant shrub species persist in the understory. If spruce is present, at approximately 100-150yrs the transition from balsam poplar to white spruce dominance begins (succession to class D). If white spruce is not present poplar persists, the stand ages and individual trees are lost to wind, disease or rot. Shrub cover commonly increases as the overstory canopy declines. Succession to class D. Flooding (probability = 0.01), represented by Option 1 in the model, causes a transition to class A. Mixed fire (MFRI = 400yrs) maintains this class. Structure Data (for upper layer lifeform) Class D 25 % Late Development 1 Open Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position PIGL ROAC VIED Upper Lower Lower Lower Min Max Open (25-59% tree cover) Open (25-59% tree cover) Height Tree Size Class Tree (> 3 m) Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. Description 150yrs+ *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 81 of 204 Open white spruce forest. Spruce gains dominance over poplar and a mixed age, open stand develops. If enough young spruce establishes as poplar declines, the canopy closes again (modeled as alternate succession to class E). Alternatively, the stand may remain open with shrubs in the understory. This class can persist in the absence of disturbance or follows an alternate succession pathway to E (probability = 0.005). Flooding (probability = 0.005), represented by Option 1 in the model, causes a transition to class A. Mixed fire (MFRI = 155yrs) causes a transition to class C. Class E Structure Data (for upper layer lifeform) 10 % Min Late Development 1 Closed Upper Layer Lifeform Cover Indicator Species* and Canopy Position Herbaceous Shrub Tree PIGL ROAC VIED Lower Lower Lower Lower Closed (60-100% tree cover) Tree (> 3 m) Height Tree Size Class Max Closed (60-100% tree cover) Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping, dominance of spruce in the overstory distinguishes class E from class C. Description 30yrs+ Closed white spruce forest. These stands may be even-aged (resulting from spruce establishment on mineral soil after a flood event (succession from class B) or mixed age (succession from class D). If succession is from class D, occasional mature balsam poplar may persist in the overstory. As the spruce canopy closes, feathermoss becomes dominant on the forest floor, reaching 80% cover. Rosa acicularis, Viburnum edule and Alnus spp. may be scattered in the stand. A low shrub and herb layer may also occupy the forest floor. This class will persist in the absence of disturbance. Flooding (probability = 0.002), represented by Option 1 in the model, causes a transition to class A. Mixed fire (MFRI = 200yrs) causes a transition to class D. Replacement fire (MFRI = 667yrs) causes a transition to class A. Insects/disease (probability = 0.02) can cause a transition to class D. Disturbances Fire Regime Group**: Fire Intervals V Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 5000 312 0.0002 0.00321 294 0.00342 Percent of All Fires 6 94 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Flooding Other (optional 2) *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 82 of 204 References Barney, R.J. 1971. Wildfires in Alaska – some historical and projected effects and aspects. In: Fire in the Northern Environment – a symposium [Fairbanks, Alaska]. P. 51-59. Boggs, K. 2000. Classification of community types, successional sequences, and landscapes of the Copper River Delta, Alaska. Gen Tech. Rep. PNW-GTR-469. Portland, OR: USDA Forest Service, Pacific Northwest Research Station. 244 pp. Duchesne L.C. and B.C.Hawkes. 2000. Fire in northern ecosystems. In: Brown, J.K. and J.K Smith (eds.) Wildland fire in ecosystems: effects of fire on flora. Gen. Tech. Rep. RMRS-GTR-42-vol 2. Ogdon, UT: USDA Forest Service, Rocky Mountain Research Station. 257 pp. Heinselman, M.L. 1981. Fire and succession in the conifer forests of northern North America. In: West, D.C., H.H. Shugart, and D.B. Botkin. Forest succession: concepts and application. Springer-Verlag, New York. Chapter 23. Murphy, K.A. and E. Witten. 2006. Riparian Spruce Hardwood. In Fire Regime Condition Class (FRCC) Interagency Guidebook Reference Conditions. Available at www.frcc.gov. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Personal communication experts’ workshop, March 2-4 2004. Fire Regime Condition Class (FRCC) interagency experts’ workshop to develop and review Potential Natural Vegetation (PNV) groups for Alaska. Anchorage, AK. Rowe, J.S., Bergsteinsson, J. L. , Padbury, G. A.,and R. Hermesh. 1974. Fire studies in the Mackenzie Valley. ALUR Report 73-74-61. Arctic Land Use Research Program, Department of Indian Affairs and Human Development, Ottawa, Canada. 123 pp. Scott, R.W. 1974. Successional patterns on moraines and outwash of the Frederika Glacier, Alaska. In: Bushnell, V.C.; Marcus, M.G., eds. Icefield ranges research project scientific results. New York: American Geographical Society (4): 319-329. Shephard, M.E. 1995. Plant community ecology and classification of the Yakutat Foreland, Alaska. R10-TP56. Thilenius, J.F. 1990. Woody plant succession on earthquake-uplifted coastal wetlands of the Copper River Delta, Alaska. Forest Ecology and Management 33/34: 439-462. Viereck L.A. 1966. Plant succession and soil development on gravel outwash on the Muldrow Glacier, Alaska. Ecological Monographs. 36(3): 181-199. Viereck, L.A. 1973. Ecological effects of river flooding and forest fires on permafrost in the taiga of Alaska. In: Permafrost - - The North American Contribution to the Second International Conference. National Academy of Sciences, Washington, DC. 60-67 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 83 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316142 Western North American Boreal Montane Floodplain Forest and Shrubland - Alaska Sub-boreal This BPS is lumped with: This BPS is split into multiple models: Western North American Boreal Montane Floodplain Forest and Shrubland was split into a boreal and sub-boreal variant for BpS modeling so that regional differences could be represented. General Information Contributors (also see the Comments field Modeler 1 Karen Murphy Modeler 2 Evie Witten Modeler 3 Kori Blankenship Map Zone 73 Forest and Woodland POBAB2 PIGL ALNUS SALIX 4/14/2008 karen_a_murphy@fws.g Reviewer Tina Boucher ov Reviewer ewitten@tnc.org Reviewer kblankenship@tnc.org Vegetation Type Dominant Species* Date Model Zone General Model Sources EQUIS ROAC VIED SHCA antvb@uaa.alaska.edu Literature Local Data Expert Estimate Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This BpS is found throughout the AK sub-boreal region adjacent to rivers. Biophysical Site Description The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): The substrate is typically well-drained sand or cobble, although finer silts and clays can be found on higher terraces, in ponds, on distal floodplains, and in lower energy systems. Permafrost is usually absent. Oxbows and other wet depressions commonly form on the floodplains, and these sites commonly develop into wetlands. Vegetation Description Primary succession on floodplains begins when new alluvial surfaces are colonized by tree, shrub and herbaceous species. Common early seral woody species include Populus balsamifera (seedlings), Picea glauca (seedlings), Alnus viridis ssp. sinuata, Alnus incana ssp. Tenuifolia, Salix barclayi and Salix alaxensis (Boggs 2000, Scott 1974, Shephard 1995, Thilenius 1990, Viereck 1966). Herbaceous species may include Equisetum spp., Chamerion latifolium, Lupinus spp. and Hedysarum spp. The next seral stage includes communities dominated by Populus balsamifera and/or Picea glauca with an understory of Alnus viridis ssp. sinuata, Salix spp. and bryophytes. On dry sites the shrub layer may be dominated by Shepherdia canadensis, Dryas octopetala, D. integrifolia and fruticose lichens (Stereocaulon spp.). The *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 84 of 204 tall shrub component of the early successional stages diminishes rapidly, probably because of decreased light from the dense tree overstory. Populus balsamifera does not regenerate in the understory and consequently, Picea glauca gains dominance in the overstory within 150yrs. Rosa acicularis and Viburnum edule are common understory shrubs on older surfaces. Disturbance Description Flooding is the primary disturbance in this BpS. Flooding can be caused by snowmelt, precipitation, ice jams and glacial runoff. Different rivers or portions of rivers may be more prone to certain types of flooding. Frequent flooding and channel migration create a pattern of gravel bars and early successional stages across the valley bottom. Sediment deposition raises the surface of the floodplain over time. As the terrace becomes farther removed from the channel, flooding becomes less frequent. Water availability on terraces plays a major role in community structure and composition. Water inputs are from overbank flow (flooding), ground water, and precipitation. Deposits with high permeability become progressively drier as they are vertically and horizontally removed from the active channels. As discussed in the Riparian Spruce Hardwood Kenai Potential Natural Vegetation Group (PNVG) model description (Murphy and Witten 2006), small, relatively infrequent, mixed severity fires characterize this system due to the sites’ proximity to rivers, which act as fire breaks (Viereck 1973, Barney 1971, Foote 1983). High moisture content of the vegetation, high percentage of deciduous species, and high relative humidity also contribute to making fires less frequent in this system than in typically adjacent upland vegetation types. This model includes two successional pathways: one with a hardwood stage that succeeds to an open spruce stand and the other in which spruce establishes on mineral soil following a flood. Either of these pathways could result in an open or closed mature spruce stand. The successional classes in this model were defined by canopy cover classes in order to make them mappable. As a result, the system may move back and forth between the open and closed successional classes regardless of age. Adjacency or Identification Concerns This model applies to forest and shrub systems in the active and inactive portions of the floodplain, but not abandoned floodplains. Oxbows and other wet depressions commonly form on the floodplains. Floodplain Wetlands are a separate ecological system and a separate BpS. Native Uncharacteristic Conditions Scale Description Linear. Issues/Problems Wetlands that occur on the floodplain are not considered in this model. Comments NOTE: 11/21/09: As a result of final QC for LANDFIRE National by Jennifer Long the max canopy closure in class B was changed from “Closed (60-100% tree cover)” to “Closed Shrub (> 75% shrub cover)” because according to LANDFIRE National rules height should only be designated for the upperlayer lifeform. This model was based on the FRCC Guidebook PNVG model for Riparian Spruce Hardwood Kenai (RSHK; Murphy and Witten 2006) and input from the experts who attended the LANDFIRE Anchorage *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 85 of 204 (Dec. 07) modeling meeting. The relative age function used in the RSHK model was not used in any class except A to comply with LANDFIRE modeling rules. Because changes to the VDDT model were relatively minor, Karen Murphy and Evie Witten were retained as modelers and Kori Blankenship's name was added. In the first draft of the model, Classes D and E were indistinguishable for mapping (because both could be open or closed canopy forest). Tina Boucher reviewed an early draft of this model and confirmed that both successional pathways could lead to either open or closed stands. To correct this, Colleen Ryan defined Class D as open and Class E as closed and added an alternative succession pathway from Class E to Class D. This is meant to represent the possibility that some closed stands could open up over time (after age 150). This change did not substantially change the class landscape percentages. Vegetation Classes Class A 5% Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position EQUIS SALIX ALNUS Upper Upper Upper Upper Structure Data (for upper layer lifeform) Min Max Cover Open Shrub (25-74% shrub cover) Open Shrub (25-74% shrub cover) Height Dwarf Shrub (< 20 cm) Tall Shrub (>1.5 m) Tree Size Class Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Description 0-4yrs Silt is deposited on the inside of river meanders following flood events. Flooding deposits seeds which germinate and take root. Equisetum spp. and Salix spp. colonize in the first year. Within five years Salix spp and balsam poplar seedlings are abundant. Plant cover is 1-2% first year. Shrub cover increases up to 40% by the fifth year, with a diverse herbaceous layer underneath. Occasionally white (or Lutz) spruce will germinate in large numbers on mineral soil after flooding, resulting in a dense, even-aged stand. Common woody species include Alnus viridis ssp. sinuata, Alnus incana ssp. tenuifolia, Salix barclayi and Salix alaxensis (Boggs 2000, Scott 1974, Shephard 1995, Thilenius 1990, Viereck 1966). Herbaceous species may include Equisetum spp., Chamerion latifolium, Lupinus spp. and Hedysarum spp. On dry sites the shrub layer may be dominated by Shepherdia canadensis, Dryas octopetala, D. integrifolia and fruticose lichens (Stereocaulon spp.). Succession to class B. Flooding (probability = 0.1), represented by Option 1 in the model, resets the age of this class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 86 of 204 Class B Structure Data (for upper layer lifeform) Min Max 15 % Mid Development 1 Closed Cover Closed Shrub (> 75% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position SALIX ALNUS ROAC VIED Shrub Tree Tall Shrub (>1.5 m) Low Shrub (20 cm to 1.5 m) Tree Size Class Upper Upper Upper Upper Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Description 5-29yrs This class is typically dominated by tall shrubs (Salix spp., Alnus spp.) and saplings with a closed canopy (>60%). Common woody species include Alnus viridis ssp. Sinuata, Alnus incana ssp. Tenuifolia, Salix barclayi, and Salix alaxensis. On dry sites the shrub layer may be dominated by Shepherdia canadensis, Dryas octopetala, D. integrifolia and fruticose lichens (Steroucaulon spp.). Saplings may consist of balsam poplar with white (or Lutz) spruce in the understory (succession to class C), or saplings may consist of pure, even-aged spruce (succession to class E). Saplings overtop shrubs at 20-40yrs, when shade-intolerant pioneer shrub species decline and shade-tolerant shrubs (Rosa acicularis(prickly rose), Viburnum edule (high bush cranberry)) become more common and have a canopy cover of 10%. Succession to class C or E. The deterministic succession pathway is to class C. The alternate succession pathway to E (probability = 0.01) represents possibility that white spruce will germinate in large numbers on mineral soil after flooding, resulting in a dense, even-aged stand. Flooding (probability = 0.03), represented by Option 1 in the model, causes a transition to class A. Class C Structure Data (for upper layer lifeform) Min Max 30 % Mid Development 1 Open Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position POBAB2 PIGL ROAC VIED Upper Middle Lower Lower Closed (60-100% tree cover) Tree Size Class Closed (60-100% tree cover) Tree (> 3 m) Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping, absence of spruce in the overstory distinguishes class C from class E. Description 30 –149yrs Balsam poplar is the dominant overstory species. White spruce (or Lutz) is commonly in the understory. Shade-tolerant shrub species persist in the understory. If spruce is present, at approximately 100-150yrs the transition from balsam poplar to white spruce dominance begins (succession to class D). If white spruce is not present poplar persists, the stand ages and individual trees are lost to wind, disease or rot. Shrub cover commonly increases as the overstory canopy declines. Succession to class D. Flooding (probability = 0.01), represented by Option 1 in the model, causes a transition to class A. Mixed fire (MFRI = 400yrs) maintains this class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 87 of 204 Structure Data (for upper layer lifeform) Class D 35 % Late Development 1 Open Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position PIGL ROAC VIED ALNUS Upper Lower Lower Lower Min Max Open (25-59% tree cover) Open (25-59% tree cover) Height Tree (> 3 m) Tree Size Class Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Description 150yrs+ Spruce gains dominance over poplar and a mixed age, open stand develops. If enough young spruce establishes as poplar declines, the canopy closes again (succession to class E). Alternatively, the stand may remain open with shrubs in the understory. This class can persist in the absence of disturbance or follows an alternate succession pathway to E (probability = 0.005). Flooding (probability = 0.005), represented by Option 1 in the model, causes a transition to class A. Replacement MFRI = 3000yrs. Mixed fire (MFRI = 800yrs) causes a transition to class C. Class E Structure Data (for upper layer lifeform) 15 % Min Late Development 1 Closed Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position PIGL ROAC VIED ALNUS Lower Lower Lower Lower Closed (60-100% tree cover) Tree (> 3 m) Height Tree Size Class Max Closed (60-100% tree cover) Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping, dominance of spruce in the overstory distinguishes class E from class C. Description 30yrs+ This class contains closed stands of white (or Lutz) spruce. These stands may be even-aged (resulting from spruce establishment on mineral soil after a flood event (succession from class A) or mixed age (succession from class D). If succession is from class D, occasional mature balsam poplar may persist in the overstory. As the spruce canopy closes feathermoss becomes dominant on the forest floor, reaching 80% cover. Rosa acicularis, Viburnum edule and Alnus spp. may be scattered in the stand. A low shrub and herb layer may also occupy the forest floor. This class may persist in the absence of disturbance, or the canopy may open up as the stand matures on some sites, causing a transition back to class D. This class may persist in the absence of disturbance or may follow an alternate succession pathway to class D after age 150yrs (probability =.005). Flooding (probability = 0.002), represented by Option 1 in the model, causes a transition to class A. Replacement MFRI = 3000yrs. Mixed fire (MFRI = 800yrs) causes a transition to class D. Insects/disease (probability = 0.02) can cause a transition to class D. Disturbances *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 88 of 204 Fire Regime Group**: Fire Intervals V Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 5000 769 0.0002 0.00130 666 0.00151 Percent of All Fires 13 86 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Flooding Other (optional 2) References Barney, R.J. 1971. Wildfires in Alaska – some historical and projected effects and aspects. In: Fire in the Northern Environment – a symposium [Fairbanks, AK]. P. 51-59. Boggs, K. 2000. Classification of community types, successional sequences, and landscapes of the Copper River Delta, Alaska. Gen Tech. Rep. PNW-GTR-469. Portland, OR: USDA Forest Service, Pacific Northwest Research Station. 244 pp. Foote, J.M. 1983. Classification, description, and dynamics of plant communities after fire in the taiga of Interior Alaska. Res. Pap. PNW-307. Portland, OR. USDA Forest Service. Pacific Northwest Research Station. 108 pp. Murphy, K.A. and E. Witten. 2006. Riparian Spruce Hardwood Kenai. In Fire Regime Condition Class (FRCC) Interagency Guidebook Reference Conditions. Available at www.frcc.gov. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Scott, R.W. 1974. Successional patterns on moraines and outwash of the Frederika Glacier, Alaska. In: Bushnell, V.C.; Marcus, M.G., eds. Icefield ranges research project scientific results. New York: American Geographical Society (4): 319-329. Shephard, M.E. 1995. Plant community ecology and classification of the Yakutat Foreland, Alaska. R10-TP56. Thilenius, J.F. 1990. Woody plant succession on earthquake-uplifted coastal wetlands of the Copper River Delta, Alaska. Forest Ecology and Management 33/34: 439-462. Viereck, L.A. 1966. Plant succession and soil development on gravel outwash on the Muldrow Glacier, Alaska. Ecological Monographs. 36(3):181-199. Viereck, L.A. 1973. Ecological effects of river flooding and forest fires on permafrost in the taiga of Alaska. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 89 of 204 In: Permafrost - - The North American Contribution to the Second International Conference. National Academy of Sciences, Washington, DC. 60-67 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 90 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316150 Western North American Boreal Lowland Large River Floodplain Forest and Shrubland This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Date 3/11/2008 Modeler 1 Kori Blankenship kblankenship@tnc.org Modeler 2 Robert Lambrecht Robert_Lambrecht@fws Reviewer .gov tom.paragi@alaska.go v Reviewer Modeler 3 Map Zone 73 Vegetation Type Forest and Woodland Dominant Species* PIGL POBAB2 SALIX ALNUS Reviewer Tom Paragi General Model Sources ROAC VIED LUPIN EQUIS Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This type is found in the AK boreal region associated with high volume interior rivers such as the Yukon, Kuskokwim, Koyukuk and Tanana. Biophysical Site Description This system includes large floodplains associated with high volume interior rivers. The flooding regime is characterized by large spring floods at break-up caused by ice jams or summer floods caused by extreme rain events, the latter sometimes in combination with glacial melt. The active flooding zone often can be several kilometers wide (NatureServe 2008). Permafrost is usually absent, but can be present in small lenses. Ice scouring and ice dams are important dynamics that result in regeneration of willow carrs on the active floodplain and less frequently on higher terraces (NatureServe 2008). Wetland development in abandoned channels is intermixed with succession on more mesic sites (NatureServe 2008). Vegetation Description Primary succession on floodplains begins when new alluvial surfaces are colonized by tree, shrub and herbaceous species. Common woody species include Populus balsamifera, Picea glauca, Alnus viridis ssp. sinuata, Alnus incana ssp. tenuifolia, Salix barclayi and Salix alaxensis (Boggs 2000, Scott 1974, Shephard 1995, Thilenius 1990, Viereck 1966). Common early seral herbaceous species may include Lupinus spp., Hedysarum spp. and Equisetum spp. The next seral stage includes communities dominated by Populus balsamifera and/or (less commonly) Picea glauca with an understory of Alnus viridis ssp. sinuata, Salix spp. and bryophytes or uniform stands of Salix alaxensis. On dry sites the shrub layer may be dominated by Dryas octopetala, D. integrifolia and fruticose lichens (Steroucaulon spp.). The tall shrub *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 91 of 204 component of the early successional stages diminishes rapidly because of decreased light from the dense tree overstory and high levels of herbivory by moose or snowshoe hares may accelerate succession (Butler et al. 2007). Populus balsamifera does not regenerate in the understory and consequently, Picea glauca gains dominance in the overstory within 150yrs. Rosa acicularis and Viburnum edule are common understory shrubs in semi-open older stands. Disturbance Description Flooding can be caused by snow melt, precipitation, ice jams and glacial runoff. Different rivers or portions of rivers may be more prone to certain types of flooding. Frequent flooding and channel migration create a pattern of gravel bars and early successional stages across the valley bottom. Sediment deposition raises the surface of the floodplain over time. As the terrace becomes farther removed from the channel, flooding becomes less frequent. Water availability on terraces plays a major role in community structure and composition. Water inputs are from overbank flow (flooding), ground water, and precipitation. Deposits with high permeability become progressively drier as they are vertically and horizontally removed from the active channels. Oxbows and other wet depressions commonly form on the floodplains, and develop into wetlands. Succession and species composition is variable due to diverse environmental conditions such as water depth, substrate and nutrient input. Aquatic bed, marsh and fen communities are common. Fire frequency in floodplain systems is considerably less than that of the surrounding terrain because channels can act as fuel breaks. Early seral vegetation is less flammable than mature boreal forest, which may develop an organic soil layer that can spread fire into the floodplain in dry years. Fires burn in an irregular pattern due to the variability of vegetation and soil moisture, resulting in a high degree of edge. Adjacency or Identification Concerns This system is found adjacent to a range of upland systems, including hardwoods (birch-dominated), upland white spruce, white spruce-hardwood, as well as all boreal black spruce types. Native Uncharacteristic Conditions Scale Description Linear. Issues/Problems Frequency of major flood events on the larger rivers seems to be lower compared with the middle 20th Century, as evident from the age of Salix alaxensis cohorts on higher terraces. With the advent of climate change and changing weather patterns, historical ice scouring/jamming and flooding events are no longer common. The decline in floods that deposit a silt cap for willow and poplar regeneration and in ice scouring that rejuvenates willows could have a major impact on reducing the amount of available winter forage for moose, an important game species in interior AK. Comments This model was based on the FRCC Guidebook Potential Natural Vegetation Group (PNVG) model for Riparian Spruce Hardwood (RISH; Murphy and Witten 2006), input from the experts who attended the LANDFIRE Fairbanks (Nov. 07) modeling meeting and was refined by Kori Blankenship and Robert Lambrecht. The resulting model is similar to RISH but with minor adjustments to some of the class ages, the removal of replacement fire in class D, and the removal of the relative age setting for mixed fire in class B, which violated LANDFIRE modeling rules. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 92 of 204 The vegetation description is closely based on the Ecological System description (NatureServe 2008), with minor edits by Robert Lambrecht. Vegetation Classes Class A 5% Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position EQUIS SALIX POBAB2 LUPIN Upper Upper Upper Upper Structure Data (for upper layer lifeform) Min Max Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Dwarf Shrub (< 20 cm) Low Shrub (20 cm to 1.5 m) Height Tree Size Class Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Description 0-4yrs Post disturbance regeneration. This class can include gravel bar, herbs, shrub regeneration and seedlings. Silt is deposited on the inside of river meanders following flood events, although it can occur on higher terraces. Flooding deposits seeds which germinate and take root. Equisetum spp. and Salix spp. colonize in the first year. Within five years Salix spp. and balsam poplar seedlings are abundant. Plant cover is 1-20% in the first year. Shrub cover increases up to 40% by the fifth year, with a diverse herbaceous layer underneath. Lupinus and Hedysarum are common herbaceous species in this stage. Succession to class B. Flooding (probability = 0.05), represented by Option 1 in the model, resets the age of this class. Class B 10 % Mid Development 1 Closed Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position SALIX ALNUS POBAB2 ROAC Upper Upper Upper Upper Structure Data (for upper layer lifeform) Min Max Cover Closed Shrub (> 75% shrub cover) Closed Shrub (> 75% shrub cover) Height Tall Shrub (>1.5 m) Low Shrub (20 cm to 1.5 m) Tree Size Class Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Description 5-19yrs Tall shrubs (Salix spp., Alnus spp., Populus balsamifera) and saplings with a closed canopy (>60%). Saplings may consist of Salix alaxensis (dominant) or balsam poplar, with white spruce in the understory (succession to class C), or saplings may consist of pure, even-aged spruce (succession to class E). Saplings overtop shrubs at 15-40yrs, when shade-intolerant pioneer shrub species decline and shade-tolerant shrubs (Rosa acicularis, Viburnum edule) become more common and have a canopy cover of 10%. Uncommonly, white spruce may germinate in large numbers on mineral soil after flooding, resulting in a dense, even-aged stand. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 93 of 204 Succession to class C or E. The deterministic succession pathway is to class C. The alternate succession pathway to E (probability = 0.01) represents the possibility that white spruce will germinate in large numbers on mineral soil after flooding, resulting in a dense, even-aged stand. Flooding (probability = 0.03), represented by Option 1 in the model, causes a transition to class A. Class C Structure Data (for upper layer lifeform) Min Max 50 % Mid Development 2 All Structures Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position POBAB2 PIGL ROAC VIED Open (25-59% tree cover) Tree Size Class Upper Middle Lower Lower Closed (60-100% tree cover) Tree (> 3 m) Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Presence of hardwoods distinguishes class C from classes D and E. Description 20-149yrs Balsam poplar is the dominant overstory species, though white spruce may co-dominate. White spruce is commonly in the understory. Shade-tolerant shrub species persist in the understory. If spruce is present, at approximately 100-150yrs the transition from balsam poplar to white spruce dominance begins (succession to class D). If white spruce is not present, poplar persists, the stand ages and individual trees are lost to wind, disease or rot. Shrub cover commonly increases as the overstory canopy declines. Stands tend to be closed but can be open depending on site conditions. Succession to class D. Flooding (probability = 0.01), represented by Option 1 in the model, causes a transition to class A. Mixed fire (MFRI = 400yrs) maintains this class. Structure Data (for upper layer lifeform) Class D 25 % Late Development 1 Open Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position PIGL ROAC VIED ALNUS Upper Lower Lower Lower Cover Height Min Open (25-59% tree cover) Tree Size Class Max Open (25-59% tree cover) Tree (> 3 m) Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Description 150-400yrs Open white spruce. Spruce gains dominance over poplar and a mixed age, open stand develops. If enough young spruce establishes as poplar declines, the canopy closes again (succession to class E). Alternatively, the stand may remain open with shrubs in the understory. Feathermosses may dominate the forest floor. This class can persist in the absence of disturbance or follow an alternate succession pathway to E (probability = 0.005). Flooding (probability = 0.005), represented by Option 1 in the model, causes a transition to class A. Mixed fire (MFRI = 150yrs) causes a transition to class C. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 94 of 204 Class E Structure Data (for upper layer lifeform) 10 % Min Late Development 1 Closed Cover Upper Layer Lifeform Height Indicator Species* and Canopy Position Herbaceous Shrub Tree PIGL ROAC VIED ALNUS Upper Lower Lower Lower Closed (60-100% tree cover) Max Closed (60-100% tree cover) Tree (> 3 m) Tree Size Class Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Description 20-400yrs Closed white spruce. These stands can be even-aged (resulting from spruce establishment on mineral soil after a flood event (succession from class B) or mixed age (succession from class D). If succession is from class D, occasional mature balsam poplar may persist in the overstory. As the spruce canopy closes feathermoss becomes dominant on the forest floor, reaching 80% cover. Rosa acicularis, Viburnum edule and Alnus spp. may be scattered in the stand. A low shrub and herb layer may also occupy the forest floor. In the absence of disturbance, this class is self-replacing, with single tree or larger openings filled in by white spruce seedlings established in the understory. If the class begins as an even-aged stand, it will become uneven-aged over time. This class will persist in the absence of disturbance. Flooding (probability = 0.002), represented by Option 1 in the model, causes a transition to class A. Mixed fire (MFRI = 200yrs) causes a transition to class D. Replacement fire (MFRI = 667yrs) causes a transition to class A. Insects/disease (probability = 0.02) can cause a transition to class D. Disturbances Fire Regime Group**: Fire Intervals V Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Avg FI Min FI Max FI Probability 9999 303 0.00010 0.00330 294 0.00341 Percent of All Fires 3 97 Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Flooding Other (optional 2) References Boggs, K. 2000. Classification of community types, successional sequences, and landscapes of the Copper River Delta, Alaska. Gen Tech. Rep. PNW-GTR-469. Portland, OR: USDA Forest Service, Pacific Northwest Research Station. 244 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 95 of 204 Butler, L.G., K. Kielland, T.S. Rupp and T.A. Hanley. 2007. Interactive controls of herbivory and fluvial dynamics over vegetation patterns along the Tanana River, interior Alaska. J. Biogeography 34: 1622-1631. Murphy, K.A. and E. Witten. 2006. Riparian Spruce Hardwood. In Fire Regime Condition Class (FRCC) Interagency Guidebook Reference Conditions. Available at www.frcc.gov. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Scott, R.W. 1974. Successional patterns on moraines and outwash of the Frederika Glacier, Alaska. In: Bushnell, V.C.; Marcus, M.G., eds. Icefield ranges research project scientific results. New York: American Geographical Society (4): 319-329. Shephard, M.E. 1995. Plant community ecology and classification of the Yakutat Foreland, Alaska. R10-TP56. Thilenius, J.F. 1990. Woody plant succession on earthquake-uplifted coastal wetlands of the Copper River Delta, Alaska. Forest Ecology and Management 33/34: 439-462. Viereck L.A. 1966. Plant succession and soil development on gravel outwash on the Muldrow Glacier, Alaska. Ecological Monographs. 36(3): 181-199. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 96 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316160 Western North American Boreal Riparian Stringer Forest and Shrubland This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Nancy Fresco Modeler 2 Colleen Ryan Modeler 3 Douglas Hanson Reviewer Marc Lee marc.lee@alaska.gov Reviewer Tom Kurkowski douglas.hanson@alaska .gov Reviewer Tom Paragi thomas.kurkowski@al aska.gov tom.paragi@alaska.go v Map Zone 73 Wetlands/Riparian PIGL BEPA PIMA POBA2 4/7/2008 nancyfresco@hotmail.c om colleen_ryan@tnc.org Vegetation Type Dominant Species* Date General Model Sources POTR5 SALIX ALNUS Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system occurs along streams throughout the boreal and sub-boreal regions of AK. Biophysical Site Description The riparian forest and shrub stringer is found as a narrow band of vegetation along streams in low gradient and low volume drainages (NatureServe 2008). Although, seasonal overbank flooding may occur, it generally does not result in shifting channels or gravel bar formation (NatureServe 2008). The well-drained soils have moderately thick to thin organic horizons at the surface (indicative of infrequent flooding), are strongly acidic, lack permafrost, and have deep water tables (Jorgenson et al. 1999). Vegetation Description The mature phase of this system is dominated by Picea glauca, Betula papyrifera and Picea mariana. Populus balsamifera and Populus tremuloides are common early seral species on some sites. Picea mariana is more common and more likely to be dominant in the northern parts of this system’s range. Common understory species include Alnus tenuifolia, Rosa acicularis, Calamagrostis canadensis, Cornus canadensis, Equisetum arvense, Hylocomium splendens and Rhytidiadelphus triquetrus (Jorgenson et al. 1999). Immediately adjacent to the stream bank and in recently burned areas, tall shrubs including Alnus tenuifolia, Salix. alaxensis, S. bebbiana and S. lasiandra tend to dominate, with Carex spp. and Calamagrostis canadensis in the understory (NatureServe 2008, Jorgenson et al. 1999). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 97 of 204 Disturbance Description Fire is the primary disturbance influencing this system. However, riparian forest and shrub stringers typically burn less frequently than adjacent vegetation types and tend to act as fire breaks. Because of the linear nature of this system, fire frequencies are strongly influenced by surrounding vegetation types. In the sub-boreal region, surrounding vegetation is more likely to be white spruce and hardwood forest. In the northern boreal region, adjacent forest is more likely to be a black spruce type that burns more frequently. However, this system may also occur in tundra areas where there is a gap in the permafrost layer along the riparian corridor. The riparian stringer in this environment will have a very low fire frequency. As a result, fire frequencies are likely to be more variable in the boreal region compared with the sub-boreal region. For this model, the overall mean fire return interval was estimated at about 170yrs, somewhat longer than the MFRI for the white spruce-hardwood systems and substantially longer than the black spruce types. This reflects the observation that fires from the surrounding forest types frequently burn into the riparian stringer types, but often do not burn across the river or stream. Patchy, mixed severity fire is thought to be more common in this type than in most other spruce-hardwood types. Seasonal overbank flooding may occur, but it generally does not result in shifting channels or gravel bar formation (NatureServe 2008). As a result, flooding typically only acts as a disturbance in a narrow zone along the bank. This zone will typically remain in a self-replacing shrub stage. This disturbance is modeled as Option 1 in the VDDT model. Beaver activity may play a role in the dynamics of this system, but it is not included in the model due to the limited scale of these impacts. Adjacency or Identification Concerns See the Disturbance Description for more information on adjacent types. Native Uncharacteristic Conditions Scale Description Linear Issues/Problems The main factor controlling the disturbance regime in this system is its ecological setting. Ideally, we would create a separate model for this system on sites where the adjacent systems are non-forested. If these sites cannot be distinguished, then it is probably not worth creating separate models for the boreal and sub-boreal regions, as the MFRI is probably similar for the two regions. A reviewer noted that the age of transition from the shrub/sapling class to the forest class may vary widely depending on the tree species present. Hardwoods will begin to overtop the shrubs around age 15yrs, but this stage will be very difficult to distinguish from the shrub stage using satellite imagery. Spruce-dominated stands may not transition from class A to B for much longer, as it may take black spruce 30yrs to reach three meters in height. Classes B and C may overlap in canopy composition. For the purposes of this model, it has been assumed that class B will have >50% hardwoods in the canopy and class C will have >50% spruce, but there will be exceptions to this rule in reality. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 98 of 204 Comments The draft version of this model was created by Nancy Fresco and Colleen Ryan based in part on input from the experts who attended the LANDFIRE Anchorage (Dec. 07) modeling meeting. Extensive review by the Alaska Dept. of Natural Resources Div. of Forestry led to extensive changes to the model. As a result, lead reviewer Douglas Hanson was added as a modeler. The other Div. of Forestry reviewers, Marc Lee, Northern Region Forest Manager, and Tom Kurkowski, GIS specialist, were listed as reviewers. Will Putnam (wputman@tananachiefs.org) reviewed an early draft of this model. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 10 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Indicator Species* and Canopy Position SALIX ALNUS CAREX CACA4 Upper Upper Lower Low-Mi Height Dwarf Shrub (< 20 cm) Seedling/Sapling <5" Tall Shrub (>1.5 m) Tree Size Class Upper layer lifeform differs from dominant lifeform. Description 0-15yrs Shrub-sapling. This stage is dominated by shrubs but trees begin to sprout and saplings may be present. Around age 15yrs, the hardwood saplings will begin to overtop the shrubs, though the spruce saplings will still be in the understory. Common shrub species include a variety of Salix spp. and Alnus tenuifolia. Herbs frequently include Calamagrostis canadensis, Equisetum arvense and Carex spp. This class includes those areas immediately adjacent to the stream bank that are dominated by shrubs maintained by frequent flooding. These areas are unlikely to burn. The 0.1 annual probability of flooding in this class is meant to represent these areas. Succession is to class B at age 15yrs. Replacement MFRI = 333yrs. Flooding (Option 1) probability = 0.1. Class B Structure Data (for upper layer lifeform) Min Max 40 % Mid Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position BEPA PIGL PIMA ALNUS Upper Upper Upper Middle Woodland (10-24% tree cover) Closed (60-100% tree cover) Dwarf Tree (< 3 m) Tree Size Class Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping, the presence of hardwoods distinguishes class B from class C. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 99 of 204 Description 16-119yrs White spruce-hardwood mix. This is a mixed white spruce (black spruce) hardwood forest. Black spruce is more common and hardwoods are less common in the northern boreal region. Early in this age range, hardwoods will dominate the overstory. Spruce species typically will not overtop the shrub layer until age 3050yrs. By the end of this age range, the canopy will typically be dominated by a mixture of birch and spruce. Species composition is highly variable in this class, depending on landscape context, site characteristics, disturbance history and available seed sources. Picea glauca, Betula papyrifera, Populus balsamifera, Alnus spp. and Salix spp. typically dominate the overstory, though Picea mariana may be common, especially in the northern boreal region. The understory commonly includes Rosa acicularis, Carex spp. and Calamagrostis canadensis. Though species composition is highly variable, this class can be distinguished for mapping purposes by canopy composition (>50% hardwoods). Succession is to class B at age 120yrs. Replacement MFRI = 500yrs. Mixed MFRI = 220yrs (system stays in class B). Class C Structure Data (for upper layer lifeform) Min Max 50 % Late Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position PIGL BEPA POBA2 PIMA Upper Upper Upper Upper Woodland (10-24% tree cover) Closed (60-100% tree cover) Dwarf Tree (< 3 m) Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Tree Size Class Upper layer lifeform differs from dominant lifeform. This class may include some senescing hardwoods in the canopy, but the canopy should be at least 75% spruce. Description 120yrs+ Mature spruce forest. This class is typically dominated by mature spruce (>50% spruce in the canopy). More northern sites will have more black spruce, while more southern sites will be mostly white spruce. However, when these stands extend to the far north of the boreal region, such as the south slopes of the Brooks Range, white spruce will dominate over black spruce. Birch frequently persists in the canopy but typically will constitute <50% of the canopy. Understory plants include Alnus tenuifolia, Rosa acicularis, Cornus canadensis, Equisetum arvense, Hylocomium splendens and Rhytidiadelphus triquetrus (Jorgenson et al. 1999). This class persists in the absence of disturbance. Replacement MFRI = 400yrs. Mixed MFRI = 220yrs (system transitions to class B). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 100 of 204 Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Min Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals III Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 434.8 243.9 0.0023 0.0041 156 0.00641 Percent of All Fires 36 64 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Flooding Other (optional 2) References Foote, M. Joan. 1983. Classification, description, and dynamics of plant communities after fire in the Taiga of Interior Alaska. Res. Pap. PNW-307. Portland, OR: USDA Forest Service, Pacific Northwest Forest and Range Experiment Station. 108 pp. Jorgenson, M.T., J.E. Roth, M.K. Raynolds, M.D. Smith, W. Lentz, A.L. Zusi-Cobb and C.H. Racine. 1999. An ecological land survey for Fort Wainwright, Alaska. U.S. Army Cold Regions Research and Engineering Laboratory, Hanover, NH. CRREL TR-99-9. 92 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 101 of 204 Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 102 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316170 Western North American Boreal Shrub and Herbaceous Floodplain Wetland This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Date 5/7/2008 Modeler 1 Tina Boucher antvb@uaa.alaska.edu Reviewer Michelle Schuman michelle.schuman@a Modeler 2 Kori Blankenship Modeler 3 Colleen Ryan kblankenship@tnc.org colleen_ryan@tnc.org Reviewer k.usda.gov Map Zone 73 Vegetation Type Wetlands/Riparian Dominant Species* METR3 CAAQ EQFL COPA28 Reviewer General Model Sources ERAN6 MYGA BENA CHCA2 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system is found throughout the boreal and sub-boreal regions. It occurs within the floodplain boundaries of the Western North American Boreal Montane Floodplain Forest and Shrubland - Boreal, Western North American Boreal Lowland Large River Floodplain Forest and Shrubland and Western North American Boreal Montane Floodplain Forest and Shrubland - Alaska Sub-boreal. Biophysical Site Description The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): Floodplain wetlands occur within the active and inactive portions of floodplain systems. Wetlands develop on poorly drained deposits, oxbows and abandoned channels and are often mosaiced with welldrained floodplain vegetation. Vegetation Description Species composition is variable due to diverse environmental conditions such as water depth, substrate and nutrient input, as well as seral stage. Aquatic bed, marsh and fen communities are common. The earliest seral stages (aquatic bed) have aquatic vegetation such as Nuphar polysepala. Aquatic vegetation generally succeeds to marsh and fen communities with species such as Menyanthes trifoliata, Carex aquatilis and Equisetum fluviatile. Later seral stages of floodplain wetland development may include Sedge-Dwarf-Shrub Bogs and Low Shrub Peatland. Common species may include Sphagnum spp., Eriophorum angustifolium, Oxycoccus microcarpus, Andromeda polifolia, Myrica gale and Betula nana. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 103 of 204 Disturbance Description Fire is likely quite rare on floodplain wetlands and was not included in the VDDT model. The following paragraph was taken from the draft North Pacific Maritime Ecological Systems description (NatureServe 2008): Frequent river channel migration and associated flooding and fluvial processes constitute the major disturbances. Floodplain wetland dynamics differ from wetlands in non-floodplain settings. Surface flooding and subsurface flow provide a more continuous flow of nutrients and create the potential for a more dynamic wetland environment. Wetland succession and species composition are variable due to diverse environmental conditions such as water depth, substrate, and nutrient input. Common seral stages include aquatic bed, freshwater marsh, fen and wet low shrub. Wetland succession beginning in open water can proceed through aquatic bed and marsh communities to fens and wet meadows. Over time, fens can succeed to Sedge-Dwarf-Shrub Bogs or Low Shrub Peatlands. At any stage in succession, flooding can set back the vegetation to open water. Less dramatic changes in hydrology (such as an increase in water table from beaver activity) can reverse the direction of succession. Though an open water stage is part of the successional pathway, it is not included in the model due to LANDFIRE mapping constraints. Floodplain wetlands can also develop through paludification on poorly drained, fine-textured deposits. In this pathway, poorly-drained shrubs would accumulate organic material in the ground layer and succeed to a fen/wet meadow or Sedge-Dwarf-Shrub bog. Adjacency or Identification Concerns Floodplain wetland vegetation includes the following classes: Freshwater Aquatic Bed, Freshwater Emergent Marsh, Wet Meadow or Herbaceous Fen, Low Shrub Peatland, and Sedge-Dwarf-Shrub Bog. These classes have also been described as unique Ecological Systems, but because floodplain wetland dynamics are different from wetland dynamics outside the floodplain, they are considered a distinct BpS and modeled accordingly. Native Uncharacteristic Conditions Scale Description Floodplain wetland vegetation tends to occur in small, linear patches. Fire is rare (not modeled) but if it occurred would likely occur in very small patches moving into the floodplain from adjacent upland systems. Issues/Problems The lack of information about the successional dynamics of this system and the variability of environmental conditions across floodplain wetlands make it difficult to quantitatively model this BpS. The class percents also likely vary from site to site making it difficult to define a historic range of variability for FRCC calculation. In a true model of ecological dynamics, class A would include areas of sparse and submerged vegetation and open water. Since these areas will be excluded from the LANDFIRE mapping process, they were excluded from the model, and the landscape percentages for the successional classes reflect only the areas that are expected to map as vegetated ( >10% cover). Comments This model was based on input from the experts who attended the LANDFIRE Fairbanks modeling meeting (Nov. 07) and refined by Tina Boucher, Kori Blankenship and Colleen Ryan. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 104 of 204 Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 20 % Early Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Tree Size Class None Herbaceous Shrub Tree Indicator Species* and Canopy Position NULUP POTAM SPARG RATR Upper Upper Upper Upper Herbaceous Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description Zero years plus Freshwater Aquatic Bed This is an early seral stage dominated by open water and aquatic vegetation occurring in oxbows, abandoned channels, or other shallow slow-moving water within the floodplain. Common species include Nuphar polysepalum, Potamogeton spp., Lemna minor, Sparganium spp., Ranunculus trichophyllus and Myriophyllum spp. Aquatic mosses such as Calliergon sarmentosum may also be present (Viereck et al. 1992). This class can persist or follow an alternate succession pathway to class B (probability = 0.15). Flooding (probability = 0.07) can reset the age of this class to zero. Class B Structure Data (for upper layer lifeform) Min Max 15 % Early Development 2 All Structures Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position METR3 EQFL TYLA COPA28 Upper Upper Upper Upper Tree Size Class Herbaceous Herbaceous None Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description One year plus Freshwater Emergent Marsh The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): Marsh systems within the floodplain are typically semi-permanently flooded or have seasonal flooding. Water is at or above the surface for most of the growing season (typically at least 10 cm above the surface). Soils are muck or mineral, and water is nutrient-rich. These systems are highly productive and have high rates of decomposition (National Wetlands Working Group 1997). Freshwater marsh vegetation is dominated by emergent vegetation such as Carex utriculata, Scirpus validus, Typha latifolia, Menyanthes trifoliata, Equisetum fluviatile and Comarum palustris (Gracz 2005, Viereck et al. 1992). Aquatic plants such as Potamogeton spp. and Myriophyllum spp. May also be common (Viereck et al. 1992). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 105 of 204 This class can persist or follow an alternate succession pathway to class C (probability = .1). Flooding (probability = 0.07) can cause a transition to class A. Class C Structure Data (for upper layer lifeform) Min Max 30 % Mid Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Tree Size Class None Herbaceous Shrub Tree Indicator Species* and Canopy Position METR3 CAAQ EQFL COPA28 Upper Upper Upper Upper Herbaceous Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description One year plus Wet Meadow or Herbaceous Fen The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): Fens are nutrient-rich and have a thick peat layer that may be floating or submerged. Standing water is usually present (Viereck et al. 1992). Dominant species may include Menyanthes trifoliata, Equisetum fluviatile, Comarum palustris, Calla palustris, Eriophorum angustifolium, and Carex aquatilis. Other common but non-dominant species include Caltha palustris, Cicuta makenzieana, Galium trifidum, Rumex arcticus and Utricularia spp., (Racine and Walters 1994). Standing water is often present. Shrubs, including Myrica gale, Salix candida, Betula nana and Alnus incana spp. tenuifolia, are occasionally present, but do not exceed 25% cover. Aquatic plants such as Myriophyllum spicatum, Hippuris vulgaris, Potamogeton spp. and Sparganium spp. may be present, and aquatic mosses are often present (Viereck et al. 1992). This system may be seral to poor fens and bogs. This class can persist or follow an alternate succession pathway to class D (probability = 0.02) or class E (probability = 0.005). Flooding (probability = 0.03) can cause a transition to class A. Structure Data (for upper layer lifeform) Class D 20 % Mid Development 2 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Min Max Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Indicator Species* and Canopy Position MYGA BENA CHCA2 SAPU15 Upper Upper Upper Upper Height Dwarf Shrub (< 20 cm) Tree Size Class Low Shrub (20 cm to 1.5 m) None Upper layer lifeform differs from dominant lifeform. Description One year plus Low Shrub Peatland The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): This mid to late seral stage includes low shrub-dominated wetlands within the floodplain. Common species *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 106 of 204 include Betula nana, Myrica gale, Ledum groenlandicum, Salix fuscescens, S. pulchra, Empetrum nigrum, Chamaedaphne calyculata and Sphagnum spp. Soils are saturated for at least a portion of the growing season. Sites may be bogs, fens, or wetlands with <40cm of peat. Myrica gale and Chamaedaphne calyculata are indicators of fen conditions. This class will persist in the absence of disturbance or follow an alternate succession pathway to class E (probability = 0.01). Flooding (probability = 0.02) can cause a transition to class A. Class E Structure Data (for upper layer lifeform) 15 % Min Late Development 1 All Structures Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree SPHAG2 VAOX ANPO ERAN6 Lower Upper Upper Upper Max Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Dwarf Shrub (< 20 cm) Low Shrub (20 cm to 1.5 m) Height Tree Size Class None Upper layer lifeform differs from dominant lifeform. Description One year plus Sphagnum peatlands, Sedge-Dwarf-Shrub Bog or poor fen. The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): This class is a late seral stage of the riparian wetland system. It includes bogs and poor fens with thick (>40cm) peat deposits. Organic soils are acidic and nutrient poor. Common species include Oxycoccus microcarpus, Andromeda polifolia, Rubus chamaemorus, Vaccinium uliginosum, Ledum groenlandicum, Betula nana, Empetrum nigrum, Carex microglochin, C. rotundata, C. rariflora, C. lasiocarpa,C. limosa, C. chordorrhiza, Eriophorum brachyantherum, E. angustifolium and Drosera spp. Sphagnum spp. are usually abundant in the ground layer. This class will persist in the absence of disturbance. Flooding (probability = 0.02) can cause a transition to class A. Disturbances Fire Regime Group**: NA Fire Intervals Avg FI Min FI Max FI Probability Percent of All Fires Replacement Historical Fire Size (acres) Avg 0 Min 0 Max 0 Sources of Fire Regime Data Literature Local Data Expert Estimate Mixed Surface All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 107 of 204 Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Flooding Other (optional 2) References Gracz, M., K. Noyes, P. North and G. Tande. 2005 Wetland Mapping and Classification of the Kenai Lowland, Alaska. http://www.kenaiwetlands.net/. National Wetlands Working Group. 1997. Wetlands of Canada. C.D.A Rubec (ed.). Ecological Land Classification Series No. 24. Environment Canada, Ottawa, and Polyscience Publications Inc., Montreal. 452 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for the Alaska North Pacific Maritime Region. Racine, C.H., and J.C. Walters. 1994. Groundwater-discharge fens in the Tanana Lowlands, Interior Alaska, USA. Arctic and Alpine Research 26: (4) 418-426. Viereck, L.A., Dyrness, C.T., Batten, A.R., Wenzlick, K.J. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 108 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316181 Western North American Boreal Herbaceous Fen - Alaska Sub-Boreal Complex This BPS is lumped with: see below This BPS is split into multiple models: This BpS was created by merging several Ecological Systems: Western North American Boreal Freshwater Emergent Marsh (in part) Western North American Boreal Herbaceous Fen Western North American Boreal Wet Meadow (in part) Western North American Boreal Low Shrub Peatland General Information Contributors (also see the Comments field Modeler 1 Torre Jorgenson Date 4/21/2008 tjorgenson@abrinc.com Reviewer Michelle Schuman michelle.schuman@a k.usda.gov Reviewer Reviewer Modeler 2 Modeler 3 Map Zone 73 Vegetation Type Wetlands/Riparian Dominant Species* MYGA BENA CHCA2 SAPU15 General Model Sources COPA28 METR3 EQFL CACA4 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system occurs throughout the sub-boreal region of AK and on sites without permafrost in the boreal region. Biophysical Site Description This model applies to sites with open hydrology and without permafrost in the Sub-boreal and Boreal regions. Typical sites include drainages and pond margins. In the early successional marsh stage, sites may be semi-permanently flooded, and seasonal flooding is common in the wetter classes. In later stages, soils are saturated for at least a portion of the growing season. An organic peat layer is usually present except in the marsh class, but peat depth is variable. Vegetation Description This system is a complex of four wetland types with varied characteristic vegetation, as follows. Freshwater Emergent Marsh Class: Freshwater marsh vegetation may be dominated by emergent sedges, forbs, or grasses. Species that often dominate or codominate include Carex utriculata, Scirpus validus, Arctophila fulva, Eleocharis palustris, Myriophyllum spicatum, Typha latifolia Menyanthes trifoliata, Comarum palustris, Hippuris vulgaris and Equisetum fluviatile (Gracz 2005, Jorgenson 1999). Arctophila fulva becomes more common in the northern portions of boreal Alaska (NatureServe 2008). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 109 of 204 Herbaceous Fen Class: Fens are nutrient-rich and have a thick peat layer (herbaceous peat or sphagnum peat) that may be floating or submerged. Common species include Menyanthes trifoliata, Potentilla palustris, Equisetum fluviatile, Potentilla palustris and Carex aquatilis (NatureServe 2008). Wet Sedge Meadow Class: Dominant species may include Carex aquatilis, C. livida, C. chordorrhiza, C. lasiocarpa, Eriophorum angustifolium, Calamagrostis canadensis, Comarum palustre, Menyanthes trifoliata, Equisetum fluviatile and E. palustre. Shrubs may be a minor component of the canopy cover and include Myrica gale, Alnus incana ssp. Tenuifolia and Salix spp. (NatureServe 2008). Low Shrub Peatland Class: Common species include Ledum palustre or L. groenlandicum, Betula nana, Rubus chamaemorus, Oxycoccus microcarpus, Myrica gale, Calamagrostis canadensis, Carex aquatilis, Comarum palustre, Salix fuscescens, S. pulchra, Empetrum nigrum and Chamaedaphne calyculata. Sphagnum spp. And/or brown mosses may be common in the ground layer (DeVelice et al. 1999, Jorgenson et al 2003). Disturbance Description The dominant factor governing the dynamics of this wetland complex system is hydrology. Changes in water level or frequency and duration of flooding can cause transitions among all four classes in this model. Each of the classes may be stable for long periods under appropriate hydrological conditions. A typical successional pathway may begin with open water or a freshwater aquatic bed (not included in this model because they are sparsely vegetated) and progress to marsh vegetation, possibly with some open water or floating vegetation. From there, the system typically moves to herbaceous fen vegetation, then to a wet sedge meadow and finally to low shrub peatland vegetation, as the site becomes drier. However, large increases in the frequency or severity of flooding may send any of these vegetation types back to the marsh system. Smaller increases in water levels may send the system back to wet meadow or fen vegetation. Hydrologic changes may also cause a site to transition out of this system entirely, as when a marsh or fen is cut off from any inflow of water, leading to paludification. This could move the site into the Boreal Sedge-Dwarf-Shrub Bog or Alaska Sub-Boreal Lowland Wet Black Spruce Complex model. Fire occurs very rarely in this system, though most of the classes may burn in very dry years. Fire effects are very complex in these wetland types. Adjacency or Identification Concerns Native Uncharacteristic Conditions Scale Description Small patch to large patch Issues/Problems Comments Vegetation Classes *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 110 of 204 Class A Structure Data (for upper layer lifeform) Min Max 5% Early Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Tree Size Class None Herbaceous Shrub Tree Indicator Species* and Canopy Position TYLA CARO6 SCTA2 Upper Middle Upper Herbaceous Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. For mapping, this class should be distinguished by existing vegetation type. Description Zero years plus Freshwater Marsh This is a freshwater marsh, typically including some open water and emergent vegetation dominated by cattails and Carex rostrata. Other emergent sedges, forbs, or grasses may dominate, and floating aquatic begetation may also be present. Other species that often dominate or codominate include Carex utriculata, Schoenoplectus tabernaemontani (syn. Scirpus validus), Arctophila fulva, Eleocharis palustris, Myriophyllum spicatum, Menyanthes trifoliata, Comarum palustris, Hippuris vulgaris and Equisetum fluviatile (Gracz 2005, Jorgenson 1999). Arctophila fulva becomes more common in the northern portions of boreal AK. This stage may be preceded by an open water/aquatic bed class (not included in this model due to mapping constraints). This class may replace itself under stable hydrologic conditions (frequent flooding). Under drying conditions, alternate succession may go to class B after two years (probability = 0.5). Class B Structure Data (for upper layer lifeform) Min Max 5% Mid Development 1 Open Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position METR3 Upper COPA28 Upper EQFL Upper Upper Tree Size Class Herbaceous Herbaceous None Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. For mapping, this class should be distinguished by existing vegetation type. Description Three years plus Herbaceous Fen This class includes fen vegetation, often with standing water. Common species include Menyanthes trifoliata, Comarum palustre (syn. Potentilla palustris), Equisetum fluviatile, Potentilla palustris and Carex aquatilis. This class may replace itself under stable hydrologic conditions (frequent flooding). Under drying conditions, *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 111 of 204 alternate succession may go to class C after age five years (probability = 0.15). Increased water level (modeled as Option 1; probability = 0.01) will set the system back to marsh (class A). Class C Structure Data (for upper layer lifeform) Min Max 40 % Late Development 1 Closed Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position Herbaceous Herbaceous Tree Size Class CAREX Upper ERAN6 Upper CACA4 Upper Herbaceous Herbaceous None Upper layer lifeform differs from dominant lifeform. Shrubs may be present in the canopy, but at <25% cover. For mapping, this class should be distinguished by existing vegetation type. Description 6+ years Wet Meadow This class represents a Wet Sedge Meadow type. Dominant species may include Carex aquatilis, C. livida, C. chordorrhiza, C. lasiocarpa, Eriophorum angustifolium, Calamagrostis canadensis, Comarum palustre, Menyanthes trifoliata, Equisetum fluviatile and E. palustre. Shrubs may be a minor component of the canopy cover and include Myrica gale, Alnus incana ssp. tenuifolia, and Salix spp. In this class, soils are saturated at some point during the growing season, but do not have standing water. Wet meadows typically have a well-developed organic mat but not deep enough to be considered peatlands (NatureServe 2008). This class may replace itself under stable hydrologic conditions (saturated soils). Under drying conditions, alternate succession may go to class D after age 25yrs (probability = 0.15). Increased water level (modeled as Option 1; probability = 0.01) will set the system back to marsh (class A). Class D 50 % Late Development 2 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position MYGA BENA CHCA2 SAPU15 Structure Data (for upper layer lifeform) Min Max Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Height Low Shrub (20 cm to 1.5 m) Tree Size Class Tall Shrub (>1.5 m) None Upper layer lifeform differs from dominant lifeform. For mapping, this class should be distinguished by existing vegetation type. Description 26yrs+ Low Shrub Peatland This class is characterized by Wet Low Shrub vegetation. Common species include Ledum palustre, L. groenlandicum, Betula nana, Rubus chamaemorus, Oxycoccus microcarpus, Myrica gale, Calamagrostis canadensis, Carex aquatilis, Comarum palustre, Salix fuscescens, S. pulchra, Empetrum nigrum and *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 112 of 204 Chamaedaphne calyculata. Sphagnum spp. and/or brown mosses may be common in the ground layer (DeVelice et al. 1999, Jorgenson et al 2003). Soils are saturated for at least a portion of the growing season, and a peat layer is usually present but variable in depth. This class is distinguished from the Boreal Low Shrub Peatland stage of the Boreal Lowland Wet Black Spruce Complex model by the increased dominance of Myrica gale, Chamaedaphne calyculata and Salix pulchra, and the decrease in Betula nana and ericaceous shrubs. This class maintains itself in the absence of disturbance. Substantial flooding/increase in water level (modeled as Option 1; probability = 0.01) will set the system back to marsh (class A). Minor flooding/increase in water level (modeled as Option 2; probability = 0.02) will set the system back to wet meadow (class C). Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals NA Avg FI Min FI Max FI Probability Percent of All Fires Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Native Grazing Wind/Weather/Stress Competition Other (optional 1) increase water table/flooding Other (optional 2) decreased water table References DeVelice, R.L., Hubbard, C.J., Boggs, K. et al. 1999. Plant community types of the Chugach National Forest. Tech. Publ. R10-TP-76. Juneau, AK: USDA Forest Service, Alaska Region. 375 pp. Gracz, M., Noyes, K, North, P. and Tande, G. 2005 Wetland Mapping and Classification of the Kenai Lowland, Alaska. http://www.kenaiwetlands.net/ *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 113 of 204 Jorgenson, M.T. et al. 2003. An ecological land survey for Fort Richardson, Alaska. Cold Regions Research and Engineering Laboratory, Hanover, New Hampshire, ERDC/CRREL TR-03019. National Wetlands Working Group. 1997. Wetlands of Canada. C.D.A Rubec (ed.). Ecological Land Classification Series No. 24. Environment Canada, Ottawa, and Polyscience Publications Inc., Montreal. 452 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 114 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316211 Western North American Boreal Black Spruce Dwarf-tree Peatland - Boreal Complex This BPS is lumped with: see below This BPS is split into multiple models: This system was created by merging several Ecological Systems: Western North American Boreal Black Spruce Dwarf-tree Peatland, Western North American Boreal Freshwater Emergent Marsh, Western North American Boreal Wet Meadow, Western North American Boreal Low Shrub Peatland and Western North American Boreal Black Spruce-Tamarack Fen Western North American Boreal Black Spruce-Tamarack Fen is lumped with Western North American Boreal Black Spruce Dwarf-tree Peatland because it occurs on similar sites and has a similar successional trajectory. The main difference is that the late seral stage would be dominated by Larix laricina but this is accounted for in the model by including both Picea mariana and Larix laricina in the later seral stage. Western North American Boreal Freshwater Emergent Marsh, Western North American Boreal Wet Meadow and Western North American Boreal Low Shrub Peatland are lumped as successional stages of this model when they occur on sites with permafrost dynamics. These three systems are also lumped into the Subboreal Fen Complex model where they occur on non-permafrost sites in the boreal or sub-boreal regions. General Information Contributors (also see the Comments field Date 4/4/2008 Modeler 1 Jill Johnstone jill.johnstone@usask.ca Reviewer Michelle Schuman michelle.schuman@a Modeler 2 Torre Jorgenson tjorgenson@abrinc.com Reviewer Lisa Saperstein Modeler 3 Reviewer Map Zone 73 Vegetation Type Wetlands/Riparian Dominant Species* PIMA LALA SALIX BENA k.usda.gov Lisa_Saperstein@fws. gov General Model Sources LEGR MYGA CAAQ SPHAG2 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system is found throughout boreal AK, in valley bottoms and broad, poorly drained floodplains (such as Yukon and Tanana Flats). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 115 of 204 Biophysical Site Description This sytem is typically found on valley bottoms and toe slopes up to eight percent slope, except on gravel substrates. Soils are poorly drained and acidic, often with a well-developed peat layer (NatureServe 2008). Much of this vegetation type is likely to be underlain by near surface permafrost (active layers less than one meter deep). The presence of permafrost and poor drainage of flat areas work in conjunction to keep soils cool and wet, restricting decomposition and aiding the accumulation of peat. Warming climates and deep-burning fires that occur late in the summer have the potential to disrupt the soil thermal regime and lead to permafrost thawing and subsidence (thermokarst). Thermokarst will lead to prolonged stages of wetland-marsh type vegetation. Re-establishment of black spruce forests is likely to occur when peatreaccumulates and the permafrost develops again, leading to a rising of the surface above the water table. Vegetation Description Boreal Lowland Wet Black Spruce forests are typically open and trees are generally stunted. Tree rooting zones are typically within the organic layer, with low nutrient availability and low rates of growth. Forests will tend to be open and stunted compared to adjacent, more well-drained areas. Common species include Picea mariana, Ledum groenlandicum, L. palustre ssp. decumbens, Betula nana, B. glandulosa, Empetrum nigrum, Vaccinium vitis-idaea, V. uliginosum, Chamaedaphne calyculata, Myrica gale, Carex spp., Eriophorum angustifolium, Menyanthes trifoliata and Sphagnum spp. (NatureServe 2008). Larix laricina may be locally abundant, particularly on less acidic sites. Disturbance Description Fire and thermokarst are the most important processes influencing Boreal Lowland Wet Black Spruce. Fire return intervals for this type generally range from 80-130yrs (this is a best guess). For the VDDT model, it was assumed that fire would occur on the less frequent end of the range provided above because the presence of moist surface fuel and the open forest canopy associated with this BpS would limit fire spread and therefore decrease the probability of any particular patch burning. Thermokarst processes play a key role in shaping the system, but the number of possible pathways and the time scale at which they operate are beyond the limit of what can be modeled with VDDT. As a result, this model has been simplified and limited in scope. This model includes Western North American Boreal Freshwater Emergent Marsh, Western North American Boreal Wet Meadow and Western North American Boreal Low Shrub Peatland as seral stages in the Boreal Lowland Wet Black Spruce system. These four classes are linked by thermokarst processes, as well as by fire dynamics. Boreal Freshwater Emergent Marsh may maintain itself on the landscape for decades, but on sites where permafrost development is likely and under appropriate hydrologic conditions, it may begin to dry out, develop an organic layer, and succeed to Boreal Wet Meadow. In this class, permafrost development begins, as mosses and organic material accumulate. This stage will succeed to Boreal Wet Low Shrub in the absence of disturbance, and permafrost and peat development will continue. In the absence of disturbance, trees will eventually begin to invade, and around age 75yrs, Boreal Lowland Wet Black Spruce or Black Spruce-Tamarack Fen will appear. In any of the three stages with permafrost, there is a low probability (modeled as 0.001) that permafrost melting will set the system back to the Freshwater Emergent Marsh class. Fire is also a possibility in the shrub and forest classes. A severe fire that impacts the peat layer may kill the shrub rootstock, setting the system back to the Wet Meadow class. If such a fire is followed by flooding (as from permafrost melting), the system may transition back to the Marsh class. If the fire is less intense, the shrubs may resprout and recover quickly. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 116 of 204 Although it is outside of the scope of this model, the system just described is part of a larger system connected by thermokarst dynamics. Under some conditions, the marsh class of this system may develop into a different type of Wet Meadow, with more bog characteristics, and permafrost may not develop again. This trajectory leads into the Ericaceous Dwarf Shrub Bog model, which could eventually lead to a forested bog or Wet Black Spruce-Tussock Woodland. In this forested stage, a permafrost layer would develop again. Alternatively, the Black Spruce Dwarf-tree Peatland or Black Spruce-Tamarack Fen could undergo paludification and transition to Boreal Sedge-Dwarf-Shrub Bog system. This can happen when restricted drainage from permafrost development (on inactive alluvial terraces, for example) leads to the establishment of Sphagnum spp. Or other peat forming mosses or sedges, and over time, peatland plants dominate the site. In either of these pathways, the forested bog or Tussock Black Spruce class has the potential to undergo permafrost melting and collapse and transition back to marsh. However, these cycles are likely to take thousands of years and are thus outside the scope of this modeling effort. Other pathways that would take the system out of this model include melting and impoundment, leading to an open water stage, or drainage of the marsh stage, leading into a white spruce-Calamagrostis canadensis system. Another pathway into this system is paludification over thousands of years, which could move a site from the Boreal and Sub-boreal Fen Complex model into this system. Adjacency or Identification Concerns This type can be found adjacent to Boreal Black Spruce Wet-Mesic Slope Woodland and Boreal Mesic Black Spruce Forest - Boreal. Boreal Lowland Wet Black Spruce is typically found on valley bottoms and toe slopes up to eight percent slope. As the slope increases past eight percent, the vegetation typically transitions to Mesic Black Spruce Forest or, on north-facing slopes, to Black Spruce Wet-Mesic Slope Woodland. Well-drained gravel sites may support Mesic Black Spruce Forest, even in low, flat areas. Native Uncharacteristic Conditions Scale Description Large patch (small patch) Issues/Problems Because the climate has been warming and drying over recent decades, thermokarst processes have been accelerating. At the same time, lakes and marshes have been shrinking and drying across the boreal region in recent decades. Many of the marshes in this region may be relicts of the earky Holocene postglacial period, rather than parts of a stable cycle of disturbance and succession. As a result, the concept of equilibrium dynamics inherent in this modeling process is difficult to apply to this system. Because thermokarst processes are accelerating, it was difficult to assign a reference probability to these processes. Over the past 5000yrs, the probability of thermokarst influencing a given point on the landscape averaged about 0.000014, compared with approximately 0.0006 in the past 50yrs. Under current climate conditions, the probability is presumably higher. Comments This model was created by Torre Jorgenson and refined by Colleen Ryan based on a more narrowly focused model created by Jill Johnstone for the Boreal Lowland Wet Black Spruce system. Jill Johnstone reviewed the final model for the wetland complex. Information provided by Amy Larsen was used in developing this model. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 117 of 204 Vegetation Classes Class A 5% Structure Data (for upper layer lifeform) Min Max Herbaceous Herbaceous Early Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Tree Size Class None Herbaceous Shrub Tree Indicator Species* and Canopy Position TYLA CARO6 EQFL Upper Upper Upper Upper Herbaceous Upper layer lifeform differs from dominant lifeform. Description 0-20yrs Freshwater Emergent Marsh. This class is characterized by herbaceous marsh vegetation such as Typha latifolia, Carex rostrata, C. aquatilis, C. utriculata, Scirpus validus, Menyanthes trifoliata, Equisetum fluviatile and Eleocharis palustris. Arctophila fulva becomes more common in the northern boreal region. Floating aquatic vegetation (Nuphar polysepalum, Potamogeton spp.) may also be present. Water level may fluctuate seasonally or between years, but is at or above the surface for most of the growing season. Areas of open water are often present seasonally. Permafrost is absent in the class. If water levels continue to be high, with frequent large fluctuations, class A may maintain itself for decades. If the hydrologic regime begins to stabilize and dry, this class may succeed to class B (Wet Meadow) in two years plus. Fire is extremely unlikely in this class. With regular seasonal flooding and high water levels, this class maintains itself. Alternate succession to class B (probability = 0.12) may occur if water levels stabilize and the site begins to dry. Class B Structure Data (for upper layer lifeform) Min Max 5% Mid Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position CACA4 CAAQ ERAN6 Upper Upper Upper Upper Tree Size Class Herbaceous Herbaceous None Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Distinguish from Class A for mapping by lack of aquatic plants and open water, presence of organic mat, mosses, permafrost, and/or dominance of sedges. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 118 of 204 Description 3-5yrs Wet Meadow. This class is characterized by dense sedges with a well-developed organic mat. Common and characteristic species include Calamagrostis canadensis, Carex aquatilis, C. bigelowii, Eriophorum angustifolium, E. russeolum, E. vaginatum, Equisetum arvense, Equisetum sylvaticum, Epilobium angustifolium, and the moss Ceratodon purpureus (Gracz 2005, Jorgenson et al. 1999). Water levels are more stable in this class than in the Freshwater Emergent Marsh. Soils are saturated at some point in the growing season but do not have standing water (NatureServe 2008). Permafrost development has begun in this class, making it subject to thermokarst processes. Fire is very unlikely in this class, due to the saturated soils. Succession to class C under typical (drying) conditions. Option 1 (probability = 0.02) represents a stable hydrologic regime that maintains class B. Option 2 (probability = 0.0006) represents increased water levels which would cause a transition to class A. Class C Structure Data (for upper layer lifeform) Min Max 45 % Mid Development 2 All Structures Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position BENA LEGR SALIX VACCI Upper Upper Lower Lower Dwarf Shrub (< 20 cm) Tree Size Class Tall Shrub (>1.5 m) None Upper layer lifeform differs from dominant lifeform. Description 6-74yrs Low Shrub Peatland Dwarf ericaceous, low or tall shrubs dominate this class. Myrica gale and Chamaedaphne calyculata are among the first shrubs to invade. Presence of sphagnum mosses tends to increase soil acidity which in turn favors the ericaceous shrubs such as Ledum groenlandicum, L. palustre ssp. Decumbens, Andromeda polifolia and Vaccinium uliginosum. Other common shrub species include Salix spp., Betula nana (B. glandulosa) and Empetrum nigrum. Permafrost is well developed in this class, making it subject to thermokarst processes. Fire is possible in this class during dry periods. Succession to class D under typical conditions. Severe replacement fire (MFRI = 175yrs) that burns into the peat layer will kill the shrubs, causing a transition to class B. Replacement fire followed by a hydrologic change that increased the water level could cause a transition to class A (probability = 0.0017). Option 2 (probability = 0.0006) represents increased water levels as a result of thermokarst processes which could kill the existing vegetation causing a transition to class A. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 119 of 204 Structure Data (for upper layer lifeform) Class D 45 % Late Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position PIMA LALA BENA MYGA Upper Upper Lower Lower Cover Height Min Woodland (10-24% tree cover) Max Closed (60-100% tree cover) Dwarf Tree (< 3 m) Tree Size Class Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. Description 75yrs+ Mature wet black spruce forest. Picea mariana or Picea mariana-Larix laricina mix. Trees are generally stunted and generally pole sized although some achieve the medium (9-21in) size class. The presence of Larix laricina is associated with higher soil pH. Common understory species include the same shrubs as in class C, as well as Calamagrostis canadensis, Carex aquatilis, C. bigelowii and sphagnum. Permafrost is well developed in this class, making it subject to thermokarst processes. The probability of fire is similar to that in the shrub class because the shrubs carry the fire. This class persists in the absence of disturbance. Overall replacement MFRI = 130yrs. Severe replacement fire (MFRI = 175yrs) will kill the shrubs, causing a transition to class B. Replacement fire followed by a hydrologic change that increased the water level could cause a transition to class A (probability = 0.0017). A mixed fire (MFRI = 333yrs) that killed most of the trees and top-killed the shrubs would cause a transition to class C. Option 2 (probability = 0.0006) represents increased water levels as a result of thermokarst processes which could kill the existing vegetation causing a transition to class A. Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 120 of 204 Fire Regime Group**: Fire Intervals IV Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 133.7 727.7 0.00748 0.00137 113 0.00886 Percent of All Fires 84 16 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Native Grazing Wind/Weather/Stress Competition Other (optional 1) continued high water level Other (optional 2) increased water level (thermokarst processes) References Foote, M. Joan. 1983. Classification, description, and dynamics of plant communities after fire in the Taiga of Interior Alaska. Res. Pap. PNW-307. Portland, OR: USDA Forest Service, Pacific Northwest Forest and Range Experiment Station. 108 pp. Gracz, M., Noyes, K, North, P., Tande, G. 2005 Wetland Mapping and Classification of the Kenai Lowland, Alaska. http://www.kenaiwetlands.net/. Jorgenson, M.T., J.E. Roth, M. Raynolds, M.D. Smith, W. Lentz, A. Zusi-Cobb and C.H. Racine. 1999. An ecological land survey for Fort Wainwright, Alaska. U.S. Army Cold Regions Research and Engineering Laboratory, Hanover, NH. U.S. Army Cold Regions Research Engineering Laboratory, Hanover, NH CRREL Report 99-9. 83 pp. Jorgenson, M.T., J.E. Roth, M.D. Smith, S. Schlentner, W. Lentz and E.R. Pullman. 2001. An ecological land survey for Fort Greely, Alaska. U.S. Army Cold Regions Research and Engineering Laboratory, Hanover, NH. ERDC/CRREL TR-01-04. 85 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Post, R.A. 1996. Functional profile of black spruce wetlands in Alaska. US EPA, Alaska Operations Office, Seattle, WA. EPA/910/R-96/006. 170 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 121 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316212 Western North American Boreal Black Spruce Dwarf-tree Peatland - Alaska Subboreal Complex This BPS is lumped with: see below This BPS is split into multiple models: This system was created by merging two Ecological Systems: Western North American Boreal Black Spruce Dwarf-tree Peatland and Western North American Boreal Sedge-Dwarf-Shrub Bog General Information Contributors (also see the Comments field Modeler 1 Torre Jorgenson Date 4/20/2008 tjorgenson@abrinc.com Reviewer Michelle Schuman michelle.schuman@a k.usda.gov Modeler 2 Colleen Ryan Modeler 3 colleen_ryan@tnc.org Reviewer Map Zone 73 Vegetation Type Wetlands/Riparian Dominant Species* VAOX ANPO VAUL LEGR Reviewer General Model Sources BENA EMNI CAREX SPHAG2 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system occurs in lowland areas throughout the sub-boreal region of AK. Biophysical Site Description This system includes hydrologically closed bogs, poor fens and organic-rich black spruce forests. Soils are poorly drained, acidic and nutrient-poor, often with a well-developed peat layer. Permafrost is generally absent. Vegetation Description Common species include Oxycoccus microcarpus, Andromeda polifolia, Vaccinium uliginosum, Ledum groenlandicum, Betula nana, Empetrum nigrum, Carex microglochin, C. rotundata, C. rariflora, C. lasiocarpa,C. limosa, C. livida, C. williamsii, Eriophorum brachyantherum, E. angustifolium and Drosera spp. Sphagnum spp. are usually abundant in the ground layer (NatureServe 2008). Disturbance Description Succession begins for this system in shallow ponds or low-lying wetlands that are hydrologically isolated, floodplain dynamics (oxbows) or thermokarst. An organic root mat typically develops and is either anchored to the mineral soil or floating on water such as a pond’s edge. Over time, peat-forming mosses and sedges may fill in the basin. As the peat layer develops, low and dwarf shrubs become established. Dwarf trees may establish on the well-developed peat and also around the margin of the peatland. The *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 122 of 204 primary disturbances in this system are changes in hydrology, which can move the system between successional classes or out of this system. A common pathway into this system is via permafrost degradation, from the Western North American Boreal Lowland Wet Black Spruce Complex or another boreal permafrost system. Permafrost degradation leading to collapse scars and thaw ponds is a common in boreal Alaska, and studies from the Tanana Flats show areas of widespread degradation (Racine et al. 1998, Jorgenson et al. 2001a, Jorgenson et al. 2001b, Jorgenson et al. 2003). Thaw ponds form when ice rich permafrost degrades and collapses forming a basin. Aquatic plants rapidly colonize the pond. Over time, marsh plants and sphagnum moss invade, creating peatland conditions. If a collapse scar is isolated, succession follows a bog development model (this system), whereas in an open hydrologic setting, succession follows a fen development model (causing a transition out of this system to Alaska Sub-Boreal Fen Complex). Pond systems may become connected as adjacent permafrost thaws. Succession to peatlands can also occur through paludification of previously forested landscapes. Restricted drainage from permafrost development (on inactive alluvial terraces, for example) can lead to the establishment of Sphagnum spp. or other peat forming mosses or sedges, and overtime, peatland plants dominate the site. Adjacency or Identification Concerns Compare with Alaska Sub-Boreal Fen Complex. Both of these non-permafrost systems may occur in the boreal region. The fen system will develop on sites with more open hydrology, while this system will develop on sites with isolated or closed hydrology. This system occurs on less than one percent of the boreal landscape. Native Uncharacteristic Conditions Scale Description Large patch (small patch) Issues/Problems The rate of thermokarst processes appears to be accelerating rapidly, leading to widespread conversion of permafrost systems to this system and other non-permafrost wetland systems. Widespread ecosystem conversion has been documented in the Tanana Flats (Jorgenson et al. 2001b), among other areas. The altered rates of thermokarst processes make it difficult to create an equilibrium model for this system, since more land area has entered this system than exited it in recent decades. Comments Vegetation Classes *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 123 of 204 Class A Structure Data (for upper layer lifeform) Min Max 10 % Early Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Tree Size Class None Herbaceous Shrub Tree Indicator Species* and Canopy Position CAAQ ERRU2 ERAN6 SPHAG2 Upper Upper Upper Lower Herbaceous Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description 0-49yrs Subarctic Lowland Bog Meadow/Marsh/Poor Fen. This early stage may exhibit a variety of herbaceous vegetation, from marsh to sedge meadow to sphagnumdominated poor fen. Common species include sedges (Carex aquatilis, C. rariflora, C. limosa, C. utriculata, Eriophorum russeolum and E. angustifolium) and Sphagnum spp., sometimes with Calamagrostis canadensis or Equisetum fluviatile. This stage could be preceded by an open water or aquatic bed system (not included here because they are sparsely vegetated). This class maintains itself in the absence of disturbance or follows an alternate succession pathway (probability = 0.01) to class B. Mid Development 1 All Structures Structure Data (for upper layer lifeform) Min Max Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Class B 20 % Herbaceous Shrub Tree Indicator Species* and Canopy Position VAOX ANPO VAUL LEGR Middle Upper Upper Upper Dwarf Shrub (< 20 cm) Tree Size Class Low Shrub (20 cm to 1.5 m) None Upper layer lifeform differs from dominant lifeform. Description 1-299yrs Sedge Dwarf Shrub Bog. Common species include Vaccinium oxycoccus, V. uliginosum, Andromeda polifolia, Ledum groenlandicum, Betula nana, Empetrum nigrum, Carex microglochin, C. rotundata, C. rariflora, C. lasiocarpa, C. limosa,C. livida, C. williamsii, Eriophorum brachyantherum, E. angustifolium and Drosera spp. Sphagnum spp. are usually abundant in the ground layer. Sucession to class C. Hydrological changes that cause a transition back to class A are modeled as Option 1 (probability = 0.001). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 124 of 204 Class C 70 % Late Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position PIMA Cover Height Structure Data (for upper layer lifeform) Min Max Woodland (10-24% tree cover) Open (25-59% tree cover) Dwarf Tree (< 3 m) Tree Size Class Tree (> 3 m) None Upper Upper layer lifeform differs from dominant lifeform. Description 300yrs+ Wet Black Spruce Forest/Forested Bog. Common species include Picea mariana, Ledum groenlandicum, Betula nana, Empetrum nigrum, Vaccinium vitis-idaea, V. uliginosum, Chamaedaphne calyculata, Carex spp., Eriophorum angustifolium and Sphagnum spp. This class maintains itself in the absence of disturbance. Hydrological changes that cause a transition back to class A are modeled as Option 1 (probability = 0.001). Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 125 of 204 Fire Regime Group**: Fire Intervals NA Avg FI Min FI Max FI Probability Percent of All Fires Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) increased water level Other (optional 2) References Boggs, K., A. Garibaldi, J. Stevens, J. Grunblatt and T. Helt. 2001. Denali National Park and Preserve Landcover mapping project. Volume 2: Landcover classes and plant associations. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage, 707 A Street, Anchorage, AK. 164 pp. Jorgenson, M.T. et al. 2003. An ecological land survey for Fort Richardson, Alaska. Cold Regions Research and Engineering Laboratory, Hanover, NH, ERDC/CRREL TR-03019. Jorgenson, M.T., J.E. Roth, M.D. Smith, S. Schlentner, W. Lentz and E.R. Pullman. 2001a. An ecological land survey for Fort Greely, Alaska. U.S. Army Cold Regions Research and Engineering Laboratory, Hanover, NH. ERDC/CRREL TR-01-04. 85 pp. Jorgenson, M.T., C.H. Racine, J.C. Walters and T.E. Osterkamp. 2001b. Permafrost degradation and ecological changes associated with a warming climate in central Alaska. Climatic Change 48: 551-579. Jorgenson, M.T., J.E. Roth, M.K. Raynolds, M.D. Smith, W. Lentz, A.L. Zusi-Cobb and C. H. Racine. 2001c. An ecological land survey for Fort Wainwright, Alaska. U.S. Army Cold Regions Research and Engineering Laboratory, Hanover, NH. CRREL TR-99-9. 92 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Racine, C.H., Jorgenson, M.T. and Walters, J.C. 1998. Thermokarst vegetation in lowland birch forests on the Tanana Flats, Interior Alaska, USA. In: Proceedings of the 7th International Conference on Permafrost, June 23-27, 1998. Universite Laval, Sainte-Foy, Quebec, Collection Nordicana, 57: 927-934. Zoltai, S.C., Taylor, S., Jeglum, J.K., Mills, G.F. and Johnson, J.D. 1988. Wetlands of Boreal Canada. Pages 99-154 in: Wetlands of Canada. Ecological Land Classification Series No. 24. Environment Canada, Ottawa, and Polyscience Publications Inc., Montreal. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 126 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316220 Western North American Boreal Black Spruce Wet-Mesic Slope Woodland This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Joan Foote Modeler 2 Colleen Ryan Modeler 3 Date joanf@mosquitonet.com Reviewer Michelle Schuman michelle.schuman@a k.usda.gov Reviewer Will Putnam wputman@tananachie colleen_ryan@tnc.org fs.org Reviewer Lisa Saperstein Lisa_Saperstein@fws. gov Map Zone 73 Vegetation Type Wetlands/Riparian Dominant Species* PIMA LEPAD VAVI EMNI 4/18/2008 General Model Sources VAUL BENA SPGI70 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system is found throughout interior boreal AK. Biophysical Site Description This BpS is found on north-facing slopes underlain by permafrost. Soils are poorly drained and acidic with a well-developed peat layer (NatureServe 2008). Vegetation Description The dominant overstory vegetation is Picea mariana. Mature trees on these sites are usually smaller than those on mesic sites. Common shrubs include Ledum groenlandicum, Ledum palustre, Betula nana (this species here includes B. glandulosa), Empetrum nigrum, Vaccinium vitis-idaea and V. uliginosum (NatureServe 2008). Herbs include Equisetum sylvaticum, Rubus chamaemorus and Carex spp. (Foote 1983). Older stands will include lichens, especially Cladina arbuscula and C. rangiferina. Mosses include Sphagnum spp., Pleurozium schreberi and Polytrichum spp. Feathermosses are typical of cooling soils, which can lead to permafrost development. Sites where permafrost is closer to the surface will have more sphagnum. Disturbance Description Fire is the dominant disturbance mechanism for this forest type. Fire is facilitated by an abundance of fine fuel and ladder fuel in this type. The overall MFRI for this model is similar to that for Western North American Boreal Mesic Black Spruce Forest - Boreal. Despite saturated soils, wet black spruce forest on *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 127 of 204 slopes lacks the standing water that limits fire in wet black spruce flats (Boreal Lowland Wet Black Spruce Complex). Fires in this forest type are usually fast-moving and patchy. High severity fire may lead to an increased deciduous tree component (Johnstone and Kasischke 2005). Adjacency or Identification Concerns This type is intermingled with the Western North American Boreal Mesic Black Spruce Forest - Boreal BpS and the Boreal Lowland Wet Black Spruce Complex BpS. Boreal Lowland Wet Black Spruce Complex is found on low flats and concave and toeslope sites. Boreal Black Spruce Wet-Mesic Slope Woodland is found on north-facing slopes, while Mesic Black Spruce Forest - Boreal is found on upper, convex slopes of other aspects. In hilly areas, Lowland Wet Black Spruce Complex typically occurs on the flats and toe slopes up to an eight percent grade (T. Jorgenson, pers. Comm.) Above the eight percent cutoff will be Boreal Black Spruce Wet-Mesic Slope Woodland on north-facing slopes or Boreal Mesic Black Spruce Forest - Boreal on other slopes. This model applies to sites with well-developed peat soils on permafrost. In contrast, mesic black spruce typically occurs on sites lacking permafrost and peat soils in the southern parts of the Boreal region. North of Fairbanks, both Boreal Mesic Black Spruce Forest - Boreal and Boreal Lowland Wet Black Spruce Complex may occur with permafrost. Native Uncharacteristic Conditions Scale Description Large patch (small patch) Issues/Problems Comments Joan Foote and Colleen Ryan drafted this model. Michelle Schuman, Lisa Saperstein and Will Putnam reviewed an early draft of this model. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 10 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Indicator Species* and Canopy Position EQSY BENA RUCH SPGI70 Upper Middle Low-Mi Lower Height Dwarf Shrub (< 20 cm) Tree Size Class Tall Shrub (>1.5 m) None Upper layer lifeform differs from dominant lifeform. Herbs or shrubs can dominate. Description 0-19yrs Herbaceous and/or shrub, open or closed. After a fire, herbs and shrubs quickly re-establish on the site. Soon, the shrubs begin to overtop the herbs. By the end of this class, a closed shrub canopy is present. Spruce seedlings are present in the understory. Common shrubs include Ledum groenlandicum, Ledum palustre, Betula nana (syn. B. glandulosa), Empetrum *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 128 of 204 nigrum, Vaccinium vitis-idaea and V. uliginosum. Herbs include Equisetum sylvaticum, Rubus chamaemorus and Carex spp. Mosses include Sphagnum spp., Pleurozium schreberi and Polytrichum spp. On some sites, alder and willow may be present. Fire is unlikely in this class due to lack of fuel and saturated soils. Any fire will be stand-replacing. Succession to class B. Replacement MFRI = 1000yrs. Class B 20 % Mid Development 1 All Structures Upper Layer Lifeform Herbaceous PIMA RUCH EMNI SPGI70 Shrub Tree Cover Indicator Species* and Canopy Position Structure Data (for upper layer lifeform) Min Max Woodland (10-24% tree cover) Closed (60-100% tree cover) Dwarf Tree (< 3 m) Height Tree Size Class Seedling/Sapling <5" Mid-Upper Low-Mid Low-Mid Lower Tree (> 3 m) Upper layer lifeform differs from dominant lifeform. For mapping, if this class can not be be distinguished from class C and D by existing vegetation type (Black Spruce Scrub and Shrub Swamp), class B can be considered a dwarf tree class. Description 20-40yrs Black spruce, open or closed, seedling/sapling. This class is dominated by black spruce saplings and shrubs. Early in this stage, the tree seedlings begin to overtop the low shrubs. By the end of this class, the trees will overtop the tall shrubs (willow and/or alder), if present. The end of this class will be a closed stand of spruce saplings. In some cases, this class may include older black spruce trees that have survived a mixed fire. Succession to class C is the primary pathway, which is followed by closed stands. If this class is open, it will succeed to class D, represented in the model by alternate succession (probability = 0.03). Replacement MFRI = 170yrs. Mixed fire (MFRI = 170yrs) maintains the system in class B. Class C 35 % Late Development 1 Closed Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position PIMA RUCH EMNI SPGI70 Upper Low-Mid Low-Mid Lower Structure Data (for upper layer lifeform) Min Max Closed (60-100% tree cover) Closed (60-100% tree cover) Tree Size Class Tree (> 3 m) Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Description 40yrs+ Black spruce, closed. This class is dominated by relatively dense sapling to pole-sized Picea mariana, typically in even-aged stands. Trees will typically range from 2.5-10cm DBH and 2-7m tall (Foote 1983). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 129 of 204 However, uneven aged stands may occur where some individuals have survived an earlier mixed fire. The canopy is nearly 100% spruce. This is typically the most flammable class because of the pulses of fuel created by the die-off of overtopped shrubs around age 40-60yrs. Abundant snags, dead lower branches and fine fuel also increase the likelihood of fire in this class. Mixed fire in this class will transition the system to class B, creating patches of mature trees in a matrix of shrubs, with spruce regeneration in the understory. This class will persist in the absence of disturbance. Replacement MFRI = 125yrs. Mixed fire (MFRI = 110yrs) causes a transition to class B. Structure Data (for upper layer lifeform) Class D 35 % Late Development 2 Open Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position PIMA PLSC70 CLADI3 SPGI70 Upper Lower Lower Lower Min Max Woodland (10-24% tree cover) Open (25-59% tree cover) Height Tree (> 3 m) Tree Size Class Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Description 40yrs+ Black spruce/lichen, open. Open canopy mature Picea mariana, with feathermoss and lichen. Mosses and lichens expand with age, as the leaf litter from the shrubs disappears. In this class, the canopy has begun to open up and lichen development has begun. Lichen species include Cladina arbuscula, C. rangiferina and Nephroma articum. Mosses include Pleurozium schreberi, Polytrichum spp., Hylocomium splendens and Dicranum spp, as well as Sphagnum spp. Low shrubs, including Vaccinium vitis-idaea, V. uliginosum and Ledum groenlandicum, are often present in the understory. Any fire in this class will kill the lichens. Mixed fire will transition the system to class B, creating patches of mature trees in a matrix of shrubs, with spruce regeneration in the understory. This class will persist in the absence of disturbance. Replacement MFRI = 170yrs. Mixed fire (MFRI = 110yrs) causes a transition to class B. Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 130 of 204 Fire Regime Group**: Fire Intervals III Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 163.9 140.8 0.0061 0.0071 76 0.01321 Percent of All Fires 46 54 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Foote, M. Joan. 1983. Classification, description, and dynamics of plant communities after fire in the Taiga of Interior Alaska. Res. Pap. PNW-307. Portland, OR: USDA Forest Service, Pacific Northwest Forest and Range Experiment Station. 108 pp. Johnstone, J.F. and E.S. Kasischke. 2005. Stand-level effects of soil burn severity on post-fire regeneration in a recently-burned black spruce forest. Canadian Journal of Forest Research 35: 2151-2163. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 131 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316240 Western North American Boreal Deciduous Shrub Swamp This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Date 4/24/2008 Modeler 1 Tina Boucher antvb@uaa.alaska.edu Reviewer Michelle Schuman michelle.schuman@a Modeler 2 Colleen Ryan Modeler 3 colleen_ryan@tnc.org Reviewer k.usda.gov Reviewer Map Zone 73 Vegetation Type Wetlands/Riparian Dominant Species* General Model Sources ALINT CACA4 ALVIS EQUIS SAPU15 SARI4 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This BpS occurs throughout the boreal and sub-boreal regions of AK. Biophysical Site Description Shrub swamps occur on poorly drained fine textured soil in lowland areas or depressions that retain standing water throughout all or most of the growing season (NatureServe 2008; Viereck et al. 1992). Soils range from muck to mineral (does not include peatlands; NatureServe 2008). This system tends to occur in transition zones between peatlands and forest systems. Specific locations may shift over time. Vegetation Description Deciduous shrub swamps are usually dominated by alders, but willows or an alder/willow mix also occur (Viereck et al. 1992). Common species include Alnus incana ssp. tenuifolia, Alnus viridis ssp. sinuata, Salix pulchra, Salix richardsonii, Calamagrostis canadensis, Equisetum spp., Potentilla palustris (Comarum palustre) and hydrophytic mosses (Viereck et al. 1992). Though this system is classified as closed by Viereck, et al. (1992), it may also occur in an open form. Disturbance Description Shrub swamps likely represent a topoedaphic climax community which will persist as long as the hydrologic conditions supporting them are maintained (Viereck et al. 1992). Some sites may move back and forth between this system and peatlands, forest, or even open water as hydrology changes, but these transitions are not directional, and are generally outside of the time scale of this model. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 132 of 204 This system typically acts as a fire break, but it could burn under drought conditions and severe fire behavior. When fire occurs, it is likely to be replacement fire. Alder may resprout following fire, allowing recovery to this system. Alternatively, fire may cause changes in hydrology that could transition the site out of this system, possibly toward an open water condition. Replacement fire that regenerated this system is estimated at a MFRI of 500yrs. Adjacency or Identification Concerns Native Uncharacteristic Conditions Scale Description Typically small patch, but some large patches occur, especially in the Tanana Flats area. Issues/Problems Comments Addition of an early herb stage was considered, but it is believed that shrubs typically recover quickly from fire. Flooding was not included as a disturbance because the dominant species are flood-tolerant. Flooding severe or prolonged enough to kill the shrubs is likely to cause a transition out of this system. Suggested reviewers for this system include: Al Batten, Torre Jorgenson and Katie Ireland. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 100 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Indicator Species* and Canopy Position ALINT ALVIS SAPU15 SARI4 Upper Upper Upper Upper Height Dwarf Shrub (< 20 cm) Tree Size Class Tall Shrub (>1.5 m) None Upper layer lifeform differs from dominant lifeform. Description Zero years plus Alpine Herbaceous-Dwarf Shrub communities may include a diverse mix of species and species assemblages. The class Indicator Species listed are not true indicators -- refer to the Vegetation Description for species information. No disturbances modeled. Open to closed tall shrub swamp. The shrub canopy is commonly 3-5m tall (Viereck et al. 1992). Overstory is typically dominated by alder, willow or an alder/willow mix. Common overstory species include Alnus incana ssp. Tenuifolia, Alnus viridis ssp. Sinuata, Salix pulchra and S. lanata. Understory can include Calamagrostis canadensis, Equisetum spp., Potentilla palustris (Comarum palustre) and hydrophytic mosses. This class persists indefinitely under appropriate hydrological conditions. Replacement fire (MFRI = 500yrs) *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 133 of 204 will reset the class to age zero as shrubs resprout. Class B Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Cover Indicator Species* and Canopy Position Height Tree Size Class Shrub Tree Upper layer lifeform differs from dominant lifeform. Description Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 134 of 204 Fire Regime Group**: Fire Intervals V Replacement Historical Fire Size (acres) Avg FI Min FI Max FI Probability 496.1 0.00202 496 0.00204 Percent of All Fires 99 Mixed Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Batten, Alan R., S. Murphy and D.F. Murray. 1978. Definition of Alaskan coastal wetlands by floristic criteria. Corvallis, OR: Corvallis Environmental Research Laboratory; EPA 80496501. 490 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck, L.A., Dyrness, C.T., Batten, A.R. and Wenzlick, K.J. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 135 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316280 Western North American Boreal Low ShrubTussock Tundra This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Jennifer Allen 4/4/2008 Jennifer_Allen@nps.go v Reviewer Lisa Saperstein Reviewer Stuart Chapin Reviewer Modeler 2 Modeler 3 Map Zone 73 Vegetation Type Upland Shrubland Dominant Species* ERVA4 CABI5 BENA SAPU15 Date General Model Sources LEPAD VACCI CACA4 ARCTA Literature Local Data Expert Estimate Lisa_Saperstein@fws. gov fffsc@uaf.edu Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This BpS occurs in the boreal and the northern section of the sub-boreal region of AK. It is found on lowland to subalpine sites. Biophysical Site Description The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): Permafrost is usually present at depths of 30-50cm. Soils are generally acidic, poorly drained, gleyed, and often with a poorly decomposed organic horizon at the surface, which may constitute most of the active layer. Frost scars are common. Tussock communities occur on gentle slopes, terraces, and old alluvial deposits. Sites are often underlain by silty mineral soils with a surface peat layer 10-40 cm thick surrounding the tussocks (Viereck et al 1992). Vegetation Description The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): Boreal Low Shrub-Tussock Tundra is a common lowland system dominated by tussock sedges and low shrubs. Eriophorum vaginatum is the primary tussock-former in most stands, but Carex bigelowii may be the dominant tussock sedge on some sites. Other indicator species include Betula nana (includes B. glandulosa), Salix pulchra, Ledum decumbens (Ledum palustre L. ssp. decumbens), Vaccinium vitisidaea, V. uliginosum, Empetrum nigrum and Carex spp. Grasses, including Calamagrostis canadensis and Arctagrostis spp., may also be present. Lichens are scarce (with the possible exception of Peltigera canina). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 136 of 204 Disturbance Description The fuel layer in sedge-shrub tussock tundra is dense and continuous and leads to large, fast spreading fires (Duchesne and Hawkes 2000, Racine et al 1987). Racine (1979) found much variation in burn intensity on a landscape scale on the Seward Peninsula, from completely unburned to intensely burned. These patterns are related to variations in topography and the composition, moisture content and soil organic accumulations of the plant communities. Fires in Eriophorum tussock tundra types tend to be light because of the wet soil profile (Wein 1971). Burns in this type usually consume all aerial woody and herbaceous plant material and litter; regeneration is vigorous via rhizomes and root sprouts. Racine (1979) found that burning was generally less severe in the tussock-shrub and sedge-shrub tundras than in the birch and ericaceous shrub tundra of the Seward Peninsula. He found that tundra burns were patchy, with unburned communities and unburned patches within burned communities. More fires occur near the forest-tundra ecotone and spread further if trees are present (Heinselman 1981). Wein (1976) reports that July and August are the most common months for lightning fires in tundra ecosystems, while Racine et al (1983, 1985) found that distinct fire seasons occur in both June and July in the Noatak River watershed. Subsidence and thermal erosion following fire is usually minimal in tundra ecosystems (Walker 1996). In most areas of tussock-shrub tundra on the Seward Peninsula, less than one half of accumulated organic soil layer was removed by fire (Racine 1979). Thaw depths increased to reach into the mineral soils, but were not greatly increased except where organics were removed. Frost features were made more conspicuous, and soil nutrient concentrations (K and P) increased locally. Mean fire return interval estimates for tussock tundra ecosystems include: -50-300yrs (personal communication FRCC experts’ workshop March 2004) -260yr (s.d. 170) fire return interval for past 1500yrs for Noatak National Preserve (preliminary data from Higuera et al. 2008) -180-1460yrs in forest shrubzone and 9320yrs in shrub subzone in northern Quebec; shorter cycle west of Hudson’s Bay/in interior zone (Payette et al 1989) -612yrs for Noatak River watershed (all vegetation types) (Racine et al 1983) -611yr fire rotation for Noatak River watershed (all vegetation below 600m which is predominantly tundra) (Racine et al 1985) -Fire interval yet to be determined (Racine et al 1987) -Rapid recovery following fire makes fire frequency difficult to determine (Wein 1971) -The fire regime of tundra systems are likely quite variable from one region to another making generalizations difficult (Viereck and Schandelmeier 1980) On interior and southcentral Alaska Tussock Tundra sites the thaw pond cycle (disturbance leads to thawing of permafrost and ponding) and paludification (Sphagnum layer buildup and saturation) are important disturbances. On the Seward Peninsula and western AK, frost action creates polygonal ground and other periglacial features and is a widespread, small-scale and continuous disturbance. Change in the arctic and subarctic climate is another source of disturbance that is currently affecting tundra ecosystems. Adjacency or Identification Concerns The Boreal Low Shrub-Tussock Tundra is similar to the Arctic Tussock Low Shrub Tundra system (and the Tussock Tundra 2 PNV) that occurs in AK’s arctic and has a longer mean fire return interval *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 137 of 204 (NatureServe 2008). Geographic location is the best determinant between these two systems. Native Uncharacteristic Conditions Scale Description Vegetation found in large patches Wien (1976) reports many tundra fires in the 1-100ha size range and few large (thousands of ha) fires. Racine (1979) reports that in 1977, lightning-caused fires burned 35,480ha on the Seward Peninsula, with one fire burning 9440ha. Jandt and Meyers (2000) report that large fires (>200,000ha) occur about every 10yrs in the Buckland Valley and surrounding highlands of the Seward Peninsula. Racine et al (1983) found that 40 fires burned 100,000ha (1000km2) in the 30,000km2 watershed of the Noatak River between 1956 and 1981. Racine et al (1985) found a minimum fire size of 0.4ha, a maximum fire size of 45,800ha and a mean fire size of 1310ha from 1956-1983 in the Noatak River watershed, an area dominated by tundra vegetation. Of the 79 fires in analyzed by Racine et al (1985), nearly half were between 1-10ha in size. Forty-three percent of wildland fires occurring in interior Alaska occur in treeless areas, primarily tundra bogs and fens (Viereck 1975). Issues/Problems Comments This model was based on the FRCC Guidebook PNVG model for Tussock Tundra 1 (TT1; Murphy and Witten 2006) and input from the experts who attended the LANDFIRE Fairbanks (Nov. 07) modeling meeting and was refined by Jennifer Allen. Much of the text in the Disturbance Description and Scale Description portion of this report were taken from the TT1 description (Murphy and Witten 2006). Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 10 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position ERVA4 CABI5 CACA4 ARCTA Upper Upper Upper Upper Herbaceous Height Herbaceous Tree Size Class None Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description 0-14yrs Mesic herbaceous-graminoid-tussock-sedge. First year following fire Eriophorum (cottongrass) and Carex spp. (sedges) regrow via rhizomes, most vascular species begin to recover, shrubs sprout from rootstock. Sedges often capture site 6-10yrs post fire. Grasses (Calamagrostis and Arctagrostis) are locally important following fire. Succession to class B. Replacement MFRI = 175yrs. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 138 of 204 Class B Structure Data (for upper layer lifeform) Min Max 75 % Mid Development 1 Closed Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position BENA SALIX ERVA4 CABI5 Shrub Tree Dwarf Shrub (< 20 cm) Tree Size Class Upper Upper Upper Upper Low Shrub (20 cm to 1.5 m) None Upper layer lifeform differs from dominant lifeform. Classes B and C should be mapped based on EVT if possible. Class B is represented by Mixed Shrub-Sedge Tussock Tundra/Bog and class C by tussock-shrub-lichen. If these types are not mapped, B can be considered a low shrub class and C a dwarf shrub class for mapping. Description 15yrs+ Low shrub-tussock. Tussocks dominated by Eriophorum (cottongrass), Carex spp. (sedges). Common shrub species include Betula nana, Salix spp. and Vaccinium uliginosum. Lichens begin to re-establish but do not reach former abundance until 50-120yrs following fire. Fire is difficult to detect even in the early stages of this class; however, the proportions of species differ from the pre-burn community, with very few lichens, fewer shrubs and more sedges, grasses and cottongrass. Former abundances of all species are typically reached 50-120yrs post fire. Lichens, if present, have <25% cover. This class can persist in the absence of disturbance or follow an alternate succession pathway to class C (probability = 0.001). Replacement MFRI = 175yrs causes transition to class A. Mid Development 1 Open Structure Data (for upper layer lifeform) Min Max Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Class C 15 % Herbaceous Shrub Tree Indicator Species* and Canopy Position BENA SALIX ERVA4 PECA60 Upper Upper Upper Upper Dwarf Shrub (< 20 cm) Tree Size Class Low Shrub (20 cm to 1.5 m) None Upper layer lifeform differs from dominant lifeform. Classes B and C should be mapped based on EVT if possible. Class B is represented by Mixed Shrub-Sedge Tussock Tundra/Bog and class C by tussock-shrub-lichen. If these types are not mapped, B can be considered a low shrub class and C a dwarf shrub class for mapping. Description 80yrs+ Dwarf shrub-lichen-tussock. Tussocks are dominated by shrubs and lichens. Species composition is similar *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 139 of 204 to that in class B, but lichen cover is >25%. This class can persist in the absence of disturbance. Replacement MFRI = 200yrs causes transition to class A. Mixed fire (MFRI = 2000yrs) maintains this class. Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Min Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals IV Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 180 9999 0.00556 0.00010 177 0.00567 Percent of All Fires 98 2 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Duchesne L.C. and B.C. Hawkes. 2000. Fire in northern ecosystems. In: Brown, J.K. and J.K. Smith (eds.) Wildland fire in ecosystems: effects of fire on flora. Gen. Tech. Rep. RMRS-GTR-42-vol 2. Ogden, UT: USDA Forest Service, Rocky Mountain Research Station. 257 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 140 of 204 Heinselman, M.L. 1981. Fire and succession in the conifer forests of northern North America. In: West, D.C., H.H. Shugart, and D.B. Botkin. Forest succession: concepts and application. Springer-Verlag, New York. Chapter 23. Higuera, P.E., M. Chipman, J. Allen, S. Rupp and F.S. Hu. 2008. Preliminary data provided by Jenifer Allen from a study of tundra fire regimes in the Noatak National Preserve since 6000 years before present. Jandt, R.J. and R.C. Meyers. 2000. Recovery of lichen in tussock tundra following fire in Northwestern Alaska. USDOI BLM-Alaska Open File Report 82. Murphy, K.A. and E. Witten. 2006. Tussock Tundra 1. In Fire Regime Condition Class (FRCC) Interagency Guidebook Reference Conditions. Available at www.frcc.gov. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Payette, S., C. Morneau, L. Sirois and M. Desponts. 1989. Recent fire history of the northern Quebec biomes. Ecology. 70: 656-673. Personal communication experts’ workshop, March 2-4 2004. Fire Regime Condition Class (FRCC) interagency experts’ workshop to develop and review Potential Natural Vegetation (PNV) groups for Alaska. Anchorage, AK. Racine, C.H., L.A. Johnson and L.A. Viereck. 1987. Patterns of vegetation recovery after tundra fires in northwestern Alaska, USA. Arctic and Alpine Research. 19: 461-469. Racine. 1979. Climate of the Chucki-Imuruk area. Pages 32-37 in H. R. Melchior, ed., Biological Survey of the Bering Land Bridge National Monument. Alaska Cooperative Park Studies Unit, University of Alaska Fairbanks, Fairbanks, AK. Racine, C. H., W. A. Patterson III, and J. G. Dennis. 1983. Permafrost thaw associated with tundra fires in northwest Alaska. Pages 1024-1029 in Proceedings, Permafrost, Fourth International Conference. National Academy Press, Washington, D.C. Racine, C.H., J.G. Dennis and W.A. Patterson III. 1985. Tundra fire regimes in the Noatak River Watershed, Alaska: 1956-83. Arctic 38:194-200. Racine, C., J.L. Allen and J.G. Dennis. 2006. Long-term monitoring of vegetation change following tundra fires in Noatak National Preserve, Alaska. Arctic Network of Parks inventory and monitoring program, National Park Service, Alaksa Region. Report no. NPS/AKRARCN/NRTR-2006/02. Viereck, L.A. 1975. Forest ecology of the Alaska Taiga. In: Proceedings of the circumpolar conference on northern ecology; 1975 September; Ottawa, ON. National Research Council of Canada: I-1 to I-22. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. Viereck, L.A. and Schandelmeier, L.A. 1980. Effects of fire in Alaska and adjacent Canada--a literature *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 141 of 204 review. BLM-Aalska Tech. Rep. No. 6. Anchorage, Alaska: U.S. Department of the Interior, Bureau of Land Management. 124 pp. Walker, D.A. 1996. Disturbance and Recovery of Arctic Alaskan Vegetation. In: J.F. Reynolds and J.D. Tenhunen eds, Landscape function and disturbance in arctic tundra. Ecological studies vol. 120, Springer Verlag Berlin Heidleberg. Wein, R.W. 1971. Panel discussion: In: Slaughter, C.W., Barney, Richard J. and Hansen, G.M. (editors). 1971. Fire in the Northern Environment – A Symposium. Sponsors: Alaska Forest Fire Counc il and Society of American Foresters. Held at: The University of Alaska, Fairbanks. April 13-14, 1971. Published by: Pacific. Wein, R.W. 1976. Frequency and characteristics of arctic tundra fires. Arctic 29: 213-222. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 142 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316290 Western North American Boreal Tussock Tundra This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Jennifer Allen 4/4/2008 Jennifer_Allen@nps.go v Reviewer Lisa Saperstein Reviewer Stuart Chapin Reviewer Modeler 2 Modeler 3 Vegetation Type Upland Grassland/Herbaceous Dominant Species* ERVA4 CABI5 VAOX CHCA2 Date General Model Sources BENA LEPAD SPHAG2 PLSC70 Literature Local Data Expert Estimate Map Zone 73 Lisa_Saperstein@fws. gov fffsc@uaf.edu Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This BpS occurs in the boreal and the northern section of sub-boreal region of AK. It is found on lowland to subalpine sites. Biophysical Site Description The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): Permafrost is usually present at depths of 30-50cm. Soils are generally poorly drained, gleyed, and often with a poorly decomposed organic horizon at the surface, which may constitute most of the active layer. Frost scars are common. Tussock bog communities occur on lowlands of boreal and boreal transition Alaska, on filled-in sloughs on flood plains and on cold, poorly drained slopes and terraces. These sites are underlain by wet, silty mineral soils with a surface peat layer 10-40cm thick surrounding the tussocks (Viereck et al 1992). Vegetation Description The following information was taken from the draft Boreal Ecological Systems description (NatureServe 2008): Boreal Tussock Tundra is dominated by sedges in a tussock growth form. Eriophorum vaginatum is the primary tussock-former in most stands and Carex bigelowii is also common. On wetter sites, Oxycoccus spp. (Vaccinium oxycoccos) and Chamaedaphne calyculata may be present. Total shrub cover is <25%, although shrubs such as Betula nana (including B. glandulosa), Ledum decumbens (Ledum palustre L. ssp. decumbens) and Vaccinium spp. may be present. Mosses (Sphagnum spp., Pleurozium schreberi, Hylocomium splendens) may form a nearly continuous mat between tussocks. Lichens may be a *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 143 of 204 significant component on some sites. Disturbance Description The fuel layer in sedge-shrub tussock tundra is dense and continuous and leads to large, fast spreading fires (Duchesne and Hawkes 2000, Racine et al 1987). Racine (1979) found much variation in burn intensity on a landscape scale on the Seward Peninsula, from completely unburned to intensely burned. These patterns are related to variations in topography and the composition, moisture content and soil organic accumulations of the plant communities. Fires in Eriophorum tussock tundra types tend to be light because of the wet soil profile (Wein 1971). Burns in this type usually consume all aerial woody and herbaceous plant material and litter; regeneration is vigorous via rhizomes and root sprouts. Racine (1979) found that burning was generally less severe in the tussock-shrub and sedge-shrub tundras than in the birch and ericaceous shrub tundra of the Seward Peninsula. He found that tundra burns were patchy, with unburned communities and unburned patches within burned communities. More fires occur near the forest-tundra ecotone and spread further if trees are present (Heinselman 1981). Wein (1976) reports that July and August are the most common months for lightning fires in tundra ecosystems, while Racine et al (1983; 1985) found that distinct fire seasons occur in both June and July in the Noatak River watershed. Subsidence and thermal erosion following fire is usually minimal in tundra ecosystems (Walker 1996). In most areas of tussock-shrub tundra on the Seward Peninsula, less than one half of accumulated organic soil layer was removed by fire (Racine 1979). Thaw depths increased to reach into the mineral soils, but were not greatly increased except where organics were removed. Frost features were made more conspicuous, and soil nutrient concentrations (K and P) increased locally. Mean fire return interval estimates for tussock tundra ecosystems include: -50-300yrs (personal communication FRCC experts’ workshop March 2004) -260yr (s.d. 170) fire return interval for past 1500yrs for Noatak National Preserve (preliminary data from Higuera et al. 2008) -180-1460yrs in forest shrubzone and 9,320yrs in shrub subzone in northern Quebec; shorter cycle west of Hudson’s Bay/in interior zone (Payette et al 1989) -612yrs for Noatak River watershed (all vegetation types) (Racine et al 1983) -611yr fire rotation for Noatak River watershed (all vegetation below 600m which is predominantly tundra) (Racine et al 1985) -Fire interval yet to be determined (Racine et al 1987) -Rapid recovery following fire makes fire frequency difficult to determine (Wein 1971) -The fire regime of tundra systems are likely quite variable from one region to another making generalizations difficult (Viereck and Schandelmeier 1980) On interior and southcentral Alaska Tussock Tundra sites the thaw pond cycle (disturbance leads to thawing of permafrost and ponding) and paludification (sphagnum layer buildup and saturation) are important disturbances. On the Seward Peninsula and western AK, frost action creates polygonal ground and other periglacial features and is a widespread, small-scale and continuous disturbance. Change in the arctic and subarctic climate is another source of disturbance that is currently affecting tundra ecosystems. Adjacency or Identification Concerns *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 144 of 204 Native Uncharacteristic Conditions Scale Description Vegetation found in large patches Wien (1976) reports many tundra fires in the 1-100ha size range and few large (thousands of ha) fires. Racine (1979) reports that in 1977, lightning-caused fires burned 35,480ha on the Seward Peninsula, with one fire burning 9440ha. Jandt and Meyers (2000) report that large fires (>200,000ha) occur about every 10yrs in the Buckland Valley and surrounding highlands of the Seward Peninsula. Racine et al (1983) found that 40 fires burned 100,000ha (1000km2) in the 30,000km2 watershed of the Noatak River between 1956 and 1981. Racine et al (1985) found a minimum fire size of 0.4ha, a maximum fire size of 45,800ha and a mean fire size of 1310ha from 1956-1983 in the Noatak River watershed, an area dominated by tundra vegetation. Of the 79 fires in analyzed by Racine et al (1985), nearly half were between 1-10ha in size. Forty-three percent of wildland fires occurring in interior Alaska occur in treeless areas, primarily tundra bogs and fens (Viereck 1975). Issues/Problems Experts at the Fairbanks meeting indicated the need for a late seral sphagnum tussock bog stage that would develop as a result of paludification in class B. The VDDT model does not include that stage because it would be hard to distinguish for mapping using LANDFIRE methods. This model is based in part on data from the Seward Peninsula, which is not part of the boreal region. In particular, much of the data on fire regimes comes from the Seward and Noatak areas, both of which are part of the arctic region. Comments This model was based on input from the experts who attended the LANDFIRE Fairbanks (Nov. 07) modeling meeting and refined by Jennifer Allen. Much of the text in the Disturbance Description, Scale Description and Succession Class Descriptions in this report were taken from the Tussock Tundra 1 PNVG model description (Murphy and Witten 2006). This model was origioanlly developed with two herbaceous classes: a Mesic Graminoid Tussock and a Wet Graminoid Tussock class. When it was determined that the LANDFIRE project would not be able to distingusish these classes using a Viereck et al. (1992) based existing vegetation map, these two classes were lumped into what is now class A. Vegetation Classes *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 145 of 204 Class A Structure Data (for upper layer lifeform) Min Max 85 % Early Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Tree Size Class None Herbaceous Shrub Tree Indicator Species* and Canopy Position ERVA4 CABI5 Upper Upper Herbaceous Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description Zero years plus Mesic graminoid tussocks dominates for the first 10yrs or longer. First year following fire Eriophorum (cottongrass) and Carex spp. (sedges) regrow via rhizomes, most vascular species begin to recover, shrubs sprout from rootstock. Sedges often capture site 6-10yrs post fire. Grasses (Calamagrostis and Arctagrostis) are locally important following fire. A high severity fire can lead to increased shrub development (class C), whereas a low to moderate severity fire typically leads to the development of a wet graminoid tussock stage. Wet graminoid tussock can dominate some sites developing about 10yrs post fire. It is the most common stage after a low to moderate severity fire. Over time paludification results in the development of a sphagnum tussock bog (the model does not include the sphagnum tussock bog stage because it would be hard to distinguish for mapping using LANDFIRE methods). This class persists in the absence of disturbance or will follow an alternate successional pathway to class B (probability = 0.001). Replacement MFRI = 175yrs. Class B Structure Data (for upper layer lifeform) Min Max 15 % Late Development 1 All Structures Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position CHCA2 VAOX ERVA4 CABI5 Upper Upper Upper Upper Dwarf Shrub (< 20 cm) None Dwarf Shrub (< 20 cm) Tree Size Class Upper layer lifeform differs from dominant lifeform. Description 10yrs+ Dwarf shrub, tussock. Most common stage after a high severity fire. Tussocks dominated by Eriophorum (cottongrass), Carex spp. (sedges). Common shrubs include Oxycoccus spp. (Vaccinium oxycoccus), Chamaedaphne calyculata, Betula nana, Ledum decumbens and Vaccinium. Lichens begin to re-establish but do not reach former abundance until 50-120yrs following fire. Fire is difficult to detect even in the early stages of this class, however the proportions of species differs from the pre-burn community, with very few lichens, fewer shrubs and more sedges, grasses and cottongrass. Former abundances of all species are typically reached 50-120yrs post fire. This class persists in the absence of disturbance. Replacement MFRI = 175yrs. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 146 of 204 Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Min Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: IV Historical Fire Size (acres) Avg 0 Min 0 Max 0 Sources of Fire Regime Data Literature Local Data Expert Estimate Fire Intervals Replacement Avg FI Min FI Max FI Probability 175 0.00571 175 0.00573 Percent of All Fires 100 Mixed Surface All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 147 of 204 Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Duchesne L.C. and B.C. Hawkes. 2000. Fire in northern ecosystems. In: Brown, J.K. and J.K. Smith (eds.) Wildland fire in ecosystems: effects of fire on flora. Gen. Tech. Rep. RMRS-GTR-42-vol 2. Ogden, UT: USDA Forest Service, Rocky Mountain Research Station. 257 pp. Heinselman, M.L. 1981. Fire and succession in the conifer forests of northern North America. In: West, D.C., H.H. Shugart, and D.B. Botkin. Forest succession: concepts and application. Springer-Verlag, New York. Chapter 23. Higuera, P.E., M. Chipman, J. Allen, S. Rupp and F.S. Hu. 2008. Preliminary data provided by Jenifer Allen from a study of tundra fire regimes in the Noatak National Preserve since 6000 years before present. Jandt, R.J. and R.C. Meyers. 2000. Recovery of lichen in tussock tundra following fire in Northwestern Alaska. USDOI BLM-Alaska Open File Report 82. Murphy, K.A. and E. Witten. 2006. Tussock Tundra 1. In Fire Regime Condition Class (FRCC) Interagency Guidebook Reference Conditions. Available at www.frcc.gov. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Payette, S., C. Morneau, L. Sirois and M. Desponts. 1989. Recent fire history of the northern Quebec biomes. Ecology. 70:656-673. Personal communication experts’ workshop, March 2-4 2004. Fire Regime Condition Class (FRCC) interagency experts’ workshop to develop and review Potential Natural Vegetation (PNV) groups for Alaska. Anchorage, AK. Racine, C.H., L.A. Johnson and L.A. Viereck. 1987. Patterns of vegetation recovery after tundra fires in northwestern Alaska, USA. Arctic and Alpine Research. 19: 461-469. Racine. 1979. Climate of the Chucki-Imuruk area. Pages 32-37 in H. R. Melchior, ed., Biological Survey of the Bering Land Bridge National Monument. Alaska Cooperative Park Studies Unit, University of Alaska Fairbanks, Fairbanks, AK. Racine, C.H., W.A. Patterson III and J.G. Dennis. 1983. Permafrost thaw associated with tundra fires in northwest Alaska. Pages 1024-1029 in Proceedings, Permafrost, Fourth International Conference. National Academy Press, Washington, D.C. Racine, C.H., J.G. Dennis and W.A. Patterson III. 1985. Tundra fire regimes in the Noatak River Watershed, Alaska: 1956-83. Arctic 38: 194-200. Racine, C., J.L. Allen and J.G. Dennis. 2006. Long-term monitoring of vegetation change following tundra fires in Noatak National Preserve, Alaska. Arctic Network of Parks inventory and monitoring program, *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 148 of 204 National Park Service, Alaksa Region. Report no. NPS/AKRARCN/NRTR-2006/02. Viereck, L.A. 1975. Forest ecology of the Alaska Taiga. In: Proceedings of the circumpolar conference on northern ecology; 1975 September; Ottawa, ON. National Research Council of Canada: I-1 to I-22. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. Viereck, L.A. and Schandelmeier, L.A. 1980. Effects of fire in Alaska and adjacent Canada--a literature review. BLM-Aalska Tech. Rep. No. 6. Anchorage, Alaska: U.S. Department of the Interior, Bureau of Land Management. 124 pp. Walker, D.A. 1996. Disturbance and Recovery of Arctic Alaskan Vegetation. In: JF Reynolds and JD Tenhunen eds, Landscape function and disturbance in arctic tundra. Ecological studies vol. 120, Springer Verlag Berlin Heidleberg. Wein, R.W. 1976. Frequency and characteristics of arctic tundra fires. Arctic, 29: 213-222. Wein, R.W. 1971. Panel discussion: In: Slaughter, C.W., Barney, Richard J. and Hansen, G.M. (editors). 1971. Fire in the Northern Environment – A Symposium. Sponsors: Alaska Forest Fire Counc il and Society of American Foresters. Held at: The University of Alaska, Fairbanks. April 13-14, 1971. Published by: Pacific. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 149 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316300 Western North American Boreal Wet Black Spruce-Tussock Woodland This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Date 3/11/2008 Modeler 1 Kori Blankenship kblankenship@tnc.org Modeler 2 Robert Lambrecht Robert_Lambrecht@fws Reviewer Stuart Chapin .gov Lisa_Saperstein@fws. gov fffsc@uaf.edu Reviewer Modeler 3 Map Zone 73 Vegetation Type Wetlands/Riparian Dominant Species* PIMA BENA LEPAD VAUL Reviewer Lisa Saperstein General Model Sources VAVI ERVA4 CABI5 RUCH Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range Boreal Wet Black Spruce-Tussock Woodland is found throughout interior AK from the southern slopes of the Brooks Range to southcentral AK (but not including the sub-boreal region) and west to the limit of tree growth (NatureServe 2008). Biophysical Site Description This type is found on north-facing slopes, gentle hills, and inactive alluvial surfaces underlain by permafrost (NatureServe 2008). Soils are poorly drained and consist of tussocks over peat or mineral soil (Jorgenson et al. 2001, Boggs and Sturdy 2005). Vegetation Description Picea mariana is the dominant tree species and occurs in open stands. Tussock-forming sedges contribute at least 25% of the vegetation cover (NatureServe 2008). Common understory shrubs include Betula nana (including B. glandulosa), Ledum palustre ssp. decumbens., Vaccinium uliginosum and V. vitis-idaea. Herbaceous species include Eriophorum vaginatum, Carex bigelowii and Rubus chaememorus (NatureServe 2008). Mosses may be abundant and include Sphagnum spp. and Hylocomium splendens (Jorgenson et al. 2001, Boggs and Sturdy 2005). Disturbance Description Fire is the primary disturbance in this system. Fire severity is generally sufficient to kill the overstory. Mean fire return interval estimates in boreal Alaska range from 25-130yrs (Rowe et al.1974, Yarie 1983, Heinselman 1978, Heinselman 1981, Viereck 1983, Viereck 1986). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 150 of 204 Under appropriate conditions, this system can originate from a very late seral stage of the Boreal Black Spruce Dwarf-tree Peatland system. Adjacency or Identification Concerns This system tends to occur on a continuum between the Boreal Black Spruce Dwarf-tree Peatland system and the Boreal Low Shrub-Tussock Tundra system. Boreal Wet Black Spruce-Tussock Woodland occurs on sites that are slightly higher and drier than tussock-shrub sites and slightly lower and wetter than wet black spruce sites. The herb and shrub classes of this system are similar in structure and composition to the herb and shrub classes of the Boreal Low Shrub-Tussock Tundra system, but the Boreal Low Shrub-Tussock Tundra system occurs where site conditions prevent trees from invading. Native Uncharacteristic Conditions Scale Description Large patch (small patch) Issues/Problems Comments This model was based on input from the experts who attended the LANDFIRE Fairbanks (Nov. 07) modeling meeting and refined by Robert Lambrecht. Torre Jorgenson provided some information on the relationships between this system and adjacent systems. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 5% Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position ERVA4 CABI5 RUCH Upper Upper Upper Herbaceous Height Herbaceous Tree Size Class None Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description 0-9yrs This class is characterized by tussock forming sedges. Common species include Eriophorum vaginatum, Carex bigelowii and Rubus chaememorus. Succession to class B. Replacement MFRI = 150yrs. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 151 of 204 Class B Structure Data (for upper layer lifeform) Min Max 20 % Mid Development 1 All Structures Cover Open Shrub (25-74% shrub cover) Open Shrub (25-74% shrub cover) Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position BENA LEPAD VAUL VAVI Shrub Tree Dwarf Shrub (< 20 cm) Tree Size Class Low Shrub (20 cm to 1.5 m) Seedling/Sapling <5" Upper Upper Upper Upper Upper layer lifeform differs from dominant lifeform. Description 10-49yrs Shrubs resprout quickly after fire, becoming the dominant, upper-level canopy layer in 10-15yrs. Common species include Betula nana, Ledum palustre ssp. decumbens., Vaccinium uliginosum and V. vitis-idaea. Succession to class C. Replacement MFRI = 150yrs. Class C Structure Data (for upper layer lifeform) Min Max 75 % Late Development 1 Open Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position PIMA BENA LEPAD VAUL Woodland (10-24% tree cover) Tree Size Class Upper Lower Lower Lower Open (25-59% tree cover) Dwarf Tree (< 3 m) Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. Description 50yrs+ This class is characterized by mature black spruce tussock forest (spruce cover generally 10-25%; tussock cover >25%). Overstory is dominated by Picea mariana. It is during this class when organic material begins to accumulate and a distinctive “active layer” appears, affecting fire behavior depending upon how dry it gets. This class can persist in the absence of disturbance. Replacement MFRI = 150yrs. Mixed fire (MFRI = 500yrs) will maintain this class. Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 152 of 204 Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals IV Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Avg FI Min FI Max FI Probability 152 667 0.00658 0.0015 124 0.00809 Percent of All Fires 81 19 Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Boggs, K. and Sturdy, M. 2005. Plant associations and post-fire vegetation succession in Yukon-Charley Rivers National Preserve. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage. Prepared For: National Park Service, Landcover Mapping Program, National Park Service-Alaska Support Office, Anchorage, AK 99501. Heinselman, M.L. 1978. Fire in wilderness ecosystems. In: J. C. Hendee, G. H. Stankey, and R. C. Lucas. Wilderness Management. USDA Forest Service, Miscellaneous Publication 1365. Heinselman, M.L. 1981. Fire and succession in the conifer forests of northern North America. In: West, D.C., H.H. Shugart, and D.B. Botkin. Forest succession: concepts and application. Springer-Verlag, New York. Chapter 23. Jorgenson, M.T., J.E. Roth, M.D. Smith, S. Schlentner, W. Lentz and E.R. Pullman. 2001. An ecological land survey for Fort Greely, Alaska. U.S. Army Cold Regions Research and Engineering Laboratory, Hanover, NH. ERDC/CRREL TR-01-04. 85 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Rowe, J.S., J.L Bergsteinsson, G.A. Padbury and R. Hermesh. 1974. Fire Studies in the Mackenzie Valley. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 153 of 204 ALUR Report 73-74-61. Arctic Land Use Research Program, Department of Indian Affairs and Human Development, Ottawa, Canada. 123 pp. Viereck, L.A. 1983. The effects of fire in black spruce ecosystems of Alaska and northern Canada. In: Wein, Ross W.; MacLean, David A., eds. The role of fire in northern circumpolar ecosystems. New York: John Wiley & Sons Ltd.: 201-220. Chapter 11. Viereck, L.A. Van Cleve, K. Dyrness, C.T. 1986. Forest Ecosystems in the Alaska Taiga. In: Van Cleve, K., Chapin F.S. III, Flanagan, P.W. (and others), eds. Forest Ecosystems in the Alaskan taiga: a synthesis of structure and function. New York: Springer-Verlag; 1986: 22-43. Yarie J. 1983. Forest community classification of the Porcupine River drainage, interior Alaska, and its application to forest management. USDA Forest Service GTR PNW-154. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 154 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316310 Western North American Boreal Alpine Dwarf-Shrub Summit This BPS is lumped with: see below This BPS is split into multiple models: This BpS is lumped with: Western North American Boreal Alpine Dwarf-ShrubLichen Shrubland General Information Contributors (also see the Comments field Modeler 1 Kori Blankenship Modeler 2 Tina Boucher Modeler 3 kblankenship@tnc.org antvb@uaa.alaska.edu Reviewer Reviewer Map Zone 73 Upland Shrubland Dominant Species* 4/16/2008 Reviewer Vegetation Type DRYAS VAUL EMNI VAVI Date General Model Sources DILA LOPR SALIX ARRU Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system is found in the alpine and subalpine areas of the boreal and sub-boreal regions in AK. To the southwest, the range includes the Ahklun Mts and to the west it extends to the Nulato hills (Ecoregions 9 and 7, Nowacki et al. 2001). Biophysical Site Description These systems occur on exposed, windswept summits and ridges on alpine and subalpine sites. Soils are thin, stony, and well drained to excessively well-drained. These sites typically do not accumulate much winter snow (Viereck et al 1992). Vegetation Description Canopy cover can be sparse due to extreme exposure. Common shrubs include Dryas spp. (e.g. Dryas integrifolia), Vaccinium uliginosum, Empetrum nigrum, Vaccinium vitis-idaea, Diapensia lapponica, Loiseuria procumbens, dwarf Salix spp. (e.g. Salix arctica, S. rotundifolia and S. reticulata), Arctostaphylos rubra and A. alpina. Exposed rock and lichens can be abundant. Common lichens include Cladina rangiferina, C. stellaris, Cetraria cucullata, Stereocaulon spp., Alectoria nigricans and Thamnolia vermiculata. Herbaceous species include Hierochloe alpina, Polygonum bistorta, Anemone spp., Festuca spp. And Luzula spp. Mosses may be present but do not contribute much cover (Viereck et al 1992). Disturbance Description Alpine shrub systems likely represent a relatively stable topopoedaphic climax, but little is known about their successional dynamics (Viereck et al. 1992). Vegetation in these areas is controlled by wind *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 155 of 204 desiccation and a short growing season. There is little information about fire in this system. It is possible that fires caused by lightning strikes could affect small patches of vegetation but likely would not spread due to the sparse cover of fine fuel and barren areas acting as fire breaks. Adjacency or Identification Concerns Adjacent systems include Western North American Boreal Alpine Dwarf-Shrubland or barren alpine classes (including talus and bedrock). Since this BpS is a combination of two systems, the identifying criteria for both systems apply: identifiers for the Alpine Dwarf-Shrub-Lichen Shrubland system are greater than 25% lichen cover and greater than 25% dwarf shrub cover, identifiers for the Alpine DwarfShrub Summit system are less than 25% vascular plant cover and abundant exposed rock. Native Uncharacteristic Conditions Scale Description Vegetation can be found in small or large patches. Fires are typically quite small and their spread is inhibited by lack of fuel continuity. Issues/Problems Comments This model was developed by Kori Blankenship in consultation with Tina Boucher. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 100 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position VAUL EMNI VAVI DRYAS Upper Upper Upper Upper Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Dwarf Shrub (< 20 cm) Low Shrub (20 cm to 1.5 m) Height Tree Size Class None Upper layer lifeform differs from dominant lifeform. Description Zero years plus Sparse, open or closed dwarf shrubs dominate. Exposed rock and lichens can be abundant. Commons species listed in the vegetation description. This BpS is relatively stable over time. Continual wind disturbance (probability = 1) maintains this BpS. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 156 of 204 Class B Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Cover Indicator Species* and Canopy Position Height Tree Size Class Shrub Tree Upper layer lifeform differs from dominant lifeform. Description Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 157 of 204 Fire Regime Group**: Fire Intervals NA Avg FI Min FI Max FI Probability Percent of All Fires Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Nowacki, G., P. Spencer, M. Fleming, T. Brock and T. Jorgenson. 2001. Unified ecoregions of Alaska. U.S. Department of the Interior, U.S. Geological Survey. Open file-report 02-297. Two page map. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 158 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316330 Western North American Boreal Alpine Mesic Herbaceous Meadow This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Tina Boucher antvb@uaa.alaska.edu colleen_ryan@tnc.org Modeler 2 Colleen Ryan Modeler 3 Upland Grassland/Herbaceous Dominant Species* 4/24/2008 Reviewer Reviewer Reviewer Vegetation Type General Model Sources CABI5 BENA ARAL2 EMNI Date Literature Local Data Expert Estimate Map Zone 73 Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system occurs on gentle slopes in subalpine and lower alpine environments throughout boreal AK (NatureServe 2008). Biophysical Site Description This system occurs on alpine and subalpine slopes. It tends to occur in small patches in a matrix with dwarf or low shrub systems. Soils are moist to mesic. Vegetation Description Carex bigelowii is the dominant species. Other common species may include Luzula confusa and lichens. Dwarf shrubs such as Arctostaphylos alpina, Empetrum nigrum, Salix pulchra and Betula nana are usually present, but contribute to <25% of the canopy cover (NatureServe 2008, Boggs and Sturdy 2005). Wetter sites may include more sedges and Salix spp. This system may form a mosaic with dwarf and low shrub systems. Disturbance Description Little is known about the disturbance regime of this system. It tends to occur adjacent to low shrub and alpine tundra systems, and even in a patchy mosaic with dwarf and low shrub systems. Because these adjacent systems burn, it is likely that some fire will carry through the Alpine Mesic Herbaceous Meadow system, especially in drier periods. Fire frequency is likely to depend on the adjacent vegetation. For this model, the fire return interval was estimated at twice that of the adjacent Western North American Boreal Mesic Scrub Birch-Willow Shrubland BpS. Little is known about post-fire dynamics, but Carex bigelowii is expected to maintain its dominance. Dwarf shrubs are likely to resprout or establish gradually over time, but will remain a minor component of *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 159 of 204 the system. Adjacency or Identification Concerns There is no species overlap between boreal and Sub-boreal herbaceous alpine meadow types. Slope position may also be different: Carex macrochaeta types usually occur just above subalpine alder on straight or concave slopes. This system often occurs in a patchy mosaic with low and dwarf shrub types (Betula nana and ericaceous dwarf shrub types). Native Uncharacteristic Conditions Scale Description Small to large patch Issues/Problems Comments Because no data are available about the dynamics of this fairly minor system, this model was based on best estimates, considering vegetation and site characteristics and the dynamics of adjacent vegetation types. Page Spencer is a suggested reviewer for this system. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 100 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position CABI5 BENA ARAL2 EMNI Upper Upper Upper Upper Herbaceous Height Herbaceous Tree Size Class None Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description Zero years plus Alpine Herbaceous-Dwarf Shrub communities may include a diverse mix of species and species assemblages. The class Indicator Species listed are not true indicators -- refer to the Vegetation Description for species information. No disturbances modeled. Mesic herbaceous. In interior AK, Carex bigelowii is the dominant species. Other herbaceous species, including Luzula confusa, are frequently present, along with a well-developed lichen layer. Dwarf shrubs such as Arctostaphylos alpina, Empetrum nigrum, and Betula nana are present, but contribute to <25% of the canopy cover (Boggs and Sturdy 2005). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 160 of 204 Although this system tends to be fairly moist, litter accumulation will provide enough fuel to carry a fire during dry periods. Because the most likely source of fire is the adjacent Boreal Low Shrub system, fire frequency for this system was assigned to be 1/2 that of the Boreal Low Shrub BpS. This BpS is relatively stable over time. Replacement MFRI = 400yrs. Class B Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Cover Indicator Species* and Canopy Position Height Tree Size Class Shrub Tree Upper layer lifeform differs from dominant lifeform. Description Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Indicator Species* and Canopy Position Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 161 of 204 Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals V Replacement Historical Fire Size (acres) Avg FI Min FI Max FI Probability 400 0.0025 400 0.00252 Percent of All Fires 99 Mixed Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Boggs, K. and Sturdy, M. 2005. Plant associations and post-fire vegetation succession in Yukon-Charley Rivers National Preserve. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage. Prepared For: National Park Service, Landcover Mapping Program, National Park Service-Alaska Support Office, Anchorage, AK 99501. Boggs, K., A. Garibaldi, J. Stevens, J. Grunblatt and T. Helt. 2001. Denali National Park and Preserve Landcover mapping project. Volume 2: Landcover classes and plant associations. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage, 707 A Street, Anchorage, AK. 164 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 162 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316351 Western North American Boreal Alpine Ericaceous Dwarf-Shrubland - Complex This BPS is lumped with: see below This BPS is split into multiple models: Western North American Boreal Alpine Dryas Dwarf-Shrubland (BpS 1634) General Information Contributors (also see the Comments field Modeler 1 Kori Blankenship Modeler 2 Tina Boucher kblankenship@tnc.org antvb@uaa.alaska.edu Modeler 3 4/16/2008 Reviewer Reviewer Reviewer Map Zone 73 Vegetation Type Upland Shrubland Dominant Species* CATE11 EMNI VAUL HAST3 Date General Model Sources ARCTO3 DRIN4 DROC LEPAD Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This type occurs from subalpine to alpine locations in the boreal and sub-boreal regions of AK. Biophysical Site Description This BpS is commonly found in alpine valleys and on sideslopes, low summits and ridges. Sites are welldrained and mesic to somewhat dry and are likely to retain late-lying snow. Vegetation Description This type eoncompasses a wide range of species and plant communities. Ericaceous or dryas (especially Dryas integrifolia and/or Dryas octopetala) dwarf shrubs typically dominate. Total lichen cover is <25% and may include Cetraria, Cladina, Cladonia and Stereocaulon. Common dwarf shrub include Cassiope tetragona (more common north of the Alaska Range), Empetrum nigrum, Vaccinium uliginosum, Harrimanella stellariana (more common south of the Alaska Range) and Arctostaphylos spp. Other shrubs that may be common include Betula nana, Diapensia lapponica, Dryas octopetala, Ledum palustre ssp. decumbens, Vaccinium vitis-idaea, Salix reticulata, S. phlebophylla, S. rotundifolia, S. arctica and Oxytropis nigrescens. Common herbaceous species include Hierochloe alpine, Arnica lessingii, Carex bigelowii, Carex microchaeta, Senecio lugens, Minuartia arctica, Anemone parviflora, Ligusticum mutellinoides ssp. alpinum, Castilleja elegans, Poa arctica, Trisetum spicatum, Silene acaulis, Saxifraga spp., Campanula lasiocarpa, Anemone parviflora, Senecio lugens and Polygonum bistorta. Mosses such as Aulacomnium palustre, Hylocomium splendens, Pleurozium schreberi and Polytrichum and Rhacomitrium spp. may be common. Cassiope and Harimanella tundra sites occur on terrain that is well protected by snow in the winter, and often remains snow covered until the middle of the growing season (Viereck et al. 1992). *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 163 of 204 Disturbance Description Alpine shrub systems likely represent a relatively stable topopoedaphic climax, but little is known about their successional dynamics (Viereck et al. 1992). Vegetation in these areas is controlled by the alpine environment and short growing season. It is possible that fires caused by lightning strikes could affect small patches of vegetation but little is known about the frequency, severity or seasonality of fires in this BpS. Adjacency or Identification Concerns This BpS is distinguished by <25% cover lichen and >20% cover dryas dwarf shrubs. Adjacent systems include Western North American Boreal Alpine Dwarf-Shrub Summit, Western North American Boreal Alpine Mesic Herbaceous Meadow, Alaska Sub-boreal and Maritime Alpine Mesic Herbaceous Meadow or barren alpine classes (including talus or bedrock). Native Uncharacteristic Conditions Scale Description Large patch Issues/Problems This BpS represents two Ecological Systems (Dryas Dwarf-Shrubland and Alpine Ericaceous DwarfShrubland) and would be more appropriately named Alpine Dwarf Shrubland. Comments This model was created by Kori Blankenship in consultation with Tina Boucher. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 100 % Early Development 1 All Structures Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position CATE11 EMNI VAUL DRIN4 Upper Upper Upper Upper Dwarf Shrub (< 20 cm) Tree Size Class Low Shrub (20 cm to 1.5 m) None Upper layer lifeform differs from dominant lifeform. Description Zero years plus Open or closed dwarf ericaceous or dryas shrubs dominate. Common species are listed in the vegetation description. This BpS is relatively stable over time. Continual wind disturbance (probability = 1) maintains this BpS. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 164 of 204 Class B Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Cover Indicator Species* and Canopy Position Height Tree Size Class Shrub Tree Upper layer lifeform differs from dominant lifeform. Description Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 165 of 204 Fire Regime Group**: Fire Intervals NA Avg FI Min FI Max FI Probability Percent of All Fires Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Boggs, K., A. Garibaldi, J. Stevens, J. Grunblatt and T. Helt. 2001. Denali National Park and Preserve Landcover mapping project. Volume 2: Landcover classes and plant associations. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage, 707 A Street, Anchorage, AK. 164 pp. Boggs, K. and Sturdy, M. 2005. Plant associations and post-fire vegetation succession in Yukon-Charley Rivers National Preserve. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage. Prepared For: National Park Service, Landcover Mapping Program, National Park Service-Alaska Support Office, Anchorage, AK 99501. DeVelice, R.L., Hubbard, C.J., Boggs, K. et al. 1999. Plant community types of the Chugach National Forest. Tech. Publ. R10-TP-76. Juneau, AK: USDA Forest Service, Alaska Region. 375 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 166 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316371 Western North American Boreal Alpine Floodplain - Lower Elevations This BPS is lumped with: This BPS is split into multiple models: Western North American Boreal Alpine Floodplain was split into a lower elevations and an upper elevations BpS model. The lower elevation model applies in the subalpine zone within the tall shrub zone. The upper elevations model applies above the elevational limit of tall shrubs. General Information Contributors (also see the Comments field Modeler 1 Tina Boucher Modeler 2 Kori Blankenship antvb@uaa.alaska.edu kblankenship@tnc.org 4/15/2008 Reviewer Reviewer Reviewer Modeler 3 Map Zone 73 Vegetation Type Upland Shrubland Dominant Species* Date General Model Sources SAAL SALIX ALVIS CHLA13 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range The BpS is found in alpine and subalpine areas in the boreal and boreal transition regions of AK. Biophysical Site Description This system includes active and inactive alpine and subalpine floodplains of glacially and non-glacially fed streams (NatureServe 2008). Soils develop on alluvium and are typically shallow and well-drained (NatureServe 2008). This model applies to Alpine Floodplains that occur just above treeline, in the tall shrub zone. Vegetation Description This system includes a range of floodplain vegetation including shrub (dwarf, low and tall), mesic herbaceous meadow, early seral forbs and barren gravel. Common species include Salix alaxensis, other Salix spp., Alnus viridis ssp. Sinuata, Betula nana, Chamerion latifolium, Lupinus spp. (L. nootkatnesis and L. arcticus), Mertensia paniculata, Crepis spp (C. nana and C. elegans) Achillea millefolium spp. Borealis, Erigeron acris and a variety of grasses (NatureServe 2008). Disturbance Description Frequent river channel migration and associated flooding and fluvial processes constitute the major disturbances in this type (NatureServe 2008). The probability of flooding is assumed to be higher in alpine floodplains compared with lower elevation floodplains because the alpine floodplains tend to have higher gradients and the landscape absorbs less runoff due to steep slopes and typically coarse substrates. The overall return interval for flooding in the alpine floodplain is estimated at about 20yrs, compared with 70*Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 167 of 204 75yrs in the floodplain forest and shrub systems. Alpine floodplains and riparian areas generally act as fire breaks. Adjacency or Identification Concerns Floodplain systems may occur in the active and inactive part of the riparian zone, but abandoned floodplains are considered part of the adjacent upland. This model applies to the lower elevation variant of this system, which typically occurs in the subalpine zone. Adjacent systems typically include Alaska SubBoreal Mesic Subalpine Alder Shrubland and Western North American Boreal Mesic Scrub Birch-Willow Shrubland - Boreal, but may also include Western North American Boreal Alpine Dwarf-Shrubland or Alaska Sub-boreal and Maritime Alpine Mesic Herbaceous Meadow, Talus and Bedrock. Native Uncharacteristic Conditions Scale Description Linear Issues/Problems The probability of flooding in the model is a best guess, not based on literature. Comments This model was developed based on input from the experts who attended the LANDFIRE Anchorage (Dec. 07) modeling meeting and refined by Tina Boucher and Kori Blankenship. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 40 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position CHLA13 LUPIN MEPA CREL Upper Upper Upper Upper Herbaceous Height Herbaceous Tree Size Class None Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description 0-14yrs Although it is not modeled, because LANDFIRE does not map sparsely vegetated areas, this class should be preceded by a sparse/gravel bar phase. This herbaceous class represents early seral vegetation that would come in on gravel bars or other sparsely vegetated areas in the floodplain. Common species include Chamerion latifolium, Lupinus spp. (L. nootkatnesis and L. arcticus), Mertensia paniculata, Crepis spp (C. nana and C. elegans) Achillea millefolium spp. borealis, Erigeron acris and a variety of grasses. Vegetation cover is generally open (10-50%) with large areas of exposed alluvium. Deterministic succession to class B. Flooding (probability = 0.08), represented by Option 1 in the model, resets the age of this class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 168 of 204 Class B Structure Data (for upper layer lifeform) Min Max 60 % Late Development 1 All Structures Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position SAAL SALIX ALVIS BENA Shrub Tree Low Shrub (20 cm to 1.5 m) Tree Size Class Tall Shrub (>1.5 m) None Upper Upper Upper Upper Upper layer lifeform differs from dominant lifeform. Description 15yrs+ Willow and alder low and tall shrubs. Common shrub species may include Salix alaxensis and other Salix spp., Alnus viridis ssp. sinuata and Betula nana. This class persists in the absence of disturbance. Flooding (probability = 0.025), represented by Option 1 in the model, causes a transition to class A. Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 169 of 204 Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals NA Avg FI Min FI Max FI Probability Percent of All Fires Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Flooding Other (optional 2) References NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 170 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316372 Western North American Boreal Alpine Floodplain - Higher Elevations This BPS is lumped with: This BPS is split into multiple models: Western North American Boreal Alpine Floodplain was split into a lower elevations and an upper elevations BpS model. The lower elevation model applies in the subalpine zone within the tall shrub zone. The upper elevations model applies above the elevational limit of tall shrubs. General Information Contributors (also see the Comments field Modeler 1 Tina Boucher Modeler 2 Kori Blankenship antvb@uaa.alaska.edu kblankenship@tnc.org 4/15/2008 Reviewer Reviewer Reviewer Modeler 3 Map Zone 73 Vegetation Type Upland Shrubland Dominant Species* Date General Model Sources SAAL SALIX ALVIS CHLA13 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range The BpS is found in alpine and subalpine areas in the boreal and boreal transition regions of AK. Biophysical Site Description This system includes active and inactive alpine and subalpine floodplains of glacially and non-glacially fed streams (NatureServe 2008). Soils develop on alluvium and are typically shallow and well-drained (NatureServe 2008). This model applies to Alpine Floodplains that occur above the elevational limit of tall shrubs. Vegetation Description This system includes a range of floodplain vegetation including shrub (dwarf, low and tall), mesic herbaceous meadow, early seral forbs and barren gravel. Common species include Salix alaxensis, other Salix spp., Alnus viridis ssp. sinuata, Betula nana, Chamerion latifolium, Lupinus spp. (L. nootkatnesis and L. arcticus), Mertensia paniculata, Crepis spp (C. nana and C. elegans) Achillea millefolium spp. borealis, Erigeron acris and a variety of grasses (NatureServe 2008). Disturbance Description Frequent river channel migration and associated flooding and fluvial processes constitute the major disturbances in this type (NatureServe 2008). The probability of flooding is assumed to be higher in alpine floodplains compared with lower elevation floodplains because the alpine floodplains tend to have higher gradients and the landscape absorbs less runoff due to steep slopes and typically coarse substrates. The overall return interval for flooding in the alpine floodplain is estimated at about 20yrs, compared with 70*Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 171 of 204 75yrs in the floodplain forest and shrub systems. Alpine floodplains and riparian areas generally act as fire breaks. Adjacency or Identification Concerns Floodplain systems may occur in the active and inactive part of the riparian zone, but abandoned floodplains are considered part of the adjacent upland. This model applies to the variant of this system that occurs above the elevational limit of tall shrubs. Adjacent systems may include Western North American Boreal Alpine Dwarf-Shrubland, Western North American Boreal Alpine Mesic Herbaceous Meadow, Alaska Sub-boreal and Maritime Alpine Mesic Herbaceous Meadow, Maritime High Alpine Herbaceous, Talus and Bedrock. Native Uncharacteristic Conditions Scale Description Linear Issues/Problems The probability of flooding in the model is a best guess, not based on literature. Comments This model was developed based on input from the experts who attended the LANDFIRE Anchorage (Dec. 07) modeling meeting and refined by Tina Boucher and Kori Blankenship. Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 35 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position CHLA13 LUPIN MEPA CREL Upper Upper Upper Upper Herbaceous Height Herbaceous Tree Size Class None Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description 0-14yrs Although it is not modeled, because LANDFIRE does not map sparsely vegetated areas, this class should be preceded by a sparse/gravel bar phase. This herbaceous class represents early seral vegetation that would come in on gravel bars or other sparsely vegetated areas in the floodplain. Common species include Chamerion latifolium, Lupinus spp. (L. nootkatnesis and L. arcticus), Mertensia paniculata, Crepis spp (C. nana and C. elegans) Achillea millefolium spp. borealis, Erigeron acris and a variety of grasses. Vegetation cover is generally open (10-50%) with large areas of exposed alluvium. This class can succeed to a low shrub state (class B) or directly to a dwarf shrub state (class C). Deterministic succession to class B. The alternate succession pathway to C (probability = 0.03) represents sites that would not support low shrubs. Flooding (probability = 0.08), represented by Option 1 in the model, resets *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 172 of 204 the age of this class. Class B Structure Data (for upper layer lifeform) Min Max 20 % Late Development 1 All Structures Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position SAAL SALIX ALVIS BENA Shrub Tree Tall Shrub (>1.5 m) Low Shrub (20 cm to 1.5 m) Tree Size Class Upper Upper Upper Upper None Upper layer lifeform differs from dominant lifeform. Description 15yrs+ Willow and alder low shrubs. Common shrub species may include Salix alaxensis and other Salix spp., Alnus viridis ssp. sinuata and Betula nana. This class may persist on some sites or may eventually transition to dwarf shrub (class C). Alternate succession to class C occurs with a probability of 0.015. Flooding (probability = 0.03), represented by Option 1 in the model, causes a transition to class A. Class C Structure Data (for upper layer lifeform) Min Max 45 % Late Development 2 All Structures Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position SALIX SARE2 DRYAS EMNI Upper Upper Upper Upper Dwarf Shrub (< 20 cm) Tree Size Class Dwarf Shrub (< 20 cm) None Upper layer lifeform differs from dominant lifeform. Description 15yrs+ On higher elevation alpine flood plains, dwarf shrubs may replace the early seral herbaceous stage. Dominant shrubs may include one or more of the following : Salix spp., Salix reticulata, Dryas spp., or Empetrum nigrum. Low willows can still be present, but cover is <25%. This class persists in the absence of disturbance. Flooding (probability = 0.03), represented by Option 1 in the model, causes a transition to class A. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 173 of 204 Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Min Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals NA Avg FI Min FI Max FI Probability Percent of All Fires Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Flooding Other (optional 2) References NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 174 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316420 Aleutian Kenai Birch-Cottonwood-Poplar Forest This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Kori Blankenship Modeler 2 Keith Boggs Modeler 3 Reviewer Reviewer Reviewer Map Zone 73 Forest and Woodland Dominant Species* 9/23/2008 kblankenship@tnc.org ankwb@uaa.alaska.edu Vegetation Type BEPAK BEPA POBA2 POBAT Date General Model Sources ALVIS SABA3 RUSP SARA2 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This BpS is commonly found on the eastern Alaska Peninsula and on Kodiak Island. Biophysical Site Description This BpS occurs at low elevations and also at the upper elevational limit of broad-leaved trees. At low elevations it is found predominantly on well-drained, gentle lower hillslopes, large moraines and old riparian terraces, although floodplain stands of cottonwood are not included in this system. In Katmai National Park and Preserve it occurs between 19-286m elevation mostly on slopes and valley bottoms (Boggs et al. 2003). When found on slopes this type is generally associated with shallow, stony soils that overlay bedrock (Viereck et al. 1992; I.B.2.c). Vegetation Description Total hardwood tree species cover is >25% dominated by Betula papyrifera var. kenaica, Betula papyrifera, Populus balsamifera ssp. trichocarpa or Populus balsamifera. Tree height ranges from 6-21m. Understory shrubs include Alnus viridis ssp. sinuata, Salix barclayi, Rubus spectabilis and Sambucus racemosa. Herbaceous species may also dominate the understory, such as Athyrium filix-femina, Calamagrostis canadensis, Calamagrostis lapponica, Chamerion angustifolium ssp. angustifolium, Equisetum spp., Gymnocarpium dryopteris and Heracleum maximum. Disturbance Description This system’s successional status is unclear. Viereck et al. (1992) suggest that some overmature Southcentral Alaska Open Paper Birch Forest (I.B.2.a) may be “reverting to open shrub communities with grassy openings (Neilenad and Viereck 1977 as cited in Viereck et al. 1992)." Also, the Open Balsam Poplar (Black Cottonwood) Forest (I.B.2.c) type identified by Viereck et al. (1992) appears to be fairly stable near tree-line. Possible disturbances include occasional fire, wind, disease and avalanches on steep *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 175 of 204 slopes. Adjacency or Identification Concerns Native Uncharacteristic Conditions This type is not departed from its “Reference Condition.” Scale Description Small to large patch size. Issues/Problems Comments This system was created for the Alaska Aleutians region and did not receive review for other regions in the state. This model was created by Kori Blankenship and Keith Boggs based on information from the draft Aleutians Ecological Systems description (NatureServe 2008). Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 100 % Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position BEPAK BEPA POBA2 POBAT Upper Upper Upper Upper Open (25-59% tree cover) Height Tree Size Class Closed (60-100% tree cover) Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Tree (> 3 m) Upper layer lifeform differs from dominant lifeform. Description Zero years plus This class represents the Aleutian Kenai Birch-Cottonwood-Poplar Forest. See Vegetation Description for more information on species composition. No disturbances modeled. Class B Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Cover Indicator Species* and Canopy Position Shrub Tree Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 176 of 204 Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Min Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: NA Historical Fire Size (acres) Avg 0 Min 0 Max 0 Sources of Fire Regime Data Literature Local Data Expert Estimate Fire Intervals Avg FI Min FI Max FI Probability Percent of All Fires Replacement Mixed Surface All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 177 of 204 Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Boggs, K., S.C. Klein, J. Grunblatt and B. Koltun. 2003. Landcover classes, ecoregions and plant associations of Katmai National Park and Preserve. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage, 707 A Street, Anchorage, AK 99501. 274 pp. Fleming, M.D., and P. Spencer. 2007. Kodiak Archipelago land cover classification users guide. SAIC at USGS Alaska Science Center, Anchorage, AK. 77 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for the Alaska Aleutians Region. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 178 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316430 Alaskan Pacific Maritime Alpine Herbaceous Dwarf Shrubland This BPS is lumped with: see below This BPS is split into multiple models: The Alpine Herbaceous Dwarf Shrubland BpS represents the following Ecological Systems: Alaska Sub-boreal and Maritime Alpine Mesic Herbaceous Meadow, Maritime Alpine Sparse Dwarf-Shrub, Maritime High Alpine Herbaceous. These types may be distinct existing vegetation communities but little is known about their successional relationships. Furthermore, these types may be difficult to map separately because they occur either adjacent or in a mosaic with one another and several of them are small patch systems. For these reasons, these types are modeled as one BpS. General Information Contributors (also see the Comments field Modeler 1 Tom DeMeo Modeler 2 Modeler 3 tdemeo@fs.fed.us Upland Shrubland Dominant Species* 7/30/2008 Reviewer Paul Hennon Reviewer Reviewer Map Zone 73 Vegetation Type EMNI PHAL4 HAST3 ARAR9 Date General Model Sources CAMA11 LUNO VASI GEER2 Literature Local Data Expert Estimate phennon@fs.fed.us Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This BpS is found from Prince William Sound south through southeast AK. Biophysical Site Description Alpine herb and shrub systems occur on a variety of sites in the subalpine and alpine zones. Much of the alpine zone will be under snow for most of the year leading to a short growing season. Environmental conditions are often harsh, particularly on exposed summits and ridge tops where vegetation may be sparse with a high proportion of exposed rock or soil. Vegetation Description This BpS represents several existing vegetation types so species composition is highly variable. Shrub species may include: Artemisia arctica, Cassiope mertensiana, Empetrum nigrum, Harrimanella stelleriana, Luetkea pectinata, Loiseleuria procumbens, Phyllodoce aleutica, P. glanduliflora, Salix arctica, S. reticulata, S. rotundifolia, Sibbaldia procumbens, Vaccinium uliginosum and Vaccinium vitisidaea. Herbceous species may include: Aconitum delphiniifolium, Anemone narcissiflora, Astragalus alpinus, *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 179 of 204 Athyrium filix-femina, Carex macrochaeta, Castilleja unalaschcensis, Chamerion angustifolium, Chamerion latifolium, Calamagrostis canadensis, Geranium erianthum, Lupinus nootkatensis, Minuartia arctica, Nephrophyllidium crista-galli, Saxifraga bracteata, Saxifraga bronchialis, Silene acaulis, Sanguisorba canadensis, Senecio triangularis, Valeriana sitchensis, Veratrum viride and Viola spp. Disturbance Description These communities tend to be quite stable over time. Soil disturbance, such as soil creep or freeze-thaw cycles, is likely the main disturbance factor although snow avalanche is possible in some areas. Wind plays a role in inhibiting vegetation growth on exposed summits and ridge tops. Reduced snow levels could allow invasion of other plants and lead to succession to another system. This type is unlikely to carry fire. Adjacency or Identification Concerns Native Uncharacteristic Conditions Scale Description Issues/Problems It is unclear whether there is a successional relationship between herbs and shrubs in some alpine types or whether soil differences result in the prevalence of different life forms on different sites. Because of the uncertainty, this system is represented by a single seral stage. Comments This system was created for the Alaska Maritime region and did not receive review for other regions in the state. This model was based on the draft Maritime Ecological Systems description (NatureServe 2008) with input by Tom DeMeo. Review comments resulted in minor additions to the description. Vegetation Classes Class A 100 % Mid Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position EMNI PHAL4 CAMA11 LUNO Upper Upper Upper Upper Structure Data (for upper layer lifeform) Min Max Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Height Dwarf Shrub (< 20 cm) Low Shrub (20 cm to 1.5 m) Tree Size Class None Upper layer lifeform differs from dominant lifeform. Herbaceous vegetation may also be dominant. Description Zero years plus Alpine Herbaceous-Dwarf Shrub communities may include a diverse mix of species and species assemblages. The Class Indicator Species listed are not true indicators -- refer to the Vegetation Description for species information. No disturbances modeled. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 180 of 204 Class B Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Cover Indicator Species* and Canopy Position Height Tree Size Class Shrub Tree Upper layer lifeform differs from dominant lifeform. Description Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Min Indicator Species* and Canopy Position Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Shrub Tree Max Cover Indicator Species* and Canopy Position Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 181 of 204 Fire Regime Group**: Fire Intervals NA Avg FI Min FI Max FI Probability Percent of All Fires Replacement Historical Fire Size (acres) Mixed Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 182 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316440 Alaskan Pacific Maritime Sitka Spruce Forest This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Tina Boucher Modeler 2 Kori Blankenship Modeler 3 antvb@uaa.alaska.edu kblankenship@tnc.org Reviewer Reviewer Map Zone 73 Forest and Woodland PISI OPHO VAOV SARA2 11/5/2008 Reviewer Vegetation Type Dominant Species* Date General Model Sources ALVIS RUSP HYSP70 RHLO70 Literature Local Data Expert Estimate Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system is found from south of Blying Sound on the Kenai Peninsula to Shuyak and Afognak Islands and the northern tip of Kodiak Island from Viekoda Bay southeast to Narrow Cape. It is also present in a narrow coastal band (likely one to several km wide) on the west side of Cook Inlet including portions of Katmai and Lake Clark National Parks. The northern extent of Sitka spruce is near the Johnson River. Biophysical Site Description This BpS occurs in an area where the boreal and maritime regions of AK overlap. Sitka spruce occurs on rolling bedrock hills and valleys where there is a strong maritime influence and warm, wet coastal storms occur. Soils tend to be well-drained, deep and well-developed (except in areas where spruce is invading meadows and shrublands). The biophysical environment of the Kodiak Archipelago is unique in that vegetation communities on these islands are still responding to the retreat of Pleistocene glaciers and ash deposition from the 1912 Katmai eruption (Fleming and Spencer 2007). Fleming and Spencer (2007, p. 5) note that: “Sitka spruce forest is actively moving south along the island of Kodiak, perhaps as much as a mile/century [the source of this claim is unknown (personal communication, Page Spencer)]. This succession replaces alder, salmon and elderberry and forb meadows with dense spruce forest. Preferred wildlife habitats with grasses, sedges and forbs and prolific berry crops are being replaced with dense conifers.” Vegetation Description Picea sitchensis is the dominant tree species and is stunted in some areas (especially on Shuyak and Afognak Islands). Oplopanax horridus and mosses are common in the understory of open stands (Fleming and Spencer 2007). Closed stands tend to have a very sparse understory that may include Oplopanax horridus, Vaccinium ovalifolium, Sambucus racemosa, Alnus crispa ssp. sinuata, Rubus spectabilis and *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 183 of 204 feather mosses (e.g. Hylocomium splendens and Rhytidiadelphus loreus) (Fleming and Spencer 2007). Calamagrostis canadensis and Menyanthes trifoliata are also present in some forest openings. In Lake Clark National Park near the northern extent of this BpS, a broader range of species is present including Picea glauca which may hybridize with Picea sitchensis. In this area, the characteristic moss understory found in the Kodiak Archipelago is absent. Disturbance Description Primary succession of Picea sitchensis forest in the Kodiak Archipelago begins with rich forb meadows. Overtime, Alnus crispa ssp. sinuata invades, fills in and Sambucus racemosa and Rubus spectabilis are added to the species mix. Eventually Picea sitchensis will start to invade. The timing of the transition to Picea sitchensis appears to be a function of the timing of the Pleistocene glacial retreat, seed source and seed advancement. As the Picea sitchensis trees become denser, the first generation closed Picea sitchensis forest develops with Vaccinium ovalifolium, Oplopanax horridus and feather mosses in the understory. After about 150-300yrs the trees start to senesce, small scale blow down occurs and a multi-age stand develops. In the forest gaps left after tree death or blowdown a rich shrub layer with Alnus crispa ssp. sinuata, Oplopanax horridus and Calamagrostis canadensis competes with the young Picea sitchensis trees. Over long time scales, these forests become more open. In areas with beaver activity, ponds can form that when drained turn into muskegs which dry and can again support Picea sitchensis trees. In the Katmai area Picea sitchensis advancement appears to be slower than on Kodiak (personal communication, Page Spencer). It is unclear how well the Kodiak successional sequence applies to Picea sitchensis forest on the south end of the Kenai Peninsula which appears to be better developed and is the likely seed source for Picea sitchensis invading the Kodiak Archipelago. Small scale wind events are the main disturbance affecting this system (larger areas of blowdown are possible after logging). Bark beetle attacks have been recorded but their effects are minor and a few human caused fires have been reported. Gap-phase dynamics drive the successional processes. As older trees senesce or are windthrown gaps are created for younger trees to exploit and a multi-age stand develops. Adjacency or Identification Concerns Open Sitka spruce forests are often intermixed with alder and forb meadows (Fleming and Spencer 2007). Native Uncharacteristic Conditions Logging on Kodiak Island and on the Kenai Peninsula south of Seldovia has created more early successional forests than would exist otherwise. Scale Description Matrix Issues/Problems This BpS is represented by a one-box model for several reasons: 1) Although Sitka spruce sites represent a unique biophysical environment with unique successional dynamics it is difficult to model them using the LANDFIRE methodology which assumes that overtime an equilibrium condition will be reached and that the percent of the landscape in each seral stage varies within a certain range (i.e. the natural or historic range of variability). These modeling assumptions do not apply to areas where spruce is actively invading. 2) In dense spruce forest where secondary succession characterized by gap dynamics has begun, the *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 184 of 204 disturbance is occurring on a fine scale as individual or small groups of trees die or are windthrown. The small forest openings created by these dynamics are likely not mappable using the LANDFIRE methodology and therefore seral stages can not be defined for me (for the LANDFIRE project seral stages must be mappable). 3) It is unclear how well the dynamics described in the Disturbance Description for the Kodiak Archipelago apply to the Kenai and west Cook Inlet parts of the distribution of this BpS. Comments This model description is largely based on work in the Kodiak Archipelago (Fleming and Spencer 2007) and conversations with Page Spencer. Vegetation Classes Class A 100 % Mid Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position PISI OPHO VAOV SARA2 Upper Lower Lower Lower Cover Height Structure Data (for upper layer lifeform) Min Max Woodland (10-24% tree cover) Closed (60-100% tree cover) Dwarf Tree (< 3 m) Tree Size Class Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Shrubs may dominate the early seral stages of this BpS. Description Zero years plus This class represents the Sitka Spruce Forest BpS. See the vegetation description for common species. No disturbances were modeled. Class B Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Cover Indicator Species* and Canopy Position Shrub Tree Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 185 of 204 Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Min Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: NA Historical Fire Size (acres) Avg 0 Min 0 Max 0 Sources of Fire Regime Data Literature Local Data Expert Estimate Fire Intervals Avg FI Min FI Max FI Probability Percent of All Fires Replacement Mixed Surface All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 186 of 204 Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Boggs et al. in preparation. Kenai Fjords User’s Guide. Fleming, M.D. and P. Spencer. 2007. Kodiak Archipelago land cover classification users guide. SAIC at USGS Alaska Science Center, Anchorage, AK. 77 pp. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 187 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316450 Alaska Sub-boreal and Maritime Alpine Mesic Herbaceous Meadow This BPS is lumped with: This BPS is split into multiple models: General Information Contributors (also see the Comments field Modeler 1 Tina Boucher antvb@uaa.alaska.edu colleen_ryan@tnc.org Modeler 2 Colleen Ryan Modeler 3 Upland Grassland/Herbaceous Dominant Species* General Model Sources LUNO VEVI ACDE2 ANNA 4/24/2008 Reviewer Reviewer Reviewer Vegetation Type CAMA11 GEER2 SACA14 VASI Date Literature Local Data Expert Estimate Map Zone 73 Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range Mountains in the sub-boreal region of AK. Biophysical Site Description This system occurs on alpine slopes above the tall shrub zone and below alpine dwarf shrub tundra (NatureServe 2008). In the Kenai Mountains, it occurs on rugged mountains, dissected mountain side slopes and ravines, on 45-55% slopes of all aspects and at elevations from 1950-2950ft (DeVelice et al. 1999). This system usually occurs on sites just above subalpine alder and below alpine dwarf shrub, most commonly on straight or concave slopes (NatureServe 2008). Vegetation Description This system is a very diverse assemblage of sedges and forbs. Carex macrochaeta and Geranium erianthum are the most typical species. Other common species include Sanguisorba stipulata, Valeriana sitchensis, Lupinus nootkatensis, Veratrum viride, Aconitum delphiniifolium, Anemone narcissiflora, Polemonium acutiflorum, Epilobium angustifolium, Castilleja unalaschensis, Artemesia arctica, Fritillaria Camschatcensis, Erigeron peregrinus, Anemone spp., Pedicularis spp., Saxifraga spp., Polygonum spp. and ferns (DeVelice et al. 1999, Viereck et al 1992). Calamagrostis canadensis may be present, but is not dominant. Disturbance Description Fire is unlikely to be a significant disturbance for this system, due to moist soils, low fuel levels, and adjacent vegetation types that rarely burn. Though this type may be found adjacent to the Western North American Sub-boreal Mesic Bluejoint Meadow, it is probably less flammable. Avalanches and snow slides are likely the most important disturbance, especially on steeper slopes. Spring *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 188 of 204 snow slides are the most likely to impact vegetation. While this system often occurs on steep slopes with high avalanche frequency, it can also occur on shoulder slopes and near summits, where snowslides are rare. For this model, the mean return interval of snowslides was estimated at 20yrs (five times the return interval of snowslides in the Alaska Sub-boreal Avalanche Slope Shrubland). Grazing may be an influence on this system in some areas. Adjacency or Identification Concerns Adjacent systems include Western North American Sub-boreal Mesic Bluejoint Meadow and Western North American Boreal Alpine Dwarf-Shrubland. Native Uncharacteristic Conditions Scale Description Large patch Issues/Problems Comments Vegetation Classes Class A Structure Data (for upper layer lifeform) Min Max 100 % Early Development 1 All Structures Cover Upper Layer Lifeform Height Herbaceous Tree Size Class None Herbaceous Shrub Tree Indicator Species* and Canopy Position CAMA11 GEER2 SACA14 VASI Upper Upper Upper Upper Herbaceous Herbaceous Herbaceous Upper layer lifeform differs from dominant lifeform. Description Zero years plus This stage is dominated by Carex macrochaeta with a variety of forbs. This class persists in the absence of disturbance. Probability of avalanche = 0.05 (20yr return interval). Class B Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Herbaceous Cover Indicator Species* and Canopy Position Shrub Tree Height Tree Size Class Upper layer lifeform differs from dominant lifeform. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 189 of 204 Description Class C Structure Data (for upper layer lifeform) Min Max 0% [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Min Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: NA Historical Fire Size (acres) Avg 0 Min 0 Max 0 Sources of Fire Regime Data Literature Local Data Expert Estimate Fire Intervals Avg FI Min FI Max FI Probability Percent of All Fires Replacement Mixed Surface All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 190 of 204 Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) snow slides Other (optional 2) References Boggs, K., A. Garibaldi, J. Stevens, J. Grunblatt and T. Helt. 2001. Denali National Park and Preserve Landcover mapping project. Volume 2: Landcover classes and plant associations. Alaska Natural Heritage Program, Environment and Natural Resources Institute, University of Alaska Anchorage, 707 A Street, Anchorage, AK. 164 pp. DeVelice, R.L., Hubbard, C.J., Boggs, K. et al. 1999. Plant community types of the Chugach National Forest. Tech. Publ. R10-TP-76. Juneau, AK: USDA Forest Service, Alaska Region. 375 pp. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Viereck et al. 1992. The Alaska vegetation classification. Pacific Northwest Research Station, USDA Forest Service, Portland, OR. Gen. Tech. Rep. PNW-GTR286. 278 pp. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 191 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316481 Alaskan Pacific Maritime Mountain Hemlock Forest - Northern This BPS is lumped with: see below This BPS is split into multiple models: Alaskan Pacific Maritime Subalpine Mountain Hemlock Woodland (1649) is lumped with Alaskan Pacific Maritime Mountain Hemlock Forest (1648). Alaskan Pacific Maritime Mountain Hemlock Forest (1648) was then split into a northern (16481) which includes fire disturbance and a southeast (16482) variant which does not. General Information Contributors (also see the Comments field Modeler 1 John Morton Modeler 2 Edward Berg Modeler 3 Tina Boucher Map Zone 73 Forest and Woodland TSME PISI PILU VAOV 11/5/2008 John_M_Morton@fws.g Reviewer Tom DeMeo ov Edward_Berg@fws.gov Reviewer Barbara Schrader Reviewer antvb@uaa.alaska.edu Vegetation Type Dominant Species* Date General Model Sources ALVIS MEFE ELPY NECR2 Literature Local Data Expert Estimate tdemeo@fs.fed.us bschrader@fs.fed.us Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range This system occurs in the Kenai, Chugach, and St. Elias mountains in the subpolar rainforest region (Alaback 1991, 1995). It occurs primarily in the maritime region, but also occurs in the sub-boreal transition between coastal and boreal forests. It extends along the Gulf Coast of AK from Kenai Fjords to Yakutat and possibly Glacier Bay and upper Lynn Canal. (North Pacific Mountain Hemlock Forest is defined as occurring from Glacier Bay south). Biophysical Site Description Within the Kenai, Chugach, and St. Elias Mountains this system occurs on mountain side slopes, shoulders, and bedrock outcrops from 0-1900ft (NatureServe 2008a). On the Chugach National Forest mountain hemlock-sitka spruce associations are found at elevations up to 1200ft and mountain hemlock associations are found several hundred feet higher (DeVelice 1999). Soils are typically shallow, welldrained and are derived from glacial and colluvial deposits as well as residual bedrock. Alaska Sub-boreal Mountain Hemlock Forest is common on north-facing slopes, and also occurs on east and west aspects. This system is uncommon on south-facing slopes (NatureServe 2008a). Vegetation Description Tsuga mertensiana may dominate or share dominance with Picea sitchensis or Tsuga heterophylla on sites along the central Gulf Coast (Prince William Sound). Cupressus nootkatensis may be present in the *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 192 of 204 overstory in isolated locations in Prince William Sound. Picea glauca or Picea X Lutzii, may be present on the inland (non-maritime) portion of the Kenai and Chugach Mountains, but have <15% canopy cover. Betula papyrifera may be common in early seral stages in inland sites.. Common shrubs include Menziesia ferruginea, Alnus viridis ssp. sinuata, Vaccinium ovalifolium, V. vitisidaea, Oplopanax horridus, Cassiope stellariana, Rubus spectabilis, Elliotia pyroliflora and Empetrum nigrum (NatureServe 2008a, NatureServe 2008b). Common herbaceous species include Cornus canadensis, Rubus pedatus, Dryopteris expansa, Gymnocarpium dryopteris, Listera cordata, Tiarella trifoliata and Streptopus amplexifolius (NatureServe 2008a, NatureServe 2008b). Common mosses include Hylocomium splendens and Pleurozium schreberi (NatureServe 2008). Sphagnum spp. may be abundant on some sites. Plant communities in this system are described by DeVelice et al. (1999). Picea sitchensis, Elliottia pyroliflorus and Nephrophyllidium crista-galli do not occur in the sub-boreal portion of the region. Disturbance Description The major disturbance processes affecting this system include soil creep, wind, snow avalanche and fungal pathogens such as red ring rot (Phellinus pini). Windthrow gap disturbances are important in both spruce and hemlock recruitment in these forests (Potkin 1997). A moderate level of disturbance probably maintains Picea sitchensis in the system. Although lightning strikes and natural fires are rare in the maritime region, wildfires can play an important role in disturbance regime of the Mountain Hemlock PNV (Potkin 1997) and other similar temperate rainforest types (Agee 1993, Franklin and Hemstrom 1981) in areas where lightning strikes do occur, such as the inland side of the Kenai and Chugach mountains and upper Lynn Canal. Radiocarbon dates of five charcoal samples from soils at various locations in the Kenai Mountains ranged from 3010 to 570yrs before present with an average of 600yrs between dates (Potkin 1997). Charcoal has been reported as present in most soil pits within the forest zone in the Kenai Mountains; this anecdotal evidence suggests the occurrence of widespread, infrequent fires in this BpS (USDA Forest Service 2002). Mountain hemlock, the climax species, is a fire “avoider”; the infrequent fires tend to be large and standreplacing (Agee 1993). In the Kenai Mountains fires travel from the valley bottom Lutz spruce stands, but often stop at the lower boundary of mountain hemlock dominated forests (Potkin 1997). Estimates of fire return intervals include: 570-3010yrs (600yr average) (Potkin 1997) (for Kenai Mountains); 1000yrs (personal communication, FRCC expert’s workshop, March 2004); 1500yrs+ (Lertzman and Krebs 1991) (Cascades and Olympic Mountains). The infrequent fires in the subalpine hemlock forests tend to be large and stand-replacing (Agee 1993). The extent of subalpine fires is partly a function of fire weather, but mainly depends on the distribution of the subalpine forests, which is often patchy and grades into shrub, tundra, rock and ice (Agee 1992). In the mountain hemlock zone in Washington State, fires have tended to be confined to individual slopes (Agee and Smith 1984). In Oregon, historic fires >3200ha have been located; however, the majority of stands found were patches of <500ha each (Dickman and Cook 1989). On the Kenai Peninsula, drought weather conditions resulting from the 1912 Katmai eruption have been suggested to contribute to large scale fires from 1913-1915, burning approximately 20,000ac on the Chugach National Forest (note that this figure includes fire in all vegetation types present on the landscape). There is almost no lightning and little human activity in the Prince William Sound area suggesting that fire is not an important factor in *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 193 of 204 this system (personal communication, Sue Kesti) for the entire area to which it applies. Adjacency or Identification Concerns The Martime and Sub-boreal Mountain Hemlock system typically occurs from sea level to treeline in coastal environments; adjacent types include Maritime Sitka Spruce and Martime Western Hemlock. In the sub-boreal region, it typically occurs in the elevation zone above the Sub-boreal White SpruceMountain Hemlock system. Native Uncharacteristic Conditions As a result of climate warming, white spruce are now recruiting above mountain hemlock in the subboreal region of the Kenai Peninsula. Scale Description Matrix Issues/Problems Vegetation communities in the mountain hemlock BpS are stable over long periods and rarely disturbed; therefore, secondary succession patterns are poorly understood (Viereck et al 1992). Fire is included in the model but may not occur (or is extremely rare) in some areas to which this model applies. Comments This model was based on the FRCC Guidebook PNVG model for Kenai Mountain Hemlock (KMHM; Murphy and Witten 2006) but the mid and late seral stages had to be collapsed so that LANDFIRE could distinguish them for mapping. Ed Berg and John Morton helped to refine the model for the Kenai Peninsula and Tina Boucher modified the description to include the rest of the range. Much of the Disturbance Description was taken from the Kenai Mountain Hemlock (KMHM; Murphy and Witten 2006) description with some modifications. Reviewers suggested that this northern Mt. Hemlock type could probably be combined with the southern Mt. Hemlock type (North Pacific Mountain Hemlock Forest). This advice was followed for the Ecological Systems classification but not for modeling purposes because the northern variant of the model includes fire disturbance but the southern variant does not. Vegetation Classes Class A 5% Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position VAOV MEFE RUSP Upper Upper Upper Structure Data (for upper layer lifeform) Min Max Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Dwarf Shrub (< 20 cm) Tall Shrub (>1.5 m) Height Tree Size Class Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Description 0-49yrs This post-disturbance class is characterized by mesic herbaceous vegetation and tall shrubs. Herbaceous species may start from seed immediately post-disturbance. Shrubs and tree seedlings become established; after *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 194 of 204 approximately 50yrs tree saplings attain the height of tall shrubs. Common shrubs include Menziesia ferruginea, Alnus viridis ssp. sinuata, Vaccinium ovalifolium and Oplopanax horridus (NatureServe 2008a, NatureServe 2008b). Succession to class B. Alternate succession (probability = 0.005) causes a transition to class C. Class B Structure Data (for upper layer lifeform) Min Max 60 % Mid Development 1 Closed Upper Layer Lifeform Herbaceous TSME PILU MEFE ALVIS Shrub Tree Cover Indicator Species* and Canopy Position Closed (60-100% tree cover) Tree (> 3 m) Height Tree Size Class Upper Upper Lower Lower Closed (60-100% tree cover) Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Description 50yrs+ This class is characterized by closed spruce-hemlock or hemlock forest. Conifers share dominance with tall shrubs in the early stages of this class. Between 170-200yrs conifers gain canopy dominance. Tsuga mertensiana is the dominant tree with at least 15% cover. This class persists in the absence of disturbance. Replacement MFRI = 1250yrs. Mixed fire (MFRI = 5000yrs) causes a transition to class C. Wind/weather/stress (probability = 0.0002) maintains this class. Class C 35 % Mid Development 1 Open Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position TSME PILU MEFE ALVIS Upper Upper Lower Lower Cover Height Structure Data (for upper layer lifeform) Min Max Open (25-59% tree cover) Open (25-59% tree cover) Tree Size Class Tree (> 3 m) Tree (> 3 m) Med. 9–20" (swd)/11–20" (hwd) Upper layer lifeform differs from dominant lifeform. Description 50yrs+ This class is characterized by open spruce-hemlock or hemlock forest. Conifers share dominance with tall shrubs in the early stages of this class. Between 170-200yrs conifers gain canopy dominance. Tsuga mertensiana is the dominant tree with at least 15% cover. This class persists in the absence of disturbance. Replacement MFRI = 1250yrs. Mixed fire (MFRI = 5000yrs) maintains this class. Wind/weather/stress (probability = 0.0002) maintains this class. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 195 of 204 Structure Data (for upper layer lifeform) Class D 0 % [Not Used] [Not Used] Upper Layer Lifeform Min Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Class E Structure Data (for upper layer lifeform) 0% Min [Not Used] [Not Used] Upper Layer Lifeform Indicator Species* and Canopy Position Herbaceous Shrub Tree Max Cover Height Tree Size Class Upper layer lifeform differs from dominant lifeform. Description Disturbances Fire Regime Group**: Fire Intervals V Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 1430 5000 0.0007 0.0002 1111 0.00091 Percent of All Fires 77 22 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Agee, J. K. 1993. Fire Ecology of Pacific Northwest Forests. Island Press publishing. 493 pp. Agee, J.K. and Smith, L. 1984. Subalpine tree establishment after fire in the Olympic Mountains, Washington. Ecology 65: 810-19. Alaback, P.B. 1991. Comparative ecology of temperate rainforests of the Americas along analogous climatic gradients. Rev. Chil. Hist. Nat. 64: 399–412. Alaback, P.B. 1995. Biodiversity patterns in relation to climate and a genetic base for the rainforests of the *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 196 of 204 west coast of North America. In R. Lawford, P.B. Alaback and E.R. Fuentes, eds. 1995. High Latitude Rain Forests of the West Coast of the Americas: Climate, Hydrology, Ecology and Conservation. Berlin: SpringerVerlag. DeVelice, R.L., Hubbard, C.J., Boggs, K. et al. 1999. Plant community types of the Chugach National Forest. Tech. Publ. R10-TP-76. Juneau, AK: USDA Forest Service, Alaska Region. 375 pp. Dickman, A. and Cook, S. 1989. Fire and fungus in mountain hemlock forest. Can. J. Bot. 67:2005-16. Franklin, J.F. and Hemstrom, M.A. 1981. Aspects of succession in the coniferous forests of the Pacific Northwest. In: West, D.C.; Shugart, H.H.; Botkin, D.B. Forest succession; concepts and application. Springer-Verlag, NewYork. Lertzman, K.P. and Krebs, C.J. 1991. Gap-phase structure of old-growth forest. Can. J. For. Res. 21: 173041. Murphy, K.A. and E. Witten. 2006. Kenai Mountain Hemlock. In Fire Regime Condition Class (FRCC) Interagency Guidebook Reference Conditions. Available at www.frcc.gov. NatureServe. 2008a. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. NatureServe. 2008b. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for the Alaska Maritime Region. Potkin, M. 1997. Fire history disturbance study of the Kenai Peninsula mountainous portion of the Chugach National Forest. Draft. USDA Forest Service, Chugach National Forest. December 5, 1997. Anchorage, AK. USDA Forest Service. 2002. Revised land and resource management plan, final environmental impact statement. Alaska Region Chugach National Forest. R10-MB-480d, May 2002. Viereck, L.A. 1979. Characteristics of treeline plant communities in Alaska. Holarctic Ecology. 2: 228-238. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 197 of 204 LANDFIRE Biophysical Setting Model Biophysical Setting 7316790 Alaska Sub-boreal White Spruce-Hardwood Forest This BPS is lumped with: see below This BPS is split into multiple models: Alaska Sub-boreal White-Lutz Spruce Forest & Woodland and Western North American Boreal Mesic Birch-Aspen Forest (when it occurs in the AK sub-boreal region) are lumped with Alaska Sub-boreal White Spruce-Hardwood Forest because it is not clear that these types occur on different biophysical settings. Although these types are mappable as separate existing vegetation types they should be mapped as one BpS. General Information Contributors (also see the Comments field Modeler 1 Karen Murphy Modeler 2 Evie Witten Modeler 3 Kori Blankenship Map Zone 73 Forest and Woodland PIGL BEPA POBA2 POTR5 2/10/2008 karen_a_murphy@fws.g Reviewer Rob DeVelice ov Reviewer ewitten@tnc.org Reviewer kblankenship@tnc.org Vegetation Type Dominant Species* Date General Model Sources MEFE ALVIS CACA4 OPHO Literature Local Data Expert Estimate rdevelice@fs.fed.us Model Zone Alaska California Great Basin Great Lakes Northeast Northern Plains N-Cent.Rockies Pacific Northwest South Central Southeast S. Appalachians Southwest Geographic Range These systems occur throughout the sub-boreal region of AK. The white spruce-hardwood forest type occurs primarily on the west side of the Kenai Peninsula on the Kenai Lowland and also at lower elevations in the northwestern Kenai Mountains. Biophysical Site Description These systems typically occur on well-drained upland terrain, including side slopes, toe slopes and inactive terraces. Soils are generally developed on surficial deposits including glacial till, colluvium, and loess. Within the Kenai Mountains, the Sub-boreal White Spruce-Hardwood system occurs at a lower elevation zone than mountain hemlock-white spruce and mountain hemlock systems. Vegetation Description The dominant canopy species are Picea glauca or Picea X lutzii (the hybrid produced where the ranges of P. sitchensis and P. glauca overlap) and Betula papyrifera. Other common canopy trees include Populus balsamifera and P. tremuloides. Picea mariana may also be present. Salix scouleriana is locally common. Common understory shrubs include Alnus viridis ssp. Sinuata, Viburnum edule, Rosa acicularis, Ribes triste, Vaccinium vitis-idaea and Linnaea borealis. Menziesia ferruginea, Vaccinium ovalifolium and *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 198 of 204 Oplopanax horridus may also be common, especially in the spruce-dominated type. Common herbaceous species include Calamagrostis canadensis, Chamerion angustifolium, Gymnocarpium dryopteris, Cornus canadensis and, especially in the spruce-dominated type, Equisetum arvense and Dryopteris expansa. Common mosses include Hylocomium splendens and Pleurozium schreberi (DeVelice et al. 1999). Disturbance Description The major disturbance processes are fire, human disturbance, blowdown, insect infestations and (in the Kenai Mountains) snow avalanches. Although lightning and natural fires have historically been infrequent, wildfire plays an important role in the disturbance regime of this system (NatureServe 2008). Under the natural fire regime, fires were infrequent, but large (USDA Forest Service 2002). Estimates of the mean fire return interval range from 600-800yrs (Berg and Anderson 2006, Berg et al. 2004, Potkin 1997, personal communication FRCC experts’ workshop March 2004). Spruce bark beetle (Dendroctonus rufipennis) infestations are a major natural disturbance of sub-boreal spruce and spruce-hardwood forests. Spruce beetles typically attack larger, slow-growing spruce, but infestations periodically escalate to epidemic levels when forest and climatic conditions are favorable for beetle expansion (NatureServe 2008). During epidemic-level infestations, beetles are less selective and may attack and kill a wider range of spruce trees. Beetle outbreaks that thin stands and produce a growth release in surviving trees occur on average every 50yrs in white and Lutz spruce forests on the Kenai Peninsula (Berg et al 2006). Spruce bark beetle outbreaks that produce a more substantial thinning occur at longer intervals, with the last two severe infestations occurring in the 1870s-1880s and 1987 –present (Berg et al 2006). The bark beetle outbreak that began in 1987 on the Kenai Peninsula has killed over 1.3 million acres of spruce (USDA Forest Service 2002). Berg (2004) found no association between spruce bark beetle mortality and fire in the past. When the canopy of these forests is thinned by heavy spruce bark beetle-mortality, bluejoint grass (Calamagrostis canadensis) and fireweed (Epilobium angustifolium) tend to increase rapidly and dominate the site for at least 10yrs (Holsten et al. 1995). Calamagrostis may proliferate rapidly from its predisturbance low level network of rhizomatous roots and may develop into a thick, seedling-excluding sod within a few years (Berg et al 2006). Boucher (2003) found that rapid spread of Calamagrostis occurs primarily on sites with deep, loamy soils. Thinning of the spruce canopy by beetle attacks also helps to maintain hardwoods in the canopy over time. Other natural disturbances include wind, avalanche and landslides. Windthrow gap disturbances are important in both spruce and hemlock recruitment in these forests (Potkin 1997). Post-fire regeneration of white spruce appears to be more successful when fires occur in mast years (Peters et al. 2005). This interaction between fire, masting and subsequent tree regeneration could have implications for historical stand structure and sucessional dynamics over time (Peters et al. 2005). Adjacency or Identification Concerns Native Uncharacteristic Conditions Adapted from Murphy and Witten (2006): The present landscape of the western Kenai Peninsula reflects human-caused fires that occurred over the last 100yrs, creating areas of early successional plant communities, which include large stands of broadleaved forests (Potkin 1997). Over 99% of the fires occurring on the Kenai Peninsula portion of the Chugach National Forest between 1914 and 1997 were *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 199 of 204 ignited by human actions (Potkin 1997). These human-caused fires have generally increased the richness and patchiness of the vegetation at the landscape scale (USDA Forest Service 2002). The large number of acres burned on the Kenai Peninsula during settlement caused conversion of some mature spruce stands to grass, brush and broadleaf tree vegetation types. Prior to the settlement period of the late 1800s, the majority of the age structures of the coniferous forest surveyed by Potkin (1997) were likely in the late successional stages (Langille 1904 in Potkin 1997) and conifers were likely dominant. Following spruce bark beetle outbreaks on the Kenai Peninsula grass and other fine vegetation increased (Holsten et al 1995). Fire spreads rapidly through this type of vegetation; indeed the majority of fires (most of which were human caused) on the Kenai Peninsula portion of the Chugach National Forest between 1914 and 1997 occurred in grassy vegetation (Potkin 1997). Standing and downed beetle killed trees increase the amount of both fine, flashy fuel and heavy fuel. Spruce bark beetle outbreaks may be increasing in frequency and severity in southcentral Alaska due to the warming climate. Berg et al. (2006) found that recent outbreaks on the Kenai peninsula were positively associated with average summer temperature in the preceding years. If this is true, current patterns of beetle attack and current fire regimes are likely atypical of reference conditions for this type. Scale Description Matrix or large patch. Issues/Problems Comments This model was based on the FRCC Guidebook Potential Natural Vegetation Group model for Coastal Boreal Transition Forest (CBTF) (Murphy and Witten 2006). Disturbances with a probability of .0001 (i.e. 10,000yr return interval) were removed from the model because their effect is insignificant in a 1000yr simulation. Class ages were adjusted slightly to make them line up along the main successional pathway, and the relative age function was not used in any class except A to comply with LANDFIRE modeling rules. These changes did not change the fire return intervals or the percent of the landscape in each class. Because only minor changes were made to the CBTF VDDT model, its original authors are included as contributors to this model, and Kori Blankenship's name was added. Much of the text in the General Information section of this description was taken from the draft Boreal Ecological Systems legend (NatureServe 2008). Vegetation Classes Class A 5% Early Development 1 All Structures Upper Layer Lifeform Herbaceous Shrub Tree Indicator Species* and Canopy Position CACA4 EQAR CHAN9 MEFE Upper Upper Upper Upper Structure Data (for upper layer lifeform) Min Max Cover Open Shrub (25-74% shrub cover) Closed Shrub (> 75% shrub cover) Height Dwarf Shrub (< 20 cm) Tall Shrub (>1.5 m) Tree Size Class Seedling/Sapling <5" Upper layer lifeform differs from dominant lifeform. Description 0-29yrs Post disturbance regeneration: herbaceous to tall shrub-sapling. Following a moderate severity burn, *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 200 of 204 vegetative reproduction of shrubs and birch and aspen from shoots and suckers. Light-seeded herbs establish where mineral soil is exposed. White and Lutz spruce seedlings are rare, but may be present if mineral soil was exposed, seed trees remained after fire and they produced a good seed crop (Foote 1983). Following severe fire on loamy soil, Calamagrostis may spread rapidly from rhizomes and capture a large percentage of the site (Boucher 2003). Mosses and lichens exist but are not an important component. If Calamagrostis captures the site, it may persist throughout this class but may become less dominant within 20yrs (Schulz 2000). Common shrubs include Menziesia ferruginea, Alnus viridis ssp. sinuata, Vaccinium ovalifolium, Oplopanax horridus, Vaccinium vitis-idaea and Linnaea borealis. Common herbaceous species include Calamagrostis canadensis, Equisetum arvense, Dryopteris expansa and Gymnocarpium dryopteris. Succession to class B. More open sites are represented by an alternate succession pathway to C (probability = 0.005) Class B Structure Data (for upper layer lifeform) Min Max 10 % Mid Development 1 Closed Cover Upper Layer Lifeform Height Herbaceous Indicator Species* and Canopy Position PIGL BEPA POBA2 POTR5 Shrub Tree Closed (60-100% tree cover) Closed (60-100% tree cover) Dwarf Tree (< 3 m) Tree Size Class Upper Upper Upper Upper Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping, presence of hardwoods distinguishes class B from class E. Description 30-129yrs Closed conifer, hardwood, or mixed. Tree saplings gain canopy dominance over shrubs. Tree species may include spruce, hardwoods or both. Rosa acicularis, Equisetum spp. and Linnaea borealis are commonly in the understory. Mosses and lichens become established. Succession to class E. Replacement MFRI = 1000yrs. Class C 15 % Mid Development 1 Open Cover Upper Layer Lifeform Height Herbaceous Shrub Tree Indicator Species* and Canopy Position PIGL BEPA POBA2 POTR5 Upper Upper Upper Upper Structure Data (for upper layer lifeform) Min Max Open (25-59% tree cover) Open (25-59% tree cover) Dwarf Tree (< 3 m) Tree Size Class Tree (> 3 m) Pole 5–9" (swd)/5–11" (hwd) Upper layer lifeform differs from dominant lifeform. For mapping, presence of hardwoods distinguishes class C from D. Description 30-129yrs Open conifer, hardwood or mixed. Young trees become dominant in the overstory. Calamagrostis, if *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 201 of 204 dominant in class A, diminishes in importance. Rosa acicularis, Equisetum spp. and Linnea borealis are commonly in the understory. Lichens and mosses become established. Succession to class D. Replacement MFRI = 1000yrs. Structure Data (for upper layer lifeform) Class D 65 % Late Development 1 Open Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position PIGL BEPA POBA2 POTR5 Upper Upper Upper Upper Min Woodland (10-24% tree cover) Height Tree Size Class Tree (> 3 m) Max Open (25-59% tree cover) Tree (> 3 m) Large 20" – 40" Upper layer lifeform differs from dominant lifeform. For mapping, absence of hardwoods distinguishes class D from C. Description 130-400yrs Open conifer. Open spruce, hardwood or mixed stands with tree density < 60%. Hardwoods, if present and mixed with spruce, lose dominance in overstory during this phase. Occasional hardwoods may remain. The understory may include various combinations of tall shrubs, low shrubs, herbs, mosses and lichens. This class persists in the absence of disturbance. Replacement MFRI = 625yrs. Mixed fire (MFRI = 3333yrs) causes a transition to C. Insects and disease can maintain this class (probability = 0.028) or cause a transition to C (probability = 0.0015) Class E Structure Data (for upper layer lifeform) 5% Min Late Development 1 Closed Upper Layer Lifeform Herbaceous Shrub Tree Cover Indicator Species* and Canopy Position PIGL BEPA POBA2 POTR5 Upper Upper Upper Upper Closed (60-100% tree cover) Tree (> 3 m) Height Tree Size Class Max Closed (60-100% tree cover) Tree (> 3 m) Large 20" – 40" Upper layer lifeform differs from dominant lifeform. For mapping, absence of hardwoods distinguishes class E from B. Description 130-400yrs Closed conifer, hardwood or mixed. Site is dominated by mature conifers with > 60% canopy closure. Hardwoods, if present and mixed with spruce, lose dominance in overstory during this phase. The understory may include various combinations of tall shrubs, low shrubs, herbs, mosses and lichens. This class persists in the absence of disturbance. Replacement MFRI = 606yrs. Mixed fire (MFRI = 2500yrs) causes a transition to D. Insects and disease can maintain this class (probability = 0.012), cause a transition to D (probability = 0.013) or cause a transition to C (probability = 0.0018). Disturbances *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 202 of 204 Fire Regime Group**: Fire Intervals V Replacement Historical Fire Size (acres) Mixed Avg FI Min FI Max FI Probability 715 5000 0.0014 0.0002 625 0.00161 Percent of All Fires 87 12 Surface Avg 0 Min 0 Max 0 All Fires Fire Intervals (FI): Fire interval is expressed in years for each fire severity class and for all types of fire combined (All Fires). Average FI is central tendency modeled. Minimum and maximum show the relative range of fire intervals, if known. Probability is the inverse of fire interval in years and is used in reference condition modeling. Percent of all fires is the percent of all fires in that severity class. Sources of Fire Regime Data Literature Local Data Expert Estimate Additional Disturbances Modeled Insects/Disease Wind/Weather/Stress Native Grazing Competition Other (optional 1) Other (optional 2) References Berg, E. 2004. White spruce fire history on the Kenai Peninsula, Alaska, based on radiocarbon-dated soil charcoal. Unpublished manuscript. Kenai National Wildlife Refuge, Alaska. Berg, E.E., J.D. Henry, C.L. Fastie, A.D. De Volder and S.M. Matsuoka. 2006. Spruce beetle outbreaks on the Kenai Peninsula, Alaska, and Kluane National Park and Reserve, Yukon Territory: Relationship to summer temperatures and regional differences in disturbance regimes. For. Ecol. Man. 227(3):219-232. Berg, E. and R.S. Anderson. 2006. Fire history of white and Lutz spruce forests on the Kenai Peninsula, Alaska, over the last two millennia as determined from soil charcoal. For. Ecol. Man 227: 275-283. Boucher, T.V. 2003. Vegetation response to prescribed fire in the Kenai Mountains, Alaska. Res. Pap. PNW-RP-554. Portland, OR: USDA Forest Service, Pacific Northwest Research Station. 59 pp. DeVelice, R.L., Hubbard, C.J., Boggs, K. et al. 1999. Plant community types of the Chugach National Forest. Tech. Publ. R10-TP-76. Juneau, AK: USDA Forest Service, Alaska Region. 375 pp. Foote, J. M. 1983. Classification, description, and dynamics of plant communities after fire in the taiga of interior Alaska. Res. Pap. PNW-307. Portland, OR. USDA Forest Service. Pacific Northwest Research Station. 108 pp. Holsten, E.H., R.A. Werner and R.L. DeVelice. 1995. Effects of spruce beetle (Coleoptera: Scolytidae) outbreak and fire on Lutz spruce in Alaska. Environmental Entomology 24(6): 1539-1547. Murphy, K.A. and E. Witten. 2006. Coastal Boreal Transition Forest. In Fire Regime Condition Class (FRCC) Interagency Guidebook Reference Conditions. Available at www.frcc.gov. NatureServe. 2008. International Ecological Classification Standard: Terrestrial Ecological Classifications. Draft Ecological Systems Description for Alaska Boreal and Sub-boreal Regions. Peters, V.S., S.E. Macdonald and M.R.T. Dale. 2005. The interaction between masting and fire is key to white spruce regeneration. Ecology 86(7): 1744-1750. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 203 of 204 Potkin, M. 1997. Fire history disturbance study of the Kenai Peninsula mountainous portion of the Chugach National Forest. Draft. USDA Forest Service, Chugach National Forest. December 5, 1997. Anchorage, Alaska Ritter, D. F. 1986. Process geomorphology. Wm. C. Brown Publishers, Dubuque, Iowa. 579 pp. Rowe, J.S. 1972. Forest Regions of Canada. Canadian Forest Service, Department of Environment. Ottawa. Inform. Can. Catalogue #FO 47-1300. Rude, M. Kenai Peninsula Spruce Bark Beetle Mitigation Program, unpublished data Schulz, B. 2000. Resurrection Creek Permanent Plots Revisited. USDA Forest Service, Forest Health Protection, Alaska Region, Anchorage, AK. Technical report R10-TP-89. 14 pp. USDA Forest Service. 2004. Forest insect and disease conditions in Alaska in 2003. U.S. For. Ser., For. Pest Manage. Tech. Rep. R10-TP-123, Anchorage, Alaska. USDA Forest Service. 2002. Final Environmental Impact Statement, Chugach National Forest Land Management Plan Revision. Available at: http://www.fs.fed.us/r10/chugach/forest_plan/feis_docs.html. Van Hees, W.W.S. and F.R. Larson. 1991. Timberland resources of the Kenai Peninsula, Alaska, 1987. U.S. For. Ser. Resour. Bull. PNW-RB-180. Portland, Oregon. 56 pp. Werner, R.A., E.H. Holsten, S.M. Matsuoka and R.E. Burnside. 2006. Spruce beetles and forest ecosystems in south-central Alaska: A review of 30 years of research. Forest Ecology and Management . 227(3):195206. Yarie J. 1983. Forest community classification of the Porcupine River drainage, interior Alaska, and its application to forest management. USDA Forest Service GTR PNW-154. Yarie J. 1981. Forest fire cycles and life tables – a case study from interior Alaska. Can. J Forest Res. 11:554-562. *Dominant Species are from the NRCS PLANTS database. To check a species code, please visit http://plants.usda.gov. **Fire Regime Groups are: I: 0-35 year frequency, surface severity; II: 0-35 year frequency, replacement severity; III: 35-100+ year frequency, mixed severity; IV: 35-100+ year frequency, replacement severity; V: 200+ year frequency, replacement severity. Sunday, November 22, 2009 Page 204 of 204