VOL. 78, NO. 30 JOURNAL OF GEOPHYSICAL RESEARCH OCTOBER 20, 1973 Distribution ofSuspended Matterin thePanama Basin WILLIAMS. PLANK,J. RONALD V. ZANEVELD, ANDI-IASONG PAX Schoolo• Oceanography,OregonState University, Corvallis, Oregon 97331 The distribution of suspended matter in the Panama basin was determined by means of light scattering and Coulter counter measurements on water samples collected at 50 hydrographic stations. The observed distribution indicates three probable sources of suspended matter: (1) the surface waters throughout the basin;' (2) erosion and runoff from the continents; and (3) bottom erosion at two sites on the Carnegie Ridge. The distribution of suspendedmatter also confirms in most. respectsthe pattern of abyssal circulation proposed by Norman P. Laird (1971). The Panama basin,boundedby the Carnegie and Cocos ridges and the coasts of Panama, Columbia, and Ecuador, has been the subject of an increasingnumber of oceanographicstudies in recent years. The circulation of the surface and intermediatelayers has been described by Stevenson[1970] and Wyrtki [1967]. Laird [1969, 1971] has hypothesized a model for abyssalcirculationin the basin basedon observations of hydrographic properties. Studies of sedimentdistribution [va,n Andel et al., 1971; Ko.wsmann, 1973; Moore et al., 1973; v.an Andel, 1973] have also provided information that has implicationsfor the abyssalcirculation in the basin. tween the Malpelo and Carnegie ridges, and ;•henspreads northward fillingthe subbasins west of the Malpelo Ridge. The distribution of oxygenand salinity in the near-bottomwater supportsthis overall pattern of circulation. An examination of the bathymetry of the ridges surrounding the basin indicates that another likely route for water entering the basin would be through the saddle over the Carnegie Ridge, which is at about 85øW and has a sill depth of about 2300 meters. Laird's studiesof hydrographicpropertiesdid not point to this saddle as a source of bottom water. However, a study of the sedimentcover in the basin [van Andel et al., 1971; Kowsmann, 1973; Moore et al., 1973] showed that this During November and December 1971 we conducteda program of optical measurements saddle is an area of active erosion and that in the Panama basin in order to delineate the distribution of suspendedparticulate matter in the basin. We hoped to use this distribution, along with hydrographic and current meter measurements,to gain further insight into the abyssal circulation and the sourcesand sinks of particulatematter in the basin. Laird [1969] hypothesizeda singlesourcefor the bottom water in the basin: the pass over the Carnegie ridge near the coast of Ecuador, which has a sill depth of about 2500 meters. The distribution of near-bottom potential temperaturesindicatesthat after enteringthe basin through this pass some of the water continues northward, parallel to the coast, filling several subbasins off the coast of Columbia. The re- maining water turns to the west, passesbeCopyright (•) 1973 by the American Geophysical Union. 7113 productsof the erosionare found along the north side of the ridge. Since the distribution of suspendedparticulateshas been utilized in several casesas an indicator of deep circulation [Jerlov, 1959, 1968; Hunkins et al., 1969; Ewing et al., 1970; Ichiye et al., 1972], the implicationof bottom erosionby northwardflowingcurrentsled us to believethat a study of suspended matter in the basinmight be useful in evaluatingthis saddle as a sourceof bottom water in the basin. Severaltechniqueshave beenusedto measure the concentrationof suspendedmatter, each of which has its limitations. Large numbers of measurementshave been made by filtering or centrifugingwater samplescollectedby sampling bottlesvaryingin sizefrom 5 liters up to hundredsof liters [Lisitzin, 1972; Ewing and Thorndike, 1965]. The difficultieswith this 7114 PLANK ET AL..' SUSPENDED MATTER -method are obvious' to obtain accurate mea- 90' 85' 80' surements, especially in the extremely clear waters found at great depths,large samplesare necessary,but the larger the samples,the more cumbersomeand time-consumingthey are to ha•ndleand analyze, and the greater is the likelihood of contamination. A modification to obtain information on the sus- pendedmatter; measurementsof light scattering and measurementsof the particle sizesby means of a Coulter counter. /• (45),(m-err) -•xI0-$ 0 .... •5 . , 5o of the filtration method that appears to hold promise is the techniqfie of in situ filtration. This method is slow, but accuracy can be improved as much as necessaryby pumping for longer times. Another techniquefor measuring suspendedmatter that has been used a great deal is the measurementof an optical property such as light attenuation or scattering [Burr, 1957; Jerlov, 1955; Thorndike and Ewing, 1967; Ochakovsky, 1966]. The difficulties involved in calibrating one such technique have been discussedby Beardsley et al..[1970]. In general,it appearsthat suchtechniquesmay be most useful for rapid surveys of large areas, and that for very accurate measurementsof the concentrationof suspendedparticles or of the particle size distribution,other methodsmay be preferable. In this study, two techniques were utilized 0o 90 = 85 = 80 = Fig. 1. B•thymetw of the Panam• basin [genera.lizedafter van Andel et al., 1971]. Stations are indicated by black dots. Letters A, B, C, and D show locations of profiles in Figure 2. Crosssectionsin Figures 3 through 6 are indicated by lines 1-1 through 4-4. Current meter locations are indicated by triangles. Bathymetric contours are in kilometers. EXPERIMENTAL PROGRAM Our study was conductedduring the 1971 cruise of the RV Yaquina to the Panama basin. At each of the 50 stationsshown in Figure 1 hydrographiccastswere made, usuallyto within 15 meters of the bottom. The water samples Scattering Cross-Section, ( u2xI0'$perI/2 ml) ;) 0 5 0 5 I Central Stahon A I0 o B Basin 4-43A C Fig. 2. Profiles of light scatteringat 45ø and scattering crosssectionof suspendedparticles at locations A, B, C, and D in Figure 1. Circles denote light scattering values. Triangles denote crosssection values. Units of abscissaapply to both parameters. PLANK ET AL.' SUSPENDEDMATTER 7115 obtainedwere analyzedfor light scatteringand direction [fi(45)] was measuredwith a Briceparticle content, in addition to temperature, Phoenix light-scattering photometer. The relaoxygen content, salinity, and the concentrations tionship between light scattering and particle of several nutrients. concentrationsin the deep sea has been discussedby Pak et al. [1971] and Beardsleyet al. ographic applications has been discussedex[1970]. In general,if the suspendedmatter is tensively [Carder, 1970; Sheldona•.d Parsons, fairly uniform in compositionand size distribu1967]. Briefly, the instrument measures the tion, the light-scattering measurementsshould height of an electronicpulse generatedby the be linearly related to total particle concentrapassageof a suspendedparticle through a small tion. These conditionsshould apply at depths orifice.The height of this pulsehas been shown greater than about 300 meters. At the two locato be proportional,within certain limits, to the tions'indicatedby trianglesin Figure 1, currents volume of the particle. In this paper 'particle STATION NUMBER diameter' means the diameter of a sphere 52 5:5 54 55 56 57 58 o having the same volume as that measuredby The use of the Coulter counter in ocean- the instrument. Particles with diametersas small as 0.4 •m can be measured with the Coulter counter. However, in the interests of rapidity and ease of analysisit was decidedto utilize a 100-•m orifice, which allows measurement of particles down to about 2 /•m. The cumulativesize distribution of particles was determined at four diameters: 2.22, 3.53, 5.60, and 10.22 •m. The volume concentration of particulate matter over this range of sizes was obtained by numerical integration, and it is this parameter whichis plottedin Figures3, 4, 5, and 6. Several investigators[Bader, 1970; Plank et al., 1972; Sheldon et al., 1972] have found the relative size frequencydistributionof particulatematter in sea water to be almost constant, especially if only samplescollectedbelowabout 300 meters are considered. The size distributions i.o :•.o :5.0 •':;i:i:!'.,!.'.:: ppmx 10-2 4.0 [] >1.4 '"•I.:•-1.4 I• I.O-I.?I•o.e-,.o [•o.•-o.e I-I< o.• LONGITUDE (west) STATION 0 5?- 5:5 NUMBER 54 55 56 57 58 of the particleswe examinedwere found in almost all cases to be nearly exponential in shape. We thus feel that an assumption of constant relative size-frequencydistribution is a good one. Measurementsof the concentrationof particles between2.2 and 10.2 •m are then proportional to the total concentrationof suspendedmatter. The proportion of the total concentrationrep- 1.0 . x•?-.o,: . :5.0 resented is difficult to determine, however, as it dependson the upper limit of sizesthat one assumesto exist in the sample and also on the shapeof the size distributionin• the submicron range. Light-scatteringmeasurementswere made on the same samplesthat were analyzed in the Coulter counter in order to obtain an inde- 4.0 • 83 8•?- 81 LONGITUDE(west) Fig. 3. Crosssectionsof the concentrationof particlesbetween2.2 and 10.2#m in ppm X 10-3 pendentmeasureof relative particle concentra- by volume (upper) and light scatteringat 45ø tion. The scattering of light having a wave- in (m str)-• X 10-3 (lower) along line 1-1 in length of 436 nm at 45ø from the forward Figure 1. were measured i meter from the bottom in an concentrationparameter in Figure 2 is geo- attempt to ga.insomeinformationabout current speedsand directionsnear the two low points on the Carnegie Ridge. metric cross section (sometimesreferred to as RESULTS Profiles of light scattering and particle concentration from the four locations marked A through •) in Figure 1 are plotted in Figure 2. These profiles are intended to indicate the general shapeof profilesfound in the varioustypes of topographic provinces in the area we are studying and to indicate the degree of correlation found between measurementsof light scattering and particle concentration.The particle STATION 56 2 :5 6 8 scatteringcrosssection),which is proportional to the % root of particle volume. The values on the abscissarepresentthe projected crosssectionalarea of the particulate matter between 2.22 and 10.22 /•m in diameter contained in 1/• ml of seawater (the size of the sample analyzed in the Coulter counter). This parameter has been previo.uslyused in studiesof the correlation of light scattering and suspended particle concentration[Beardsleye'i al., 1970]. Note that the valueson the abscissa in Figure 2 apply to both parameters. Figures 3, 4, 5, and 6 are cross sectionsof NUMBER 9 I0 II 12 0 2øS o- 5oN LATITUDE Fig. 4. Cross sections of the concentration of particles between 2.2 and 10.2 gm in ppm X 10-2 by volume (upper) and light scattering at 45ø in (m str) -• X 10-• (lower) along line 2-2 in Figure 1. PLANK ET AL.: SUSPENDEDMATTER STATION 5-21 5-7 7117 NUMBER 26 39 48 z• 5 6 7 I.O 90 88 86 LONGITUDE STATION 5-21 •l 5-7 I 90 26 I 88 I 84 82 (wesl) NUMBER 5• 48 I I 86 I 4, 5 6 7 I 84 LONGITUDE Fig. 5. Cross sections of the coricentration of particles between 2.2 and 10.2 /zm in ppm X 10-0 by volume(upper) and light scatteringat 45ø in (m str)-• X 10-a (lower) along line3-3in Figure1. light scatteringand particle concentrationalong the lines marked 1-! through 4-4 in Figure 1. Particle concentrations on these cross sections are shown in units of ppm X 10-' by volume. For comparisonwith other authors whose results are reported in terms of mass concentration, we should note that, for an assumedparticle density of 2.75 g/ml (the density of clay particles), I ppm by volume equals2.75 mg/1. Figure 7 shows contours of integrated particle volume in the lower of the water 100 and 500 meters column. The values are obtained by numerically integrating the volume of particulate matter from the bottom to 100 and 500 meters above the bottom. The contour value representsthe amountof particulatematter, in cubic microns,containedin a columnof water « cm• in cross-sectional area and 100 meters high (for the upper figure; 50'0metershigh for the lower). DISCUSSION It {S interestingto comparethe volume concentrationsof particles we measuredwith concentrationsobtained by other workers who have made measurementsover a wider range of sizes or who have used other techniques to obtain particle concentrations. 7118 [PLANEET AL.' SUSPENDED MATTER STATION 3,7 49A 3,9 $4 NUMBER 41 43, , 17 19 about 0.012 ppm to 0.014 ppm, and the minimum concentrationsin deeper waters decreased to less than 0.006 ppm. Consideringthe difference in the size rangesexamined,our measure- 16 0.6 ments are then not inconsistent with those of Sheldonet al. [1972]. In a study of suspendedmatter in the Caribbean Sea utilizing a gravimetric analysis,Bassin et al. [1972] filtered water from 5- and 30direr 1.2 :3:1.8 Niskin 2.4 I I I 3,_øS o - 3,5 3,7 49A 3,9 • and obtained minimum ma• compareswell with the valuesobtainedby both LATITUDE STATION 3,3, bottles concentrations(at depths of about 3 km) of less than 50 ttg/1. This is equivalent (for an • . _ assumed particle density of 2.75 g/ml) to a '• EILo•2 i•oe-•o ....Do6 os1--1>o6 volume concentration of 0.018 ppm. This value (•e I NUMBER 41 43, 19 I? Sheldon and us (assuming our measurements 16 ,.•..• ....• -.,:..........•:•.•::•,-:..•,••f•. 85 ø 80 ø 90 ø 85 ø 80 ø 90 ø 85 ø 80 ø 90 ø 0.6 ioo • PARTICLEVOLUME 1.2 •'-"P k /I '••'•••':•-:• I - 3.0 I [ i • I I [ I [ L•TITUDE Fig. 6. Cross sections of the concentration of particles between 2.2 and 10.2 •m in ppm X 10-2 by volume (upper) and light scattering at 45ø in (m str) -• X 10-a (lower) along line 4-4 in Figure 1. 500 •. PARTICLE VOLUME Sheldon et al. [1972] determined volume concentrations between diameters of I and 100 :?[]<2 km •>3 km ttm by meansof a Coulter counter over a wide range of depths and in several areas of the world. At 500 meters the volume concentrations they obtained varied from about 0.03 to 0.15 ppm. At 5000 meters the minimum concentrations they foufid were lessthan 0.02 ppm. If, as Sheldonfound, particle size spectra in deep waters show roughly equal amounts of material in logarithmically equal-sizedintervals, then our measurements between 2.2 and 9C• 85 ø 80 ø 10.2 ttm shouldamount to about 34% of their measurements between I and 100 ttm for water with the same total concentrationof particles. Our measurements at 500 meters averaged Fig. ?. The integrated volume of suspended particles between 2.2 and 10.2 gm in the lower 100 meters of the water column (upper) and the lower 500 meters of the water column (lower). Units of the contours are 2 /•ma X m/ml. PLANK ET AL.: SUSPENDEDMATTER are •quivalent to 34% of Sheldon's),although any comparison of gravimetric and Coulter counter results is, of course, complicatedby the different size range of particles examined by the two methods.These comparisonsindicate that our measurements over a narrow size range are not only valid in a relative sense,but are alsovaluablein indicatingroughlythe magnitude of the total amount of suspendedmatter present. The crosssectionsof Figures 3-6 show that high concentrationsof suspendedmatter occur primarily in four locations: 1. In the surface layers throughout the basin. 7119 ously, sediment samples indicate that this is the case.Current directionwas predominantly alignedwith the axesof the two passes,and in both cases180ø reversalsoccurred,indicating that the currentshave a tidal component.It is probable that these two passesare sites of advectiveand diffusiveprocesses of sufficiently high energyto erodeand maintain in suspension increasedamountsof particulatematter. The presenceof higher concentrations of particles throughout most of the water columnover the CarnegieRidge that we seein Figure 4 may be a result of the confluenceat this point of all three sourcesof suspendedmatter. That is, settling from the surface, erosion, and runoff from the shelf and bottom erosionin the vicinity 2. Over the Carnegie Ridge in the two north-south sections(Figures 4 and 6). These of the sill at about 1øS latitude all contribute high concentrationsseem to occur throughout particles to the water column near station 3, a greater range of depths in the section near resultingin high concentrationsfrom the surthe coast (Figure 4). face to the bottom. High concentrationsnear 3. On the east-westsectionin Figure 5, in the bottom both to the north and south of the the intermediate and deep waters near the Carnegie Ridge is this crosssectionmay be a coast and between 86øW and 88øW, to the result of the erosion of particles from the northwest of the saddle. saddle by the oscillatory currents mentioned 4. Near the bottom in the Peru trench above. The particles may then be transported (Figure 3) and in the deep coastalbasinto the to the north and south by gravity or advection or both. north of the CarnegieRidge (Figure 4). Three probable sources of the suspended The profiles in Figure 2 show that on the matter found in the basin are thus indicated: continentalrise (location A) and near the sad1. The •urface layers throughout the basin dle at 85øW (location C) there are wellcontributeparticlesthat may settle through the developednepheloidlayers near the bottom, water column to the bottom. as might be expectedif active erosionis occur2. The continentsuppliesmaterial by runoff ring. In the deepcoastalbasin (locationB) the and erosionon the shelf. The particle 'clouds' increasein particle concentrationis smallerin near the coast between about 300 and 1300 magnitudeand is spreadover a greaterdepth. meters in Figure 3 may indicate that the The profiles fromthe centraibasin(location Guayas River, located at the same latitude as D) demonstratethe characteristics of an area this section,is an important sourceof suspended where there is probably little erosionor sediparticles. mentation taking place in the size rangesof 3. Bottom erosion in the saddle at 85øW and probably also in the pass near the coast is injecting particlesinto the water column.The current meter measurements taken at the locations particleswe examined. Valuesof light scattering and particle concentrationare low and relatively constantbelow400 meters. In Figure7 the contoursof integratedparti- shown in Figure 1, although not of sufficient cle volumeshowa high gradientnear the coast. duration (6-7 hours) to determine the long- This indicates,first, that the continent is a term circulation, did show velocities between 7 and 10 cm/sec I meter above the bottom. Consideringthe fact that, with such a short record, the probability of seeingmaximum velocitiesis very small, it is likely that velocities high enoughto erode bottom sedimentsdo occur at these sites, and, as has been discussedprevi- majorsourceof particulatematterand,second, that much of this matter is being depositedin the deepcoastalbasin.The contoursalsoshow regions of highparticlecontent centered around the CarnegieRidgesaddleat 85øW. It appears that material eroded from the sill is probably beingspreadboth to the north and the south 7120 PLANK ET AL.: SVSPENDED MATTER and is settling out of the water column on the on the compositionand index 0f refraction of flanks of the ridge. The contoursin Figure 7 the particles. In conclusion,the observed distribution of also indicate tongues of suspendedparticles extendinginto the northwest portion of the particulate matter in the Panama basin has basin. Two explanationsfor these tongues in general confirmed the pattern of abyssal circulationsuggested by Laird [1971], although seemplausible. it does appear that the saddle at 85øW 1. The particles are erodedfrom the bottom longitude is a site of reasonably high-energy in the vicinity of the saddle at 85øW and are dynamicprocesses. However,we cannotevaluinjected by lateral mixing into the bottom ate precisely,on the basisof our measurements, water comingfrom the passnear the coast. the contributionof theseprocesses to the supply 2. The particles are eroded in the saddleat of bottom water to the baSin. 85øW and are transportedto the north by Since the work done in this study was conbottom water entering the basin over the centrated around the two saddles in the Carnesaddle. Until more direct current meter measure- mentsare availableit is difficultto say exactly what the relationshipis betweenthe observed distribution of suspended matterand the longterm mean abyssal circulation. One recent esti- mate of the settlingrate of a 5-•m sphereof density2.65 in seawater (0øC, 34%) is 1.17 X 10-• cm//sec,or about 370 meters per year [Gibbs et al., 1971]. SinceLaird's estimateof renewal timefor the basinwasabout175years, if a particleof this sizeis transportedto a region where there is no upward-directedturbulent flux of particlesfrom the bottom, it will gieRidge,•heparticledistribution in thenorthern part of the basinis as yet not well defined. Acknowledgments. This investigation has been supported by contract N00014-67-A4)369-0007 under project NR 083-102 with Oregon State University. REFERENCES Bader, H., The hyperbolic distribution of particle sizes, J. Geophys. Res., 75, 2822, 1970,. Bassin, N. Jay, John E. Harris, and Arnold H. Bouma, Suspendedmatter in the Caribbean Sea: A gravimetric analysis, Marine Geol., 12, M1, 1972. settletoo rapidlyto be of any helpin determin- BeardsIcy,G. F., Jr., HasongPak, Kendall Carder, ing the mean circulation in the basin. The significance of the light-scattering measurements relative to circulation is also difficult and Bo Lundgren, Light scattering and suspended particles in the eastern equatorial Pacific Ocean, J. Geophys. Res., 75(15), 1837, 1970. to evaluate.In viewof the fact that no particles Butt, Wayne V., On the attenuation of light in smallerthan 2.22 •m were examinedin the the sea, J. Mar. Biol. Ass. U.K., 36, 223, 1957. Coulter counteranalyses,the good correlation Carder, K. L., Particles in the eastern equatorial Pacific Ocean: Their distribution and effect •etween lightscattering andparticle concentra- upon optical parameters, Ph.D. thesis, Oregon tions shownin Figures2-6 indicatesthat either State University, Corvallis, 1970. the particlessmallerthan 2.22 •m do not con- Ewing, M., and E. M. Thorndike, Suspended tribute significantly to the scatteringmeasure- matter in deep ocean water, Science, 147, 1291, ments (which would be true if there were few 1965. circulation as do the Coulter counter measure- Ewing, M., S. L. Euttreim, J. I. Ewing, and X. Le Pichon, Sediment transport and distribution in the Argentine basin, 3, Nepheloid layer and processesof sedimentation,Ph•s. Chem. Earth, 8, 51-77, 1970. Gibbs, R. J., M.D. Matthews, and D. A. Link, The relation between sphere size and settling velocity, J. $ed. Petrol., 41(7), 7, 1971. Hunkins, K., E. M. Thorndike, and G. Mathieu, Nepheloid layers and bottom currents in the ments. It would seemto be important in the Arctic Ocean,J. Geophys.Res., 7•(28), 6995, of themor if they had lowindicesof refraction) or else the factors which influence the observed distributions(e.g., erosive forces, advection, diffusion)affect both the larger and smaller particlesin a similar way. In any event, the scattering measurementslead to the same conclusionsabout particle sourcesand sinks and future to extend our size distribution measure- ments to smallerparticlesand to utilize any available techniquesthat provide information 1969. Ichiye, Takashi, N. Jay Bassin, John E. Harris, Diffusivity of suspendedmatter in the Caribbean Sea, J. Geophys. Res., 77(33), 6576, 1972. PLANK ET AL.: SUSPENDEDMATTER $erlov, N. G., The particulate matter in the sea as determined by means of the Tyndall meter, Tellus,7,218, 1955. Jerlov, N. G., Maxima in the vertical distribution of particlesin the sea, Deep Sea Res., 5, 178, 1959. Jerlov, N. G., Optical Oceanography,194 pp., Elsevier, New York, 1968. Kowsmann, R. O., Coarse componentsin suP- 7121 Plank, W. S., Hasong Pak, and $. Ronald V. Zaneveld, Light scattering and suspended matter in nepheloid layers, Y. Geophys. Res., 77(9), 1689,1972. Sheldon,R. W., and T. R. Parsons,A practical manual on the use of the Coulter counter in marine science, 66 pp., Coulter Electronics, Toronto, 1967. ' Sheldon,R. W., A. Prakash, and W. H. Sutcliffe, face sediments of the Panama basin, eastern Jr., The size distribution of particles in the equatorial Pacific, J. Geol., in press, 1973. Laird, Norman P., Anomalous temperature of bottom water in the Panama basin, J. Mar. Res.,27(3),355, 1969. Laird, Norman P., Panama Basin deep water: Properties and circulation,Y. Mar. Res., 29(3), 226, 1971. Lisitzin, Alexander P., Sedimentation in the world ocean, Soc. Econ. Paleontol. Mineral. Spec.Publ., 17, 218 pp., 1972. Moore, T. C., Jr., G. R. Heath, and R. O. Kowsmann, Biogenicsedimentsof the Panama basin, J. Geol.,in press,1973. Ochakovsky,Yu. E., On the dependenceof the total attenuation coefficient upon suspensions in the sea, U.S. Dep. Commerce,Joint Publ. Res.,Rep.,36(816),16,1966. Pak, Hasong, J. Ronald V. Zaneveld, and G.F. ocean,Limnol. Oceanogr.,17(3), 327, 1972. Stevenson,Merritt, Circulation in the Panama bight, J. Geophys.Res., 75(3), 659, 1970. Thorndike, E. M., and M. Ewing, Photographic nephelometersfor the deep sea, in Deep-Sea Photography,edited by J. B. Hersey, pp. 113116,JohnsHopkins Press,Baltimore, Md., 1967. van Andel, Tjeerd, G. Ross Heath, Bruce T. Malfait, Donald F. Heinrichs, and John I. Ewing, Tectonics of the Panama basin, eastern equatorial Pacific, Geol. Soc. Amer. Bull., 82, 1489,1971. van Andel, Tjeerd H., Texture and dispersal of sedimentsin the Panama Basin, J. Geol., in press,1973. Wyrtki, Klaus, Circulation and water massesin the eastern Pacific Ocean, Int. J. Oceanol. Limnol., 1(2), 117,1967. Beardsley, Jr., Mie scattering by suspended clay particles,J. Geophys.Res., 76(21), 5065, 1971. (ReceivedApril25,1973; revised June 25, 1973.)