Geomorphology 139–140 (2012) 384–402 Contents lists available at SciVerse ScienceDirect Geomorphology journal homepage: www.elsevier.com/locate/geomorph Structural organization of process zones in upland watersheds of central Nevada and its influence on basin connectivity, dynamics, and wet meadow complexes Jerry R. Miller a,⁎, Mark L. Lord a, Lionel F. Villarroel a, Dru Germanoski b, Jeanne C. Chambers c a b c Department of Geosciences & Natural Resources, Western Carolina University, Cullowhee, NC 28723, United States Department of Geology and Environmental Geosciences, Lafayette College, Easton, PA 18042, United States Rocky Mountain Research Station, USDA Forest Service, 920 Valley Road, Reno, NV 89512, United States a r t i c l e i n f o Article history: Received 24 June 2011 Received in revised form 4 November 2011 Accepted 6 November 2011 Available online 12 November 2011 Keywords: Fluvial geomorphology Sensitivity Connectivity Groundwater recharge Great Basin a b s t r a c t The drainage network within upland watersheds in central Nevada can be subdivided into distinct zones each dominated by a unique set of processes on the basis of valley form, the geological materials that comprise the valley floor, and the presence or absence of surficial channels. On hillslopes, the type and structure (frequency, length, and spatial arrangement) of these process zones is related to the lithology and weathering characteristics of the underlying bedrock. Process zones dominated by sediment accumulation, storage, and groundwater recharge are associated with less resistant rocks that weather into abundant but relatively small particles. Sediment transport and runoff-dominated zones are associated with resistant, sparsely fractured rocks that produce limited but larger clasts. The type and structure of process zones along axial valleys depend on the characteristics of the process zones on the hillslopes. Numerous sediment storage-dominated reaches leads to a relatively high number of unincised fans located at the mouth of tributaries along the axial valleys and to frequent and lengthy unincised valley segments, both of which disconnect large sections of the drainage basin from channelized flows. In contrast, a relatively high density of transport-dominated process zones leads downstream to the incision of side-valley fans and axial valley deposits as well as a high degree of basin connectivity (defined by the integration of surficial channels). Connectivity also is related to the lithology of the underlying bedrock, with higher degrees of connectivity being associated with volcanic rocks that presumably yield high rates of runoff. Lower levels of connectivity are associated most frequently with extensively fractured, locally permeable sedimentary and metamorphic rock assemblages. Thus, basins underlain by volcanic rocks appear to be more sensitive to incision and produce more dynamic channels in terms of the rate of channel/valley modification than those underlain by other lithologies. Considerable attention has been devoted in recent years to the management of wet meadow ecosystems that serve as important riparian habitats within upland basins of central Nevada. The data presented here show that, within basins characterized by a high degree of connectivity, areas of wet meadow are minimal. Where they do exist in these basins, they tend to be severely degraded by incision or gully erosion and will be difficult to manage given the dynamic nature of the axial channel processes. Wet meadows within basins that exhibit a low degree of connectivity and high sediment storage-to-transport ratios on hillslopes will likely be more responsive to management activities because of the reduced threats of channel incision and, presumably, a larger supply of groundwater flow to the meadows created by an extensive network of recharge sites. Importantly, human activities that lead to an increase in basin connectivity can negatively impact downstream meadows through a decrease in groundwater recharge and an increase in stream dynamics, in spite of the fact that these activities may be physically separated from the wet meadow areas. © 2011 Elsevier B.V. All rights reserved. 1. Introduction The Great Basin forms the largest section of the Basin and Range Physiographic Province and is composed of a series of north–northwest-trending mountain ranges separated by alluviated, ⁎ Corresponding author. Tel.: +1 828 227 2269. E-mail address: jmiller@wcu.edu (J.R. Miller). 0169-555X/$ – see front matter © 2011 Elsevier B.V. All rights reserved. doi:10.1016/j.geomorph.2011.11.004 intermountain basins formed by high angle extensional faulting. The ranges within the central Great Basin comprise >40% of the landscape and reach elevations 1500 to 2000 m above the surrounding valley floors. Precipitation varies significantly with elevation but may reach 55 cm at upper elevations and decreases to as little as 20 cm within the intermountain valleys (Chambers and Miller, 2004). As a result of the limited precipitation, particularly at lower elevations, riparian areas within the Great Basin encompass b1% of the total landscape and are found mostly within small (b100 km 2) J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 upland watersheds. Despite their limited size, riparian areas support most of the region's native biodiversity including the Lahontan cutthroat trout (Oncorhynchus clarki henshawi), which is listed as threatened under the U.S. Endangered Species Act (Dunham et al., 1997). Unfortunately, an estimated 50% of the riparian areas in the Great Basin, as in many semi-arid regions, are in poor ecological condition (Jenson and Platts, 1990). One of the highest priority ecosystems for management and restoration are riparian meadow complexes. These meadow ecosystems are hydrogeomorphic features characterized by elevated water tables and riparian grasses and broad-leafed plants (i.e., riparian obligate graminoids and forbs). Emphasis on their management and restoration results from the fact that they represent unique habitats that in many cases are being degraded by 385 ongoing stream incision and its associated impacts on shallow groundwater flow. Numerous geomorphic, hydrologic, and ecologic studies have been conducted in the region during the past decade with the intent of providing the information necessary to develop sound management practices for riparian areas (Chambers et al., 1998, 1999; Castelli et al., 2000; Chambers and Linnerooth, 2001; Germanoski et al., 2001; Martin and Chambers, 2001a,b, 2002; Chambers and Miller, 2004,2011). These studies have shown that many wet meadow complexes are located upstream of alluvial fans formed at the mouth of tributaries (referred to here as side-valley fans) and other forms of valley constriction (Fig. 1). The valley, then, is characterized by a distinct segmented pattern with regards to valley form, Side-valley fan A B C Fig. 1. (A) Side-valley fan located at mouth of tributary. During the Holocene, fan prograded across valley and inhibited downstream flow of water and sediment; (B) Aerial view of wet meadow complex located upstream of side-valley fan; and (C) Channel incised into wet meadow complex. Photograph is looking downvalley toward side-valley fan show in (A) and (B). Reach is classified as an incised alluvial valley. 386 J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 Table 1 Summary of basin sensitivity types identified by Germanoski and Miller (2004); table modified from Chambers et al. (2004). Category Geology Geomorphic characteristics Watershed sensitivity Group 1 Flood-dominated Tertiary volcanic rock Very high Group 2 Deeply incised channels Tertiary volcanic rock Group 3 Fan-dominated Sedimentary and meta-sedimentary rock Group 4 Pseudo-stable channels Intrusive igneous, and sedimentary rock • • • • • • • • • • • • • • High relief basins Narrow bedrock controlled Minimal sediment storage Multiple, discontinuous terraces Large, high relief basins Incised channels/trenched fans Multiple, semi-continuous terraces Lower relief basins Large side-valley fans Metastable channels with low incision values, but active downcutting Moderate/minor incision Potential for catastrophic incision via groundwater sapping Cobbles or smaller bed material Multiple, discontinuous terraces where incised processes, and deposits where wet meadows are present. Along the constricted reach, the channel typically exhibits a steep gradient and is bound by bedrock and coarse-grained alluvium and/or colluvium often associated with side-valley fans. This high gradient reach typically links higher and lower elevation stream segments creating a “stepped” longitudinal channel profile. Stratigraphic studies in nearly 30 basins revealed that fan deposition primarily occurred during an intensive period of hillslope erosion extending from ~2580 ± 70 to 1980 ± 60 YBP (Miller et al., 2001). During this period many sidevalley fans temporarily blocked the downvalley movement of water and sediment producing upstream zones of sediment accumulation and low valley gradients. Downstream of the fans, reworked fan and other alluvial sediments were deposited along the valley floor creating an episode of channel/valley aggradation marked by a distinct alluvial unit that is often several meters in thickness. Since about 1980 YBP, stream systems throughout the region have exhibited periods of stability that were separated by episodes of channel incision often by the headward migration of knickpoints (Miller et al., 2001, 2004, 2011). Detailed hydrologic studies that involved sediment coring, seismic and ground penetrating radar surveys, and the installation of numerous piezometers (more than 90 at one site) have been carried out within five wet meadow complexes (Lord et al., 2011). These analyses have shown that wet meadow formation requires (i) a significant upstream supply of water, (ii) some form of valley constriction that leads to reduced downvalley flow rates and upward flow patterns, and (iii) fine-grained valley fill deposits that retard groundwater flow to create shallow water tables. Thus the integrity of meadow complexes is strongly dependent on the magnitude of groundwater recharge and the flow of groundwater along the drainage network as well as the magnitude of channel incision that has the potential to lower water table levels within the meadow complexes. Germanoski and Miller (2004) found that the rates, magnitude, and nature of channel incision vary spatially between basins and along the axial valley of any given basin. Such differences in response are often explained in terms of landform sensitivity, which is defined by Brunsden and Thornes (1979) as “the likelihood that a given change in the controls of the system will produce a sensible, recognizable, and persistent response [in the landform of interest].” This definition of sensitivity includes two distinct components of change: (i) the likelihood for a geomorphic feature such as a stream reach to change as governed by a set of resisting and driving forces, and (ii) the ability of a stream to remain in an equilibrium state (Schumm, 1991; Downs and Gregory, 1995). The rate of change also is important because, while a stream may be in a state of disequilibrium, it may exhibit little change if the driving and resisting forces are such that adjustments occur at an exceedingly slow rate. The analysis by Germanoski and Miller (2004) showed that the examined watersheds (nearly 20 in total) could be subdivided into Low to moderate Low to moderate Moderate to high four distinct groups characterized by different degrees of landscape sensitivity and channel dynamics (Table 1). The observed differences in sensitivity represent a powerful management tool as they provide insights into (i) the likelihood that a given river or meadow will respond to future disturbances, (ii) the timing, duration, rate, and nature of the response, and ultimately (iii) the potential for a given system to be stabilized or restored (Chambers et al., 2004; Germanoski and Miller, 2004). However, three short-comings of the sensitivity analysis were noted. First, differences in basin dynamics could not be entirely explained using commonly applied methods of basin morphometric analysis. Second, it was apparent that channel, valley floor, and meadow ecosystem sensitivity varied along the riparian corridor of a given basin. Classifying the sensitivity of an entire watershed did not adequately describe the spatial variations in geomorphic response that occurred within a basin or the controls on channel/valley sensitivity within a given reach of the drainage system. Third, the analysis did not directly attempt to integrate groundwater flow patterns and processes into the study. An improved understanding of recharge mechanisms, local water table elevations, and their relationships to streams permits a more holistic approach to sensitivity. Since the early 1990s, the classical continuum view of river systems has been questioned and often replaced by a hierarchical perspective of a drainage network (Frissell et al., 1986; Kishi et al., 1987; Grant et al., 1990; Montgomery and Buffington, 1993; Grant and Swanson, 1995; Brierley and Fryirs, 2001, 2005). This conceptual shift is related to the realization that hierachical classifications are extremely useful in understanding the interactions between processes functioning over differing temporal and spatial scales and for capturing abrupt spatial variations in channel and valley form, processes, and behavior that are not adequately described by the classical continuum approach. While these classification systems differ in their specifics, the unifying theme is that drainage networks consist of channel and valley floor environments that can be subdivided into progressively smaller units, each of which are morphologically homogeneous with respect to landforms, processes, and the controlling factors of geology, vegetation, substrate, and hillslope influences (Grant and Swanson, 1995). Commonly included categories range from localized channel-scale units (defined on the basis of various channel-bed features such as pools, riffles, bars, etc.), reach-scale units (defined according to the nature of both the channel and valley floor), and larger scale units ranging up to and beyond the entire drainage basin. Application of the hierarchical approach has been found to be particularly well suited to the investigation and management of mountainous streams where factors external to the channel play a more significant role in controlling channel and valley form than is the case for lowland watersheds (Montgomery and Buffington, 1993; Grant and Swanson, 1995; Montgomery and Buffington, 1997). J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 These external (nonfluvial) constraints include passive factors such as bedrock outcrops as well as mass wasting and other hillslope processes that produce localized accumulations of colluvial and alluvial materials (Grant and Swanson, 1995). The significance of these external constraints is that semicontinous downstream changes in channel/valley form and process observed along many lowland rivers are replaced by a segmented drainage net where individual segments terminate abruptly along the network and may be repeated multiple times throughout the drainage system. Often referred to as process zones, these reach-scale units include segments of the stream channel and adjacent valley floor with homogenous landforms and processes. The objective of this study is to determine if the hierarchical approach can more effectively explain spatial variations in geomorphic response to late Holocene perturbations and the present distribution of wet meadow complexes. We examine the linkages between channel dynamics, wet meadow complexes and the spatial structure and connectivity of process zones at reach and basin scales. Variations in the spatial structure (type, length, and spatial arrangement) of delineated process zones on basin- and reach-scale sensitivity and the factors that control these variations are also examined. Connectivity, as used here, refers to the degree to which water and sediment can be transferred from one zone to the next downstream zone on the surface through a distinct channel. Thus the adopted definition of connectivity does not include groundwater flow. 2. Methods 2.1. Basin selection Germanoski and Miller (2004) conducted an extensive study of landscape sensitivity within 18 watersheds in central Nevada. They found that the watersheds could be subdivided into four groups with respect to the style of geomorphic adjustment and the sensitivity of the axial channel to change during the late Holocene (Table 1). The current study utilizes eight watersheds that are considered to be representative of these four groups. The studied basins are Barley Creek, Birch Creek, Cottonwood Creek, the north and south branch of Corcoran Canyon, Kingston Canyon, Pine Creek, San Juan Creek, 387 Table 2 Basin characteristics of studied watersheds. Kingston C. Birch Cr. San Juan Cr. Cottonwood Cr. N. Corcoran C. S. Corcoran C. Barley Cr. Pine Cr. South Twin R. Basin area (km2) Basin relief (m) Divide elevation (range, m) Drainage density (km/km2) Drainage frequency (#/km2) 60.46 46.09 27.34 20.54 8.06 7.03 89.22 31.66 49.90 1546 1400 1044 1268 508 555 1005 1309 1674 3497–2597 3290–2067 3117–2228 3341–2292 2757–2292 2805–2292 3212–2457 3601–2737 3588–2701 3.21 4.11 3.37 3.53 3.59 4.42 2.97 3.31 4.42 11.5 17.5 11.7 11.1 14.3 18.4 9.10 11.8 18.4 and the South Twin River (Fig. 2). The north and south branches of Corcoran Canyon were examined separately because the two subbasins exhibit differences in the number and area of contained wet meadow complexes. General topographic and morphologic characteristics of the studied basins are provided in Table 2. 2.2. Delineation, characterization, and mapping of process zones Elements of the hierarchical classification systems presented by Frissell et al. (1986), Montgomery and Buffington (1993), and Grant and Swanson (1995) were modified, combined and fit to the characteristics of the watersheds in central Nevada for this study. In general, the watersheds were subdivided into hillslope, low-order valley, axial valley, and side-valley fan components (Fig. 3). Each component was then subdivided further into process zones on the basis of the composition of the valley fill or underlying geological materials (bedrock, colluvium, alluvium), and the presence or absence of a distinct channel throughout the length of the reach (Fig. 3). The classification resulted in 13 process zones (Fig. 3). Delineation and mapping of process zones involved a multistepped approach beginning with the classification and mapping of distinct reaches of the drainage network on mylar overlays using B 4 1 3 A 2 7 6 8 5 25 km Fig. 2. (A) Distribution of the Basin and Range Physiographic Province in the U.S.; (B) map showing the location of the study basins in central Nevada. 1 — Kingston Canyon; 2 — Cottonwood Creek; 3 — San Juan Creek; 4 — Birch Creek; 5 — Barley Creek; 6 — Corcoran Creek; 7 — Pine Creek; and 8 — South Twin River; aerial imagery for part B from Google Earth™. 388 J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 Fig. 3. Classification system used to categorize process zones within the study basins. 2006, 1:12,000-scale, georectified aerial photographs. Where an accurate designation could not be determined on the georectified photos (in two-dimensions), 1993 photographs allowing stereo viewing were consulted. Google Earth™ images also were used to gain insights into the nature of the valley system. Once mapped, paper copies of the mylar overlays were used to check the accuracy of the mapped process zones in the field. The maps were then edited to reflect map revisions made during field inspections (see Fig. 4 as an example). The number and length of each process zone type was subsequently determined from the constructed maps and combined with basin morphometric data to assess frequency (#/km 2) and density (total length in km/km 2) of the process zones within the individual study basins. All length–area measurements were made with a Planix digital planimeter. Downstream patterns in process zone types were determined by examining the frequency with which specific zones were linked to one another. The process zone maps also were compared to existing maps of bedrock geology to assess geological controls on the nature and structure of the process zones. Field characterization of the process zones involved the random selection of ~10 characterization points for the predominant types of process zones within each of the study basins. Some process zones, such as incised alluvial valleys that comprise large sections of the axial channel network, were more intensively characterized in order to capture downstream differences in channel morphology. Other process zones were either too dangerous to access (e.g., steep bedrock valleys), or rarely occurred in the basins (e.g., bedrock hollows) and were not quantitatively characterized in the field. Data were collected at a total of 389 sites; the number of reaches characterized within an individual basin ranged from 20 to 108. The parameters collected at each characterization point included descriptors of hollow/valley/channel form, bed material lithology, hydrologic features, and grain size distribution of the bed material (Table 3). Cross section/valley measurements were made using the tape/stadia rod method, whereas gradients were measured using a laser level. Particle size distribution of the bed material was determined using an Excel-based grain size analysis algorithm developed by the U.S. Forest Service Stream Technology Center (http://www.stream.fs.fed. us/publications/software.html) on 50 randomly selected clasts measured with a gravelometer. Approximately 9700 clasts were examined within the seven basins (field data were not collected from the South Twin River because of access and other problems). The long, intermediate, and short axes of the ten largest clasts also were measured at each site. Hydrologic features, such as flow conditions (water depth) and springs, were recorded where present. 2.3. Characterizing process zone connectivity The geomorphic and hydrologic connectivity of the system are highly dependent on the time scale under consideration, particularly in semiarid environments such as that in central Nevada. Our previous work in the region, for example, demonstrates that an integrated drainage system may be hydrologically connected only during the seasonal snowmelt period, but disconnected during the late summer months when the channel possesses both perennial and ephemeral reaches (Lord et al., 2011). Over longer timescales (years to decades), sections of the drainage network may become incised, thereby increasing connectivity between the zones, or become filled, creating a discontinuous drainage system with decreased connectivity between process zones. For this analysis, the focus is on channel connectivity of the system over a period of years, and connectivity is operationally defined as the existence of a recognizable, physically integrated channel (Hooke, 2003). Thus, process zones that do not have a channel (e.g., unincised colluvial or alluvial valleys, or unincised side-valley fans) represent zones that disconnect upstream portions of the drainage network from those farther downstream. While some surface water connection may occur during extreme hydrological events where channelized systems are lacking, the ability of unincised reaches to transfer water, sediment, and other constituents downstream is significantly less than along channelized reaches of the drainage network. Connectivity of the system was quantified by mapping and measuring the amount of connected basin area immediately up- and downstream of selected tributaries to the axial drainage in terms of both an absolute value and percentage of the total upstream basin area. We had originally intended to determine the relationships between such parameters as process zone length, slope, abundance, particle size, contributing basin area, etc. using multivariate statistical methods. However, the inability to safely access steep, bedrock hollows and valleys locally covered by a thin veneer of loose sediment prevented collection of the necessary data. Nonetheless, basic statistical/empirical data (e.g., data means, ranges, and standard deviations) combined with field observations and the available cartographic information provide insights into the relationships between geology, process zone type and structure, and drainage net connectivity. All statistical analyses were conducted using SYSTAT 9. J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 389 Fig. 4. Map of process zones produced for Kingston Canyon. 3. Results 3.1. Process zone characteristics 3.1.1. Hillslope hollows Hillslopes within the upland watersheds possess three types of U-shaped depressions that are commonly referred to in studies of Table 3 Summary of collected parameters. • Basin/subbasin • Latitude/longitude (GPS) • Grain size distribution of bed material (D50, D84, size of 10 largest); bedrock exposures • Clast lithology (for 50 clasts and 10 largest) • Valley and channel slope • Valley width (where applicable) • Water width and depth (if present) • Channel or hollow cross profile • Bank material composition; stratification • Bank failure mechanisms • Groundwater seepage; springs • Channel bed features (pools, riffles, etc.) • Maximum hillslope gradients • Landforms present on valley floor • Underlying bedrock geology basin morphometry as zero-order channels (Tsuboyama et al., 2000; Sheridan and Spies, 2005). These hillslope features are referred to here as ‘unincised hollows’, ‘incised hollows’, and ‘bedrock hollows.’ The morphologic and sedimentologic characteristics of each are described in Table 4 and below. 3.1.1.1. Unincised hollows (UH). The most frequently observed type of process zone on hillslopes is called ‘unincised hollows’. These features represent the first recognizable, upslope segment of the drainage network. They are devoid of an integrated surface channel with welldefined banks (Fig. 5A). Rather, the floor of the depression is characterized by angular, well-sorted, highly porous gravels of relatively small size (D50 of ~ 17 mm, Fig. 6). Sagebrush (Artemisia tridentata spp), wild rose (Rosa woodsii), and other upland species partially cover axial deposits that typically exhibit thicknesses on the order of 0.5 to >1 m. Bedrock outcrops within the floor of the unincised hollows are rare, although they may occur along the margins. Slope of the unincised hollows is variable, but on average is relatively high in comparison to the other types of process zones as a result of their position within the drainage network (Fig. 6). 390 J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 Table 4 Summary of geomorphic and hydrologic process zone characteristics. Process zones on hillslopes Process zones within low-order valleys Process zone Morphology Sediment storage Dominant process(es) Unincised hollow (UH) U-shaped depression oriented roughly parallel to slope Incised hollow (IH) U-shaped depression oriented roughly parallel to slope; channel incised in center of depression U-shaped depression oriented roughly parallel to slope V-shaped valley oriented roughly parallel to slope; channel does not possess welldefined banks; channel often floored by bedrock with local knickpoints present Broad, rectangular shaped channel formed in resistant, scarcely fractured bedrock Rectangular to asymmetrical channel formed in thick alluvial deposits; channel often characterized by pools, riffles, and point bars Well-sorted, angular sediment of limited lithologic variation; sediment typically b1–2 m thick Well-sorted, angular sediment of limited lithologic variation along margins of bedrock-floored channel No continuous layer of sediment present; occasional large clasts present Very little sediment found within the valley Deposition of sediment from adjacent hillslopes; significant recharge with limited runoff Rapid transport of water and sediment derived from adjacent hillslope; incision associated with single, catastrophic event Rapid transport of water and sediment derived from adjacent hillslope Transport of water and sediment delivered to the channel and valley from upstream channel reaches and adjacent hillslopes No continuous layer of sediment present; occasional large boulders located along depression Thick alluvial sediments (often thicker than 5 m) within channel bed and adjacent floodplains; sediments exhibit wide range of lithologies, moderately sorted, and moderately rounded Transport of water and sediment delivered to the channel and valley from upstream channel reaches and adjacent hillslopes Transport of surface water and sediment delivered to the channel from upstream reaches. Significant groundwater recharge through channel bed where coarse grained and a deep water table; sediment is periodically deposited within channel and floodplain, and moved during large events Deposition of sediment delivered to the valley floor during flood events; significant zone of recharge Transport of sediment through continuous channel; recharge into bed sediment locally important Bedrock hollow (BH) Unfilled valley (UV) Bedrock valley (BV) Process zones within axial valleys Process zones within axial valleys Process zones associated with side-valley fans Incised alluvial valley (IAV) Unincised alluvial valley (UAV) Incised colluvial valley (ICV) Low-relief valley floor surface with no continuous channel Rectangular to asymmetric channel with variable bank heights formed in colluvial and side-valley fan sediments Incised alluvial/ colluvial valley (IACV) Rectangular to asymmetric channel with variable bank heights formed in colluvial and side-valley fan sediments Anabranching valley (ANBV) Multi-channel system in which channels are separated by vegetated islands; numerous debris dams present Fan-shaped accumulations of sediment at mouth of tributaries; no continuous channel present Fan-shaped accumulations of sediment at mouth of tributaries; channel extends from fan apex to toe Unincised side-valley fan (UF) Incised side-valley fan (IF) The highest density (km/km 2) and frequency (#/km2) of the unincised hollows (measured as total length/number of the hollows divided by basin area) are found in Kingston Canyon, Cottonwood Creek, San Juan Creek, Birch Creek, and the South Twin River (Fig. 7). In constrast, Barley Creek and Corcoran Canyon exhibit a low density and frequency of unincised hollows. 3.1.1.2. Incised hollows (IH). These features exhibit a well-defined channel that typically exceeds 1 to 2 m in depth and several meters in width and that is floored by bedrock. In every other way, incised hollows are morphologically similar to unincised hollows. In comparison to unincised hollows, incised hollows are scarce (b0.4/km 2); they are most common, and exhibit the highest densities, within the Pine Creek basin and to a lesser degree the South Twin River basins (Fig. 7). 3.1.1.3. Bedrock hollows (BH). Bedrock floored, U-shaped depressions, shown in Fig. 5C, are called ‘bedrock hollows’. Although they are morphologically similar to the sediment-filled, unincised hollows, bedrock hollows lack continuous accumulations of unconsolidated debris. Detailed measurements of their form (cross-sectional morphology and gradient) were not collected in the field because of the dangers of traversing slick bedrock on steep slopes. Nonetheless, field observations suggest that they exhibit similar dimensions, form, and gradients as the other types of hollows. Bedrock hollows Surface underlain by alluvial deposits often >5 m thick; significant zone of sediment accumulation Channel bed sediment overlying colluvial material of considerable thickness; colluvial valley fill associated with past episodes of hillslope erosion and deposition Channel bed sediment overlying colluvial material of considerable thickness; often characterized by colluvial and alluvial sediments on opposite sides of channel Thin (b2.5 m thick) deposits consisting of cobble-sized sediment with occasional boulders overlying bedrock Thick (> 10 m) accumulations of alluvial and colluvial sediments Thick (> 10 m) accumulations of alluvial and colluvial sediments Transport of sediment through continuous channel; some deposition on alluvial floodplain; recharge into bed sediment locally important Spatially and temporally variable episodes of deposition and erosion; highly dynamic valley floors Deposition of sediment; significant zone of groundwater recharge Represent long-term zone of deposition; currently, zone of sediment and water transport occur most frequently and exhibit the highest densities within the Barley, Pine, Corcoran, and South Twin River basins (Fig. 7). 3.1.2. Low-order valleys Two types of process zones, referred to here as ‘bedrock valleys’ and ‘unfilled valleys’, receive water and sediment from hillslopes and hillslope hollows. Both are found at relatively high elevations within the watersheds and are considered low-order channels by the classical Horton (1945) and Strahler (1952) drainage net classification systems. 3.1.2.1. Unfilled valleys (UV). Process zones that exhibit a V-shaped cross-sectional valley form and are characterized by hillslopes that grade directly into a bedrock-dominated channel are referred to as ‘unfilled valleys’ (Fig. 5B). Alluvial and colluvial deposits within the valley bottom are limited. Locally, small knickpoints are present in the bedrock, indicating channel bed incision. The unconsolidated sediment that is present tends to possess a larger median grain size and is more poorly sorted than sediment found in hillslope hollows (Fig. 6). The average gradient of unfilled valleys is approximately half of that observed for the upslope hollows. These features are relatively common, typically occurring at higher frequencies and densities than the bedrock valleys described below (Fig. 7). 3.1.2.2. Bedrock valleys (BV). In contrast to unfilled valleys, bedrock valleys exhibit relatively wide valley floors that often reach widths J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 A B C D E F 391 Fig. 5. Photographs showing typical morphology and composition of the major process zones mapped within the study basins. (A) unincised hollow; (B) unfilled valley; (C) bedrock hollow (U-shaped feature) draining into bedrock valley); (D) rugged nature of bedrock outcrop associated with bedrock valleys; (E) incised alluvial valley; and (F) incised colluvial valley. of 5 to >10 m (Fig. 5C). Unconsolidated sediment, typically eroded from the underlying bedrock, tends to be larger and more poorly sorted than is found in the other types of process zones (Fig. 6). Bedrock valleys occur most frequently and exhibit the highest densities in the Barley, Corcoran, and the South Twin River basins (Fig. 7). They are also common in the Kingston Canyon watershed where they are predominately located on carbonate rocks (as will be discussed in more detail below). 3.1.3. Axial valleys Higher-order, lower elevation segments of the drainage network are characterized by relatively wide valleys (measured in 10s of meters) that have been partially filled with alluvial and colluvial sediments (Figs. 1C and 5E). These process zones are collectively referred to as ‘axial valleys’. It is important to note, however, that they not only occur along the predominant (main) channel within the watershed, but often extended into the lower reaches of a tributary basin where they serve as the axial drainage route within that subbasin. Axial valleys are subdivided into distinct process zones on the basis of (i) the composition of the valley fill, and (ii) the presence or absence of a continuous channel (Fig. 3). The four recognized types are described below. 3.1.3.1. Incised alluvial valley (IAV). In most basins, the majority of axial valley floors consist of incised alluvial valleys (Fig. 5E). Incised alluvial valleys are characterized by a continuous (integrated) channel cut into alluvial sediments that comprise the majority of the valley fill. Core data collected from five basins in the area demonstrate that the thickness of the valley fill is often measured in a few tens of meters. Sedimentologically, the fill is highly variable, ranging 392 J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 Fig. 6. Slope and grain size characteristics of channel or valley materials for predominant types of process zones. Values based on data from all seven study sites for which field data are available. Largest clast values based on intermediate diameter of 10 largest clasts found at characterization site. Filled rectangle — mean; box — standard deviation; line across box — median intermediate diameter; whiskers — 10 and 90% exceedence values. UH — unincised hollow; IH — incised hollow; BH — bedrock hollow; UV — unfilled valley; BV — bedrock valley; ICV — incised colluvial valley; IACV — Incised alluvial/colluvial valley; IAV — Incised alluvial valley; and UAV — Unincised alluvial valley. from coarse gravel-sized material to fine-grained silt- and claydominated units (Lord et al., 2011). Bed material within the channels exhibits a wide range of particle sizes, but is typically larger than clasts found in unincised hollows and smaller than those found in bedrock hollows and valleys (Fig. 6). Lithologically, the clasts are more diverse than the particles found in higher elevation process zones reflecting a wider range of source rocks. The average slope of incised alluvial valleys is variable, but is relatively low (b0.1) in comparison to other types of process zones found along the axial valley (Fig. 6). The frequency of incised alluvial valleys is similar across the study basins with the exception of Corcoran Canyon (Fig. 8). The high frequency within the north and south Corcoran Canyon basins reflects (i) the relatively small area of the basin (Table 2), and (ii) incised alluvial valley segments that are frequently disrupted by colluvial landforms. The density of incised alluvial valleys is more variable between basins than other types of axial valley zones, with larger densities found in the Birch Creek, Barley Creek, and Corcoran Canyon basins (Fig. 8). The depth of channel incision below the valley floor is highly variable, among basins and along the riparian corridor of a given basin, ranging from a few tens of centimeters to more than 5 m. Downstream variations within Kingston Canyon, Cottonwood Creek, San Juan Creek, and Birch Creek tend to alternate between deeply incised reaches and reaches with limited entrenchment, until reaching the basin mouth where incision tends to increase to a maximum. As an example, observed downstream variations in incision along Kingston Canyon are shown in Fig. 9. 3.1.3.2. Incised colluvial valleys (ICV). These features, shown in Fig. 5F, represent the second most abundant type of axial valley (Fig. 8). They are characterized by a continuous channel cut into colluvial valley fill or fan deposits that slope from the hillslope toward the valley axis. Incised colluvial valleys are located in two primary locations: (i) within tributaries well upstream of the tributary basin's mouth, and (ii) along the main channel at the toe of side-valley fans. These process zones, on average, are slightly steeper and possess slightly larger material than is found within the incised alluvial valleys (Fig. 6). A few reaches within the Kingston Canyon, Birch Creek, San Juan Creek, and Cottonwood Creek basins consist of incised alluvial/colluvial valleys. As the name implies, these process zones are transitional in character between incised alluvial and incised colluvial valleys in that the channel is cut into both alluvial and colluvial valley fill deposits. They tend to possess higher channel gradients and coarser bed material than is found along the other types of axial valley process zones (Fig. 6). 3.1.3.3. Unincised alluvial and colluvial valleys (UAV/UACV). In contrast to incised alluvial and colluvial process zones, unincised alluvial and unincised alluvial and colluvial valleys lack an integrated channel network. Although the valley floor deposits may locally possess shallow (b1 m deep) gully systems, they are typically characterized by a low relief valley floor covered in sagebrush and other vegetation. The surface sediments in most instances are composed of fine-grained (siltdominated) deposits. In a few locations, however, upstream axial channels transport sediment onto the valley floor on a semiannual J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 393 Fig. 7. Density and frequency of selected hillslope and low-order channel process zones stratified by studied basin. UH — unincised hollow; IH — incised hollow; BH — bedrock hollow; UV — unfilled valley; and BV — bedrock valley. basis, allowing for localized valley floor aggradation. These reaches tend to exhibit, on average, the lowest channel/valley floor gradients and the smallest particle sizes of any process zone type (Fig. 6). 3.1.3.4. Anabranching channels (ANBV). Downstream segments of the axial channel in Pine Creek and the South Twin River exhibit an anabranching pattern that was not observed in the other study basins. Fig. 8. Density and frequency of axial valley process zones stratified by studied basin. ICV — incised colluvial valley; IAV — incised alluvial valley; IACV — incised alluvial/colluvial valley; IABV — anabranching valley; UAV — unincised alluvial valley; and UACV — unincised alluvial/colluvial valley. 394 J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 Incision depth (m) 4 system (called ‘incised side-valley fans’), and (ii) fans that are devoid of a defined channel or that possess a channel only in apex or medial areas of the fan before flowing onto the fan surface (referred to as ‘unincised side-valley fans’) (Table 4). With the exception of Birch Creek, the number of side-valley fans per unit basin area is consistent across the study basins, ranging from ~0.7 to 1.5/km 2. Birch Creek possesses a high number of side-valley fans (Table 5), many of which are located within wide tributary basins (in comparison to the other studied watersheds). In contrast to the density of fans within the basins, significant differences exist in the proportion of incised to unincised fans between the basins (Table 5). A higher percentage of unincised fans occur in the Kingston Canyon, Cottonwood, San Juan, and Birch Creek basins than is found within the other studied watersheds. 3 2 1 0 % Distance downstream Fig. 9. Downstream variation in maximum depth of incision below valley floor in Kingston Canyon. Data points represent location of measured cross sections. Trend, which exhibits considerable variability in the depth of incision along the axial valley, is typical of basins with high storage-to-transport ratios for hillslope process zones and lower degrees of connectivity including Birch Creek, Cottonwood Creek, and San Juan Creek. 3.1.5. Downstream patterns The sequence of process zones that occurs within the studied basins was quantified by recording the frequency at which selected process zone types are linked to one another at the downstream end. The data reveal that the downstream pattern of process zone types is similar between the basins. The most commonly observed sequence is characterized by the flow of unincised and incised hollows into unfilled valleys that are linked to either side-valley fans or incised alluvial valleys. Further downbasin, incised alluvial valleys commonly alternate with incised colluvial valleys associated with large side-valley fans that extend across the valley floor (Fig. 10). The pattern is characterized by the splitting and rejoining of the channel around vegetated islands forming a multichannel system. Formation of individual anabranches is typically related to channel aggradation and filling upstream of woody debris dams, followed by overbank flooding, floodplain erosion, and channel avulsion. The net product is a highly dynamic valley floor characterized by localized cutting and filling that creates multiple, discontinuous terraces with terrace scarps on the order of 1 to 2 m or less in height (Germanoski and Miller, 2004). This process zone type is characterized by minimal sediment storage in floodplains or terraces within the valley as the channel bed is frequently composed of bedrock, and floodplains and terraces consist of thin (b2.5 m) cobble-sized alluvium that can apparently be mobilized by floods having a recurrence interval measured in years (Germanoski and Miller, 2004). 3.1.6. Geological controls on process zones In order to quantify bedrock–process zone relations, the rock type underlying each of the delineated process zones within the Kingston Canyon, Birch Creek, Cottonwood Creek, and San Juan Creek basins was determined using existing geologic maps (the other basins are underlain exclusively by volcanic rocks and thus were not included in this analysis; Fig. 11). The collected data are presented in Fig. 12 in terms of the percent to which a given process zone is associated with a specific rock type. However, the basins differ in the degree to which they are underlain by a given geological unit. Thus, the normalized percentage for which a specific process zone is associated with a given lithology is also provided. The normalized values are based on the percent of the basin that the rock type underlies. Fig. 12A demonstrates that there is a preference for bedrock hollows, bedrock valleys, and unfilled valleys in Kingston Canyon to form on carbonate rocks and, to a lesser degree, siliciclastic sedimentary rocks. There also appears to be a preference for unincised hollows to form on siliciclastic 3.1.4. Side-valley fans Accumulations of alluvial and colluvial sediments at the mouth of tributary channels are common within the upland watersheds of central Nevada (Fig. 1). The size (length, area), gradient, and composition of these side-valley fans is highly variable and, as shown by Germanoski and Miller (2004), is dependent on basin size and shape as well as tributary size, relief, and gradient. These side-valley fans can be subdivided into two types: (i) those in which the tributary channel traverses the length of the fan and joins the axial channel Table 5 Summary of meadow, process zone and connectivity characteristics of studied basins; groupings are modified from Germanoski and Miller (2004). Unincised-toincised ratio # and area of meadows (km2) % Meadowto-basin area Character of lower-most reacha 90 0 2/0.02 0.06 66 86 0.06 2/0.02 0.04 98 61 85 0.02 1/0.06 0.07 0.21 0.13 0.82 100 74 81 14 9 234 100 89 55 0 0 0.15 2/0.05 5/0.07 5/0.1 0.62 1.00 0.22 ANBV in bedrock controlled canyon ANBV in bedrock controlled canyon IAV; >4 m incision at mouth IAV IAV IAV in narrow bedrock canyon 1.06 88 32 28 0.10 1/0.02 0.10 0.76 75 49 35 0.07 5/0.18 0.30 1.02 87 42 60 0.18 2/0.09 0.33 Category Basins Dominant bedrock Storage-totrans. ratio Group 1 Flood-dominated Pine Cr. Volcanic 0.52 99 29 South Twin R. Volcanic 0.56 95 Barley Cr. Volcanic 0.43 N. Corcoran S. Corcoran Birch Cr. Volcanic Volcanic Siliciclastics Carbonates Intrusives Other Metamorphic Siliciclastics Carbonates Volcanic Group 2 Deeply incised channels Group 3 Fan-dominated Cottonwood Cr. Kingston C. San Juan Cr. a IAV — Incised alluvial valley; ANBV — anabranching valley. % basin connected # of fans % fans incised IAV; incised several meters at mouth IAV in narrow bedrock canyon IAV; incised several meters at mouth J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 Unincised hollow Bedrock valley Unfilled valley Incised Unincised colluvial alluvial valley valley Unincised fan Unincised hollow Unincised hollow Unincised hollow Incised fan Bedrock hollow Wet Meadow Incised alluvial valley 395 Incised alluvial valley Incised colluvial valley Unfilled valley Incised alluvial valley Incised alluvial valley Unincised fan Unfilled valley Unfilled valley Fig. 10. Schematic diagram of the most common downstream patterns in process zones observed within the studied basins. Diagram based on statistical analysis of downstream linkages. rocks. In contrast, a relatively high number of unincised hollows and unfilled valleys are located on volcanic rocks within the Birch Creek basin in spite of the fact that volcanic assemblages underlie less than about 1% of its area. Comparison of the actual to normalized data (Fig. 12D) suggests that these process zones are preferentially located on volcanic rocks within the San Juan Creek basin as well. Geology also can influence basin dynamics by controlling the size of the alluvial/colluvial sediment found within the basin. An examination of more than 9700 clasts in seven of the basins (all but the South Twin River) reveals that clasts from highly fractured metamorphic (metasedimentary) and siliciclastic rocks are generally finer when all process zones are considered, whereas clasts from quartzite and intrusive rocks tend to be larger (Fig. 13). The size of carbonate clasts varies between basins but tends to be relatively large in Kingston Canyon and Birch Creek, where carbonate rocks are relatively abundant, and fine-grained where carbonate rocks are rare. The size of volcanic particles also varies widely between the basins, but tend to be of relatively moderate size. Fan-dominated basins Deeply-incised basins 3.2. Connectivity Geomorphic connectivity was operationally defined by the existence of a continuous channel within a given process zone that allows for the channelized transport of water and sediment through the reach. Fig. 14A was produced by measuring the relative proportion of the drainage area that was connected (and disconnected) upstream of all major tributary confluences within the basin. The figure illustrates that (i) the degree of connectivity measured at the basin mouth varies significantly (from ~74 to 100%) between the eight studied watersheds and (ii) the way in which connectivity changes downvalley varies between the basins. The Pine Creek, South Twin River, Barley Creek, and North Corcoran Creek basins, which are underlain by volcanic rocks, exhibit the highest degree of connectivity (exceeding 95% throughout the basin). Cottonwood Creek possesses a fully connected drainage network in the upstream areas of the basin, but significant areas are not integrated with the axial channel downstream. As a result, about 15% of the basin is disconnected with the axial channel at the basin mouth. Flood-dominated basins Fig. 11. Bedrock composition of studied basins (modified and updated from Germanoski and Miller, 2004). 396 J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 Fig. 12. Proportion of process zones underlain by a given rock type in Kingston Canyon (A), Birch Creek (B) Cottonwood Creek (C), and San Juan Creek (D). Graphs on right show relative frequency of zone type normalized according to percent of basin area underlain by given lithology and illustrates tendency of process zone type to be associated with specific rock type. The loss of connectivity begins in the upstream-most areas of the Kingston Canyon, Birch Creek, San Juan Creek, and South Corcoran Creek basins and semisystematically declines before acquiring a variable but moderate (~10–20%) or high (20–30%) degree of disconnectivity throughout the middle and lower portions of the basins. At the basin mouth, about 15 to 25% of the upstream basin area in these watersheds does not contribute water or sediment to the axial drainage via a channel. Inspection of the process zone maps (e.g., Fig. 4) reveals that connectivity of the drainage network is primarily disrupted by two types of process zones: unincised side-valley fans and unincised valley floors. J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 Kingston Canyon (n= 2500) Cottonwood Creek 397 Birch Creek (n= 1550) San Juan Creek (n= 1400) (n= 1700) Other Basins Total Clasts (n= 9700) (n= 1350) (n= 750) (n= 850) Fig. 13. Basic statistics for intermediate particle diameter measured within all process zones within the 7 basins for which data are available. Data stratified by clast lithology where multiple lithologies exist (i.e., within Kingston Canyon, Birch Creek, Cottonwood Creek, and San Juan Creek). Graph for total clasts represent statistics stratified by lithology for all clasts examined within the studied basins. Filled rectangle — mean; box — standard deviation; line across box — median intermediate diameter; whiskers — 10 and 90% exceedence values; and n represents number of clasts used in statistical analysis. 4. Discussion 4.1. Basin sensitivity and dynamics An objective of this investigation was to expand on the work of Germanoski and Miller (2004) to gain a more detailed understanding of landform sensitivity and dynamics in upland watersheds in central Nevada. The hierarchical approach used here was particularly aimed at quantifying spatial variations in channel/valley form and process observed in the earlier analysis. Upstream reaches of Kingston Canyon, for example, are clearly affected by side-valley fans; whereas side-valley fans are truncated by a deeply incised channel near the basin mouth. As a result, classification of the entire watershed into a single sensitivity type proved problematic, prompting Germanoski and Miller (2004) to subdivide Kingston Canyon into two distinct sensitivity types (fan-dominated and deeply incised, respectively). 398 J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 A B % Connected 120 100 80 60 40 y = -133.85x + 97.59 R² = 0.47 20 0 0 0.05 0.1 0.15 0.2 Area of meadows (km2) Fig. 14. (A) Downstream changes in percent of drainage connected by surface channels. Symbols represent individual meadows within the studied watersheds; (B) relation between basin connectivity and total meadow area (sum of all meadow areas) within the 9 studied basins. Relationship is statistically significant. They hypothesized that a hierarchical approach would largely eliminate the problems of classifying basin sensitivity imposed by spatial variations in process and form. At the watershed scale, the hierarchical approach allowed the eight investigated basins (counting north and south Corcoran Canyon as a single catchment) to be subdivided into three broad groups in terms of landscape sensitivity to change and basin dynamics. These groups are consistent with the groups described by Germanoski and Miller (2004) as flood-dominated channels (group 1), deeply incised channels (group 2), and fan-dominated channels (group 3) (Table 5). Two basins, San Juan Creek and Birch Creek, categorized by Germanoski and Miller (2004) as deeply incised and pseudostable channels, respectively (Table 1), are categorized as fan-dominated channels using the hierarchical approach in this analysis. Reclassification of the two basins reflects (i) a focus by Germanoski and Miller (2004) on downstream segments of the riparian corridor where a majority of meadow complexes are found and (ii) a more detailed analysis of reach-scale processes during this study and the spatial variations in valley/channel form and process throughout the entire watershed. We believe that the hierarchical approach used here is more effective from a management perspective because it not only considers the general (average) form and process rates of the riparian corridor, but classifies basins according to the type and structural arrangement (frequency, density, spatial pattern) of process zones that are present within any given part of the basin. Understanding of the process zone types and structure subsequently allows for a more effective prediction of such processes as channel incision, avulsion, and the preservation of wet meadow ecosystems along any given segment of the riparian corridor. Moreover, the process zone approach provides important insights into the spatial and temporal variability of runoff and recharge processes along the drainage network, as will be described in more detail elsewhere. It is important to note, however, that the work conducted by Germanoski and Miller (2004) showed that clear differences in channel incision and watershed-scale parameters exist between the studied drainages that can be related to ecological processes (Chambers et al., 2004) and that these differences are recognized in the analysis performed here. The most dynamic basins studied in central Nevada, referred to as ‘flood-dominated’ basins (Table 5), include Pine Creek and the South Twin River. Both basins possess an anabranching channel pattern within the lower reaches of the watershed. The pattern is characterized by high rates of sediment transport that allows for localized zones of deposition and erosion behind debris dams and for the rapid change of the valley floor through the production and erosion of low-relief, discontinuous terraces. The alluvial sediments are transported over a shallow bedrock surface and through a bedrock canyon that forms the local hillslopes. Meadow complexes are absent within the anabranching channel reach and are extremely limited within other parts of the basin (Table 5). The basins are underlain exclusively by volcanic rocks (Fig. 11) that generate abundant, relatively large clasts (mean intermediate diameter is >50 mm) (Fig. 13). On hillslopes, these sediments are primarily stored within unincised hollows, which occur with moderate frequencies and densities (Fig. 7). At the base of the hillslopes, side-valley fans occur with a frequency similar to that within the other watersheds (with the exception of Birch Creek); however, nearly all of these are located in headwater areas and are incised (Table 5). Within reaches characterized by anabranching channels, incision is limited by shallow bedrock. Upstream in more headwater areas valleys are filled with alluvial and colluvial deposits. Nearly all of these valleys have been incised creating incised alluvial and incised colluvial valley reaches that on a basin scale are present at relatively low densities and frequencies (Fig. 8). Unincised valleys, which limit the connectivity between hillslope and axial channel areas, are limited in number and length. Upstream of the anabranching channels, then, the riparian corridor is prone to rapid and pervasive incision associated with a dynamic channel. From a process perspective, hillslope process zones can be grouped into reaches that (i) have a tendency to store sediment and allow for infiltration and groundwater recharge (unincised hollows), and (ii) lack extensive unconsolidated deposits, but can effectively transport water and sediment through the reach to downslope areas (including incised hollows, bedrock hollows, unfilled valleys, and bedrock valleys). Fig. 15A shows that the ratio of zone storage-totransport (per unit basin area) is below 1 for Pine Creek and the South Twin River and moderately low in comparison to the other studied basins. Thus, the nature of the upper drainage network is structurally organized to transport relatively large amounts of sediment temporarily stored in unincised hollows downstream, across incised alluvial fans, to the axial channel. It is then transported through the incised alluvial and colluvial channel segments to the downstream anabranching channel reaches. These anabranching reaches occur within relatively steep, narrow bedrock influenced valleys that prevent the storage of large volumes of sediment. Thus, the dynamic nature of the anabranching channels in the lower reaches of the watershed appears to be related to (i) an abundance of sediment that can be effectively transported off the hillslopes by significant runoff and (ii) a high degree of connectivity in the basin (>95%, Fig. 14A) that allows for its continued movement toward the basin mouth. The high degree of connectivity may also enhance the discharge of channelized flows that promote incision of the side-valley fans and valleys filled with alluvial and colluvial sediments. The net result of the basins' architecture is a highly dynamic channel system in downstream basin areas and upstream valley segments that respond relatively quickly to changing environmental conditions by means of channel incision. As is the case for Pine Creek and the South Twin River, the Barley Creek and Corcoran Canyon basins have deeply incised channels (Table 5) and are underlain exclusively by volcanic rocks (Fig. 11). However, the volcanic units found in Pine Creek and the South Twin River basin differ from those found in Barley Creek and Corcoran J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 A B Fan-dominated basins Deeply-incised Flood-dominated basins basins Fig. 15. (A) Ratio of density of storage versus transport-dominated process zones located on hillslopes. Lower ratios indicate tendency to transport sediment and water off hillslopes; (B) ratio of the total length of unincised-to-incised (incised) process zones within the studied basins. Lower ratios associated with highly connected drainage network and higher proportion of incised channels. Canyon in that they generate smaller clasts (mean intermediate diameter is b50 mm) (Fig. 13). The rocks also produce large quantities of sand/grus-sized sediment (Barley Creek) and finer material from poorly welded tuffs (Corcoran Canyon) as evidenced from thick alluvial deposits exposed along the incised alluvial valleys. Both the frequency and density of unincised hollows are relatively low (Fig. 7), suggesting that the amount of sediment stored on the hillslopes is limited. However, the frequency and density of process zones that are conducive to water and sediment transport are relatively high in comparison to the other watersheds. As a result, the sediment storage-to-transport ratio is lower than for the other basin types (Fig. 15A), suggesting that sediment can be effectively transported off the hillslopes. In contrast to Pine Creek and the South Twin River, the Barley Creek basin is characterized by the highest density of incised alluvial valleys (Fig. 8). Anabranching channels do not occur. Not only are incised alluvial valleys common, but the depth of incision is (at least locally) high, exceeding 5 m at the basin mouth within Barley Creek. The incised alluvial valleys within the north and south Corcoran basins are less entrenched, although the axial valleys are frequently characterized by large, upstream-migrating headcuts exceeding a few meters in height. Meadow complexes occur along the riparian corridor within Barley Creek and Corcoran Canyon, but they have been severely degraded by channel/gully incision. In the case of Barley Creek, the meadow near the basin mouth is preserved only along the valley margins and represents a local zone of groundwater discharge distant from the deeply incised axial channel. Five meadows exist in south Corcoran (Table 5), but each is being dissected by numerous gullies with migrating headcuts that began on narrower and steeper parts 399 of the valley floor (Miller et al., 2011). The fine-grained sediments underlying the meadows prevented a pervasive drop in the water table caused by incision, permitting persistence of meadows (Lord et al., 2011). In Barley Creek and Corcoran Canyon, the dissection has resulted in relatively small meadows (generally b0.02 km 2). Connectivity within these basins is high, exceeding 95% (Table 5). Thus, once reaching the base of the hillslopes, sediment can be transported through the side-valley fans (nearly all of which are incised, Table 5) and through the incised alluvial and colluvial valleys before exiting the basin. It is not entirely clear why these basins lack anabranching channel reaches, but it may be related to the more deeply buried bedrock near the basin mouth, lower valley side relief along the axial drainage, and lower availability of stored sediment on the hillslopes. Given the similarities in the underlying bedrock and process zone type and structure on the hillslopes, the degree of incision along the lower valleys in Corcoran Canyon might be expected to be significantly greater, approaching that observed along Barley Creek. It appears, however, that incision was limited within both the north and south Corcoran basins by (i) their relatively small size (b10 km2), (ii) the relatively low elevation of the drainage divides that limits the accumulation of snow and spring runoff and (iii) their relatively low relief (Table 2). The remaining basins are fan-dominated (Table 5) and include Kingston Canyon, Cottonwood Creek, San Juan Creek, and Birch Creek. They exhibit a number of similarities at both the basin and reach scale. These similarities include (i) a high frequency and density of unincised hollows in comparison to the other studied basins (Fig. 7); (ii) high hillslope storage-to-transport ratios (Fig. 15A); (iii) a relatively high percentage of unincised fans (Table 5); (iv) a relatively high frequency and density of unincised valleys leading to a high unincised valley-to-incised valley length ratio (Fig. 15B); (v) large areas of the basin that are disconnected from the axial drainage (Fig. 14A); (vi) a loss of connectivity that typically begins in the upstream-most areas of the basin, decreases to the middle of the basin, and subsequently remains constant or decreases downstream; (vii) an alternating pattern of incised alluvial valleys with incised colluvial valleys along the riparian corridor within the upper reaches of the basin (e.g., shown on Fig. 4). The incised colluvial valleys are typically associated with large side-valley fans that project across the valley floor and that create a ‘stepped’ longitudinal channel and valley profile (as discussed in the introduction); (viii) channels that are incised into the valley floor within the lower reaches of the basin; (ix) relatively large (>0.04 km 2) wet meadow complexes, often located with the upper or middle segments of the basin (Table 5); and (x) a variety of underlying bedrock types, including sedimentary, metamorphic, and intrusive rocks (Fig. 11). The nature of the incised alluvial and colluvial valleys within upstream basin areas indicates that these parts of the basin have a high potential for axial channel incision and meadow degradation centered on the removal of the ‘steps’ that occur along the incised colluvial valleys. However, the combined character of the hillslope process zones, the underlying rock types, and the high degree of basin area that is disconnected from the axial valley limits the rate and magnitude of incision that can and has occurred. Immediately upstream of the basin mouth, an increase in basin area and discharge has led to deeper channel incision (e.g., shown for Kingston Canyon, Fig. 9), and the steppedlongitudinal profile of the channel has largely been replaced by a more continuously sloping channel bed. In addition, downstream meadows have been replaced by drier plant communities and are significantly degraded. These downstream areas have a lower potential for further incision, but are more dynamic in terms of the potential rate of incision following an environmental disturbance. 4.2. Geological controls on process zone structure and connectivity Data presented above suggest that the nature and behavior of the drainage system along axial valleys depends in part on the type and J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 spatial structure of the process zones on the hillslopes and the degree of connectivity within the watershed. It is therefore important to understand the factors that control both connectivity and the nature of the process zones on the hillslopes. The formation of unincised/incised hollows appears to have two requirements. First, the resistance of the underlying bedrock to erosion must be such that it allows for the development of U-shaped depressions oriented semiparallel to the hillslope. Second, erosion of the bedrock must produce an abundant supply of relatively small particles (a median clast diameter of ~20 mm) that accumulate within the depression. Fig. 12A shows that within Kingston Canyon these two factors are met most often by siliciclastic rocks and, to a lesser degree, carbonates. Within the Birch and San Juan Creek basins, hillslope hollows have a distinct tendency to form on volcanic rocks that weather into easily eroded, relatively small clasts. In contrast, bedrock hollows and bedrock valleys are favored by resistant rock units that produce limited, but larger clasts and steep (locally vertical), jagged outcrops (Fig. 5C,D). Fig. 12A shows that within Kingston Canyon these characteristics are predominantly met by carbonate lithologies. The fact that unincised hollows and bedrock hollows and valleys form on carbonate and volcanic rocks indicates that the lithology of the rock units may not be as important in controlling the type of process zone that is present as the weathering characteristics of the materials. In the case of carbonates, some units are highly fractured, allowing them to be more easily eroded and to produce an abundance of relatively small clasts. These clasts are frequently transported to and deposited along downstream incised alluvial valleys where they locally comprise as much as 70% of the >2.8 mm sediment fraction. Other carbonate rocks are poorly fractured and break into large clasts and boulders (Fig. 5C). Differences in weathering characteristics, and their influence on the distribution and type of process zone present on the hillslopes, are also apparent for metamorphic, intrusive, and volcanic rocks. Bedrock hollows and valleys are present on these rock types where they are locally more resistant to erosion (Fig. 12). In most areas, however, these rock units are relatively easy to erode and, as shown by the normalized data in Fig. 12, tend not to produce bedrock hollows and valleys. Unfilled valleys also form preferentially in easily eroded rock types. In fact, unincised hollows are most often linked downstream with unfilled valleys (Fig. 10), and both are rarely associated with bedrock hollows and valleys. In other words, unincised/incised hollows and unfilled valleys commonly form a couplet that are most frequently found on easily eroded rock types. Where rocks are more resistant, bedrock hollows and valleys occur. The net result, then, is that there is a tendency at the basin scale for the density of unincised/incised hollows and unfilled valleys to be inversely related to bedrock hollows and valleys (Fig. 7). The downstream alteration of unincised hollows to unfilled valleys is presumably driven by an increase in contributing drainage area to the drainage net that leads to particle movement. More specifically, as contributing area and presumably runoff increases, a threshold is reached at which the generated particles can be entrained and transported downslope. The consistent ability to entrain and transport sediment leads to the development of V-shaped, sediment-limited valleys in comparison to those found upslope. Interestingly, the contributing area required to produce unfilled valleys varies between rock types (Fig. 16A), indicating that bedrock influences runoff. Volcanic rocks require the lowest contributing area to produce an unfilled valley, whereas siliciclastic and carbonate rocks tend to require the most (5 to 7 times more than for volcanic rocks). The implication is that higher rates of runoff are associated with volcanic lithologies. These data are consistent with three additional observations. First, unincised fans and unincised valleys are most often associated with upstream contributing areas underlain by carbonates and, to a lesser degree, siliciclastics. Thus, these rock types tend to generate basins characterized by a large degree of A 0.16 0.14 Contributing area (km2) 400 0.12 0.10 0.08 0.06 0.04 0.02 0.00 B Fig. 16. (A) Mean upstream area contributing flow to head of unfilled valley stratified by rock type. Low values indicate that smaller areas are required to transport sediment downslope, presumably as a result of higher runoff; (B) annual flood hydrograph developed for Kingston Canyon and San Juan Creek by Amacher et al. (2004). Hydrographs indicate a tendency for higher peak flows and more rapid runoff (shorter basin lag times) for basins underlain by volcanic rocks. Increased runoff from volcanic basins is consistent with lower sediment storage to transport ratios, a higher percentage of incised side-valley fans, a lower unincised-to-incised valley ratio, and a higher degree of basin connectivity. disconnected drainage. Second, basins underlain by volcanic rocks tend to exhibit more extensively incised fans and axial valley areas. Finally, hydrographs presented by Germanoski and Miller (2004) showed that annual peak discharges tend to be higher and peak more rapidly in San Juan Creek, underlain predominately by volcanic rocks, than in Kingston Creek, underlain by a large percentage of carbonates and highly fractured metasedimentary rocks (Fig. 16B). While rock type appears to influence the nature of the process zones on the hillslopes, the type and distribution of process zones that are present may also influence connectivity. Unincised hollows exhibited few signs of surface runoff within subbasins of Kingston Canyon and Birch Creek hydrologically monitored during a threeyear period. This was not unexpected given the highly porous nature of the unconsolidated sediment within the depressions. Thus, a high density of unincised hollows would be expected to enhance groundwater recharge and reduce runoff and, therefore, downstream incision of alluvial and colluvial deposits. This suggestion is supported by the relatively high frequencies and densities of unincised hollows (Fig. 7) found in basins with a high percentage of unincised fans (Table 5), unincised valleys (Fig. 8), and disconnected drainage (Table 5, Fig. 14A). 4.3. Factors influencing meadow distribution A total of 56 meadow complexes have been identified in upland watersheds of central Nevada. The geomorphology, stratigraphy, J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 and hydrology of many of these meadows have been extensively studied using a variety of methods, ranging from the coring and description of the valley deposits, to the evaluation of groundwater data collected from extensive arrays of monitoring wells and piezometers, to the application of ground penetrating radar and seismic surveys (Jewett et al., 2004; Lord et al., 2011). These analyses have demonstrated that each meadow complex is a unique feature and cannot be unequivocally described by any single model. Nonetheless, nearly all of the meadows result from a combination of factors, including an adequate supply of water to the reach and a mechanism that retards the downvalley movement of groundwater through the valley fill and causes groundwater levels to rise to the valley surface. A reduction in groundwater flow may be produced in several ways, the most prominent of which are the vertical and/or lateral constriction of the valley fill by bedrock and a change in grain size of the valley-fill deposits. Reductions in grain size and the low permeability of the resulting deposits are commonly associated with deposition upstream of side-valley fans and bedrock protrusions (Germanoski et al., 2011). Thus, the majority of meadows are located upstream of large side-valley fans that project across the valley floor (Fig. 1). In addition, most of the meadows exhibit relatively impermeable finegrained sediment near the ground surface in comparison to deeper deposits. These fine-grained surface materials (i) retard the ability of water to rapidly discharge and drain the underlying aquifer and (ii) create artesian flow conditions and springs that allow meadows to remain saturated over long periods of time (Lord et al., 2011). The above data show that the location of meadow development is dependent in part on both watershed scale and local factors (Germanoski et al., 2011). Meadows are unlikely, for example, to form where rapidly evolving anabranching valleys occur as they possess neither the fine-grained sediment required for meadow formation nor the constrictions required to retard groundwater flow and produce the necessary upward flow gradients. Meadows are scarce in deeply incised alluvial valleys, such as is within the Barley Creek basin, because the entrenched channel typically allows for direct hydrologic connection between the channel and permeable subsurface units, rapid drainage of the shallow aquifer system, and lowering of the groundwater table. Unlike flood-dominated basins with anabranching channels, some deeply incised alluvial valleys supported meadow complexes prior to incision. With incision, the meadows that existed became lost or highly degraded and typically occur as small features along the valley margins fed by groundwater that is poorly integrated with the entrenched channel (Lord et al., 2011). Earlier we argued that the process zones that comprise the lower axial valleys reflect the nature of the underlying bedrock and the type and structure of the process zones on the hillslopes. It follows, then, that meadow formation and distribution should also reflect the type and structure of process zones within the basin. Such relations do exist. For example, 78% (18 of 23) of the meadows within the studied basins are found along unincised valleys or incised alluvial valley floors characterized by shallow depths of incision, and which possess upstream drainage areas exhibiting a relatively low degree of connectivity (b90%). In fact, >20% of the upstream basin area is disconnected from the axial drainage network for a majority of the meadows. At the basin scale, 47% of the variation in total meadow area can be accounted for by the degree of basin connectivity within the watershed (Fig. 14B), a high percentage given the local requirements for meadow formation. The relationship between connectivity and meadow formation and area is related to several factors. First, meadow development is promoted by groundwater recharge and enhanced groundwater flow. Clearly, groundwater recharge is promoted by the movement of water from a tributary basin across the surface of an unincised fan or unincised valley reach. In addition, basins with numerous unincised fans, unincised valleys, and a relatively high degree of disconnected basin area also exhibit a high frequency and density of 401 unincised hollows. The broad U-shaped nature of the hollows combined with their loose, highly permeable accumulations of sediment are likely to increase the amount of upland recharge. In fact, we argue that the relatively high magnitude of recharge that occurs within areas characterized by numerous unincised hollows reduces the ability of surface flows to erode and incise downstream, side-valley fan deposits. Second, channel incision, particularly across side-valley fans, is less severe along axial valleys within basins characterized by a relatively low degree of connectivity (e.g., Kingston Canyon, Cottonwood Creek, Birch Creek, and San Juan Creek). Thus, incision of the meadow deposits and degradation of the meadow complexes are less likely. These observations are significant in that they suggest that any activity within the upland areas that promotes an increase in connectivity within the basin may negatively affect meadow complexes, in spite of the fact that the meadows are spatially separated from the disturbance. 5. Summary and conclusions The drainage network within the studied watersheds of central Nevada was subdivided into distinct process zones on the basis of valley form, the inherent geological deposits, and the presence or absence of surficial channels. Each type of process zone is dominated by a suite of geomorphic and hydrologic processes that produce the morphologic characteristics upon which it is classified. U-shaped, unincised hollows, for example, are characterized by sediment deposition and storage as well as groundwater recharge; whereas Vshaped, bedrock-floored unfilled valleys are dominated by sediment transport and runoff. On hillslopes, the type and spatial structure of the process zones are related to the lithology and weathering characteristics of the underlying bedrock. Unincised hollows and unfilled valleys, to which the unincised hollows are frequently linked, are associated with less resistant rock types (including siliciclastic, volcanic, and some carbonate formations) that weather into abundant but relatively small particles. Bedrock valleys and hollows are associated with resistant, poorly fractured rocks that produce limited but larger clasts. The type and spatial arrangement of process zones within lower elevation (axial) valleys is dependent on the characteristics of the process zones on the hillslopes. A high frequency and density of unincised hollows are associated with numerous unincised sidevalley fans as well as frequent and lengthy unincised valley segments that disconnect large sections of the drainage basin from channelized surface flows. In contrast, a relatively high density of transportdominated process zones (including unfilled valleys, bedrock hollows, and bedrock valleys) leads downstream to the incision of side-valley fans and axial valley deposits. Connectivity is related to the lithology of the underlying bedrock, with higher degrees of connectivity being associated with volcanic rocks that presumably yield high rates of runoff. Low levels of connectivity are associated most frequently with sedimentary and metamorphic assemblages, particularly carbonates that are locally permeable where they are extensively fractured. At the basin scale, watersheds can be separated broadly into three general groups (flood-dominated, deeply incised, and fandominated) on the basis of the underlying bedrock geology, the ratio of sediment storage to transport-dominated process zones, the degree of connectivity, and the severity of axial valley incision. Meadow complexes are most extensive within basins characterized by a high ratio of sediment storage to transport-dominated process zones on hillslopes and by a relatively low degree of connectivity in comparison to the other studied watersheds. The more extensive nature of meadows in these basins is thought to be related to (i) lower runoff from sedimentary, intrusive, and metamorphic rocks in comparison to volcanic terrains that reduces the severity of incision along the riparian corridor; (ii) enhanced groundwater recharge in numerous and lengthy unfilled hollows on the hillslopes; and 402 J.R. Miller et al. / Geomorphology 139–140 (2012) 384–402 (iii) enhanced groundwater recharge upon unincised side-valley fans and unincised valleys. From a management perspective, basins underlain by volcanic rocks appear to be more sensitive to incision and produce more dynamic channels in terms of the rate of channel/valley modification than those underlain by other lithologies. Where meadow complexes exist in these basins, they are likely to be severely degraded and will be difficult to manage given the dynamic nature of the axial channel processes. Meadows within basins that exhibit a low degree of connectivity and high sediment storage-to-transport ratios on hillslopes will likely be more responsive to management activities because of the reduced susceptibility to channel incision and because of a large supply of groundwater flow to the meadow through an extensive network of recharge sites. 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