United States Department of Agriculture, Forest Service Pacific Southwest Research Station Sierra Nevada Research Center 1000 Years of Forest History in the Glass Creek Watershed, Eastern Sierra Nevada: Interpreting the influence of fire, climatic change, and environmental change on subalpine forest structure and composition The Research: Much progress has been made in the past two decades in elucidating the key role that recurring wildfire plays in determining forest structure and composition. While other forces such as climate change and landscape evolution obviously influence vegetation over longer terms, their relative importance as co-architects at the management-timescale of decades to centuries remains little investigated. We are studying forest history in a midelevation watershed in the eastern Sierra Nevada, the Glass Creek watershed. The forest extends from lodgepole pine at low elevations, old-growth eastside mixed conifer with red fir at middle elevations, to whitebark pine at treeline. A large wet meadow with high species diversity dominates the valley floor. Aside from domestic sheep grazing in the meadow, the watershed has been minimally disturbed by recent human activities; even fire suppression does not seem to have influenced fire regimes. The watershed has been significantly influenced, however, by repeat eruptions of the nearby Inyo Craters, whose most recent activities covered the watershed with many meters of tephra ~750 years ago. In related studies, we have been documenting the influence of climate on vegetation at nearby treeline elevations and the influence of climate on conifer invasion of the meadow. In this study, we investigate the main forest zones of this watershed as a case study in dissecting the role of climate, fire, and volcanism in shaping stand structure and composition. Using dendrochronological, ecological plot, and fire-scar analyses in the forest zones, we are reconstructing forest structure, climate relations, and fire history over the last 1000 years. We find evidence for influence of each of the forces (fire, climate, and eruption) on forest dynamics, with relative importance changing over time. In some cases, the influences are confounding, in that they have similar effects ecologically, and are thus difficult to separate. Important conclusions emerging from these studies include 1) the role of each force is significant and vegetation diversity cannot be adequately understood with reference to single one, 2) extreme events and variability in events determine significant ecological pathways that become set for hundreds of years, and 3) third order change is evident, that is - vegetation changes are evident in response to physical systems that are themselves changing. Our Mission: Sierra Nevada Ecosystems are complex and our knowledge of them is incomplete. As a result, the long term outcome of any given land and resource management strategy is uncertain. We will provide assistance to land managers and policy makers by addressing this management dilemma through targeted research, emphasizing an integrated, ecoregional approach to examine particular physical, ecological, and socioeconomic issues, across a range of appropriate spatial and temporal scales specific to each issue. This unit will represent the collective research expertise and interests of scientists located in Fresno, Davis and Albany as well as other scientists within the Pacific Southwest Research Station. With a full spectrum of research, from long term, fundamental research to short-term, tactical applications, this Center is intended to support conservation, restoration, and sustainable utilization of the lands within the Sierra Nevada ecoregion. Objectives: Evaluate the relative roles of fire, climate change, and volcanic eruptions as architects of forest structure and composition over the past 1000 years. Application of Research Results: This study is part of a complex of studies we are conducting in the Glass Creek watershed designed to provide information for resource managers about the relative influence of climate, fire, and volcanism on forest dynamics. Under conditions of changing climates, and altered fire regimes, decision-makers need accurate information about how forests respond to these forces. The Glass Creek Watershed is part of a larger landscape analysis by the Inyo National Forest, and management decisions will benefit from information provided in the suite of studies we are undertaking there. Location: Eastern Sierra Nevada: Glass Creek watershed, headwaters of the Owens River. Publications: PDFs available at: http://www.fs.fed.us/psw/programs/snrc/staff/millar • Poster: Millar, CI; Westfall, RD; Delany, DL; King, JC; and Alden, HA. 2004. HighElevation Response of Conifers to Climate Change in the Sierra Nevada and Western Great Basin, USA: Treeline Elevation is Not the Primary Effect. Poster: Millar, CI; Westfall, RD; Delany, DL; King, JC; and Alden, HA. 2004. Climate as an Ecosystem Architect; Responses of High-Elevation Conifers to Past Climate Variability. http://www.fs.fed.us/psw/programs/snrc USDA Forest Service Pacific Southwest Research Station Sierra Nevada Research Center Albany Location: 800 Buchanan Street Albany, CA 94710 For further information contact: Dr. Connie Millar cmillar@fs.fed.us Mailing Address: P.O. Box 245 Berkeley, CA 94701 Dr. Peter Stine, Program Manager Phone: 510-559-6300 Fax: 510-559-6440 Davis Location: 2121 Second Street Suite A101 Davis, CA 95616 Phone: 530-759-1700 Fax: 530-747-0241 Fresno Location: 2081 E. Sierra Avenue Fresno, CA 93710 Phone: 559-323-3200 Fax: 559-297-3355