THIS REPORT NOT TO BE QUOTED WITHOUT PRIOR REFERENCE TO THE COUNCIL *
International Council for the Exploration of the Sea C.M.1994/G:6
Demersal Fish Committee
This document is areport of a Study Group of the International Council for the Exploration of the
Sea and does not necessarily represent the views of the Council. Therefore, it should not be quoted without consultation with the General Secretary.
* General Secretary
ICES
Palaegade 2-4
DK - 1261 Copenhagen K
Denmark
TABLE OF CONTENTS
•
Section
1.
INTRODUCTION
1. 1 Terms of Reference . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
.
1.2
Members of Study Group .
2.
COMMON DATA BASE .
3.
SPECIES DISTRIBUTION
3.1
Data Used and Methods
3.2
Hake
3.3
Horse Mackerel
3.4
Mackerel
3.5
Lepidorhombus whiffiagonis
3.6
Lepidorhombus boscii
3.7
Lophius piscatorius
3.8
Lophius budegassa
4.
GEAR COMPARISON
5.
CO-ORDINATION OF BOTTOM TRAWL SURVEYS
.
.
.
.
.
.
.
.
.
.
.
6.
REFERENCES
TABLES 2-4
FIGURES 2-3.30
.
.
.
Page
4
5
6
3
4
2
3
3
1
1
2
3
3
3
•
•
1 INTRODUCTION
1.1 TERMS OF REFERENCE
At the 1993 ICES Statutory Meeting the Council decided (C.Res.
1993/2:32), that the Study
Group on Bottom Trawl Surveys in Sub-areas VI, VII and VIII and Division IXa (Chairman: Mr
J.C. POULARD, France) will work by correspondence in 1994 to : a) assemble data from different national research vessel surveys in a common data base; b) describe, on the basis of survey results, the seasonal distribution and relative abundance of hake, mackerei, horse mackerei, anglerfish (Lophius budegassa and L. piscatorius), and megrim (Lepidorhombus whiffiagonis and L. bosciJ) in the area; c) promote collaboration between nationallaboratories in co-ordinating boUom trawl surveys in the area.
1.2 MEMBERS OF THE STUDY GROUP
M. Azevedo
F. Borges
M de F. Cardador
A. Cascalho
J. Casey
P. Connolly
H.Dornheim
S. Ehrich
A. Eltink
M.J. de Figueiredo
B.W. Jones
M. Martins
F.J. Pereiro
G. Pestana
J.C. Poulard (Chairman)
F.Sanchez
M. Walsh .
Portugal
Portugal
Portugal
Portugal
United Kingdom
Ireland
Germany
Germany
Netherlands
Portugal
United Kingdom
Portugal
Spain
Portugal
France
Spain
United Kingdom
2 COMMON DATA BASE
No progress has been made in the definition and location of the data base. However, collating survey data using the IBTS data format exchange was undertaken. The table 2 gives the inventory of data collated up to now.
3 SPECIES DISTRIBUTION
3.1 DATA USED AND METHODS
Three sets of surveys results were extracted from the available data to describe the distribution of species during quarters, they are as folIows:
1
- the first quarter of 1990 : distribution of hake, horse mackerei, mackerel and megrim
(Lepidorhombus whiffiagonis) , in ICES divisions Via, Vllb and Vllj, from data provided by the
German survey (table 3);
- the fourth quarter of 1990 : distribution of hake, megrim (Lepidorhombus whiffiagonis and L.
bosciJ), anglerfish (Lophius piscatorius and L. budegassa) , to the south of 51
N, from data provided by the French, Spanish and Portuguese surveys; :
- the fourth quarter of 1992 : distribution of hake, horse mackerei, mackerei, megrim
(Lepidorhombus whiffiagonis) and anglerfish (Lophius piscatorius and L. budegassa) , to the north of 43° N, from data provided by Scottish and French surveys.
A detailed description of the survey design and methods has been given previously (ICES CM
1991/G :13). The main features of the surveys used are given in table 3. It must be noticed that several different gears were used according to the country during these surveys. Although a gear comparison was carried out in autumn 1992 (see § 4) between the bottom trawl type BAKA
44/60 used during Spanish surveys and the GOV 36/47 trawl used during French surveys, no attempt was made to standardize the gear in the absence of information about the performances of other gears. It must be noted that for hake (table 4) there is almost no difference between the fishing power of BAKA and GOV trawls, on the contrary BAKA trawl catches about three times as much megrim than GOV.
i
The distribution of the hauls through the surveyed area is showed on figure 3.1. There is no overlapping area between the different surveys, except a small one in 1992 between French and Scottish surveys around the 51° N (figure 3.1 Cl. Trawl stations completed in September
1990 by Spain and France and in September 1992 by France were incorporated into the data sets used.
Seven categories have been defined for the numbers of fish caught per hour fishing. Several scales were used, each of them was fitted to the abundance level of individual species to get the best description of its distribution.
The length frequency distributions are given when length data were available or the total number caught were large enough. They were obtained by pooling tow length compositions without applying any depth stratified weighting factors.
.
3.2
HAKE (FIGURES 3.2-3.7)
South of 51
N, the hake distribution shows a patchy pattern in autumn. Higher densities corresponded to O-group (Le. fish approximately less than 15 cm in length) which was generally caught at a depth comprised between 50 and 200 meters. The overall distribution of hake during the fourth quarter shows in fact the position of its major nursery grounds. The highest densities were recorded in Villa in 1990 and in VlIg,h in 1992.
3.3
HORSE MACKEREL (FIGURES 3.8-3.11)
North of 49° N, horse mackerel seems more abundant along the shelf edge during the first quarter than during the fourth one. In both quarters the 2+ group made up the bulk of the catches.
•
•
•
•
In the Bay of Biscay, higher densities were recorded at depth less than 100 m during the fourth quarter. The total catch can be split into four age groups (0 to 3+), the younger one being mainly found in shallow waters.
3.4
MACKEREL (FIGURES 3.12-3.15)
The highest catch numbers (more than ten thousands per hour) were found predominantly off the western coast of Ireland and south of Cornwall during the first quarter. High number distribution seems more even during the first quarter than during the fourth one along the shelf edge in the northern part of the area.
Densities were low in the Bay of Biscay compared to those recorded in the north.
3.5 Lepidorhombus whiffiagonis (FIGURES 3.16-3.20)
This species occurred mainly in two areas. The main one is centered on divisions VlIg,h,j and extended northwards to divisions Vllb and Via. In sub-area VIII densities were generally lower.
The second area was confined to division Vlllc with higher densities occurring mainly at a depth comprised between 100 and 200 meters. The length frequency distributions show no large difference over the surveyed area.
3.6 Lepidohombus boseii (FIGURES 3.21-3.22)
The distribution area of L.
boseii was Iimited to divisions Vllic and north of division IXa. There was an increasing trend in abundance from east to west in division Vlllc. The species occurred mainly in the 200-400 meters depth range . Outside of this area the abundance of very low and the species was not reported from surveys carried out north of 51 0 N.
L.
boseii was
There is some indication that BAKA trawl used by Spain is more efficient at catching both megrim species than the other gears used. This can affect their distribution pictures.
3.7 Lophius piseatorius (FIGURES 3.23-3.27)
The species was rather evenly distributed in northern area (divisions Via and Vllbj) during the fourth quarter of 1992.
L.
piseatorius was on the contrary scattered in division Vllh and sub-area
VIII during the fourth quarter of 1990 and 1992. The species was found fairly abundant in the eastern part of division Vlllc during the fourth quarter of 1990.
3.8 Lophius budegassa (FIGURES 3.28-3.30)
L.
budegassa abundance distribution showed high catch values in the southern part of division
Vllib and also in division VII/c. The species was scattered through the northern part of the surveyed area and was almost not found north of 55 0 N.
4 GEAR COMPARISON
A comparison between the bottom trawl type BAKA 44/60 used by the R.V Cornide de Saavedra
(Spain) and the GOV 36/47 used by the R.V Thalassa.(France) during surveys (Anon., 1991)
3
was carried out in the south of the Bay of Biscay between the 26 September and the 4 October
1992. Nineteen pairs of simultaneous tows lasting 30 minutes each were completed.
Preliminary results showed that for hake both gears have almost the same catch efficiency
(table 4), the length compositions being similar. However, Baka trawl caught somewhat three times as many megrim as the GOV trawl and less than twice as many anglerfish. The length distributions of these species were similar although there was some indication that the Baka trawl caught a higher proportion of small anglerfish and a smaller proportion of L. baseii.
5 CO-ORDINATION OF BOTTOM TRAWL SURVEYS
Not enough progress has been made in the co-ordination of bottom trawl surveys in the area.
However, it should be possible to get a good coverage of the northern part of the area (Le. from
VIa to Villa) during the first quarter by co-ordinating surveys carried out by Seotland, Ireland,
England and Germany. The whole area is already eovered during the fourth quarter beeause many national surveys are eonducted by Seotland, Ireland, Netherlands, Franee, Spain and
Portugal.
6 REFERENCES
Anon. 1991. Report of the study group on the co-ordination of bottom trawl surveys in Subareas VI, VII and VIII and Division IXa. ICES, Doe. C.M. 1991/G: 13.
•
•
4
Table 2 Bottom trawl surveys in ICES Sub-areas VI, VII, VIII and division IXa : summary of survey data collated.
Quarter
Countrv
France
Germany
Year
1987
1988
1989
1990
1991
1992
I
1987 Vla,Vllb,j,e,Vllla
1989 Vla,Vllb,j,e,Vllla
1990.
Vla,Vllb,j,e
1991 Vla,Vllb,i,e
11
Vllla-d
Vlle-j, Vllla-d
111 IV
Vllla-d
Vllla-d
Vllla-d
Vlle-j, Vllla-d
Vlle-j, Vllla-d
Portugal*
Scotland
Spain*
1990
1991
1992
1993
1994
1990
1991
Via
VIa
Via
IXa
IXa
IXa
IXa
Vla,Vllb,j
Vla,Vllb,j
Vlllc,IXa
Vlllc,IXa
* only selected species available
•
Table 3 Main characteristics of the surveys used to describe the species distribution.
Year
/Quarter
Country
1990 I st Germany
1990 IV th Portugal
Spain
France
1992 IV th France
Scotland
Gear
HGS180 P
Norwegian CampeIl trawl
Baka 44/60
GOV 36/47
GOV 36/47
GOV 36/47
Survey dates Depth range
1/03 - 18/03
27/10 -6/12
10/09 - 14/10
25/09 - 10/11
18/09 - 30/10
18/11 - 5/12
(m)
65 -450
15 -708
37 - 444
16 - 568
17 - 576
50 - 205
Number of hauls
71
123
120
193
171
38
Table 4 Conversion coefficients for number/hour from BAKA to GOV trawls.
SPECIES
Merluccius merluccius
Micromesistius poutassou
Lepidorhombus boseii
Lepidorhombus whitfiaqonis
Anqlerfish
Traehurus traehurus
GOV/BAKA
1.06
0.51
0.34
0.26
0.69
2.18
Figure 2. ICES Sub-areas and Divisions in the area surveyed by the Study Group.
61
60 - \ - _ i - - + - - - j _ - i - i - _ - i - - i - _ . i - _ - I I : :.•·....
I·········
60
59 59
5B 5B b /
-If,,······;..·.• i···..
·;·········1 b /
56 56
55
54
+······1
53 -1 ,(../+ ,
55
54
53
52 -1·..···....;..·········1···········1 ···-\·····.. ····~r
51
52
51
50
-1········1···· ..
·-1-········ 50
49
4B
-f--i--+-----!-,.-----L--+..,,---+---I---S=\._
49
4B
47
46 -I
4 5
-I
; ..;.
,
;
+
!
1
f--+:-~--+-<v
47
46
45
44
43
42
41
40
-I
3B -I
; ; ··..
·;··········1
39
-1 :.
+· .. ,·
+······1
, ; , I····..·..·
3 7 -I ; ..;.
; , ,
···:·:c-e~~::·····t· ......
36
-f--j--+--j--t--+-.--r-i-"'--r"""'-,r---r--r-+--+-'fL-,--1-
15 14 13 12 11 10 9 a
7 6 5 4 3 2 1 0 1
37
1
44
43
42
6
•
..
..
.,
..
.s
15
0\
••
Figure 3.1 Survey areas and loeation of the trawl stations used to deseribe species distribution during A the first quarter of 1990; B the fourth quarter of 1990;
C the fourth quarter of 1992.
, .
1
·········t········l·····+·········:··········+-··_··+··....
51
--1.-1
.....J::::j.!
:L
L. .
..
(y.:.:.,I,'._..
:I, . . . . . . . . • . .
..
••
..
..
5 •
..
11 10
..
"
••
I
=r~~=-~-~='H--ic-
C
Area surveyed during the lourth quarter 011992
+ Scottish survey (18111-5112)
)( French survey (18109-30110)
4' -1---i--.;...-i-.....- .....
....;..c.,...-....-.-.....,,..........- .....- ....- .....,..........
-f-
11 10
r
Figure 3.2 Hake distribution observed during the first quarter of 1990.
56
58
57
60
59
14
-+_-L_-L_-L_---L_---L_---L_--l._--l._--l._--l._--l._
.
...,
··········1··
.....+ ;
; i ..............•...............,
+
+
..........:
.L
l l
.............. 1
, , .
___
_
___~___I~~1_61 i ..1.
: ;
1.
.'
.
..·.··.:.·.·.·.·.·.·.·.·.·.!,·.·.·..
.
.
I
........................J.....
60
59
.............+ .1
1
.+ .
·······1 r
...•.•L ..•........, •.••••••••.••.....•.•......
+ :, j••••.•...
.............,
"... ".1
i
58
57
.........l.
,
~
.. ;.1
'.:....•....
......"."" t·· .. : ...... :
56
55
54
55
54
53
52
53
52
51 v~ .•....•.
+
51
+
1-50
50
49 49
48 48
47
46
45
44
Merluccius merluccius
1 st QUARTER 1990
Numberlhour o l\)
1-50
51-100
~ 101-200
• 201-400
• 401-800
• >800
47
46
45
44
43-t--+--,---+--r--+-.,-~-~-~---.---.---.---,r----,r---11-
15 14 13 12 11 10 9 8 7 6 5 4 3 2 1
7
•
•
•
•
Figure 3.3 Hake distribution observed during the fourth quarter of 1992.
..
59
58
-i , , .
57
56
55
54
53 -i , ; ; .
52
51
50
49
48
47
46
45
44
43
15 14
Merluccius merluccius
4th QUARTER 1992
Numberlhour
13
0
1-50
~
51-100
101-200
201-400
401-800
>800
12 11 10 9 8 7 6 5 4 3 2 1
49
48
47
46
5
44
50
51
52
53
54
55
56
57
59
58
60
8
Figure 3.4 Hake distribution observed during the fourth quarter of 1990.
o
-+---'----I--.,.-'---L--::=-',...--'----+---z...L.---"'----''-----''---'---+-52
47
51
49
5 0
-1
+ \.: :
!:•........: ,..
~~
.......•.•...............:"'..:
....
"....
..
•
.. ...• \1}
..
~ .
..
+
48
-1
+ +
+···················i············· ·············"·······:··················t··:···············i·;:t'\.
46
4 5
-I·················t··················+················ + ; .
44
43
42
Merluccius merluccius
4th QUARTER 1990
41
40
39
38
Numberlhour
0
1-100
@
101-200
\1} 201-400
401-800
801-1600
39
40
>1600
38
41
37 37
................
.........
·,t
36
13 12 11 10 9 8 7 6 5 4 3 2 1
42
43
4
45
46
47
51
50
49
48
•
•
9
•
•
Figure 3.5 Length frequency distribution of hake during the first quarter of 1990.
German data
Numberofhauls :71
10
9
7
% 5
4
3
2 o
10 15 20 25 30 35 40 45 50 55 60 65 70 75 80 85 90 95 100 105
Lenglh (em)
Figure 3.6 Length frequency distribution of hake during the fourth quarter of 1992.
Scottish data
Num ber of hauls : 38
12
%10
8
6
20
18
16
14
4
2
0
0 5 10 15 20 25 30 35 40 45 50 55 60 65 70 75 80 85 90 95 100 105
Length (ern)
French data north of 48° N
Num ber of hauls : 62
20
18
16
14
12
%10
4
2 o o
5 10 15 20 25 30 35 40 45 50 55 60 65 70 75 80 85 90 95 100 105
Length (ern)
French data south of 48° N
Num ber of hauls : lOg
12
%10
8
6
4
2
0
20
18
16
14
0 5 10 15 20 25 30 35 40 45 50 55 60 65 70 75 80 85 90 95 100 105
Length (ern)
10
Figure 3.7 Length frequency distribution of hake during the fourth quarter of 1990.
French data north of 48° N
Nurn ber 01 hauls : 69
20
18
%10
8
6
4
2
0
16
14
12
0 5 10 15 20 25 30 35 40 45 50 55 60 65 70 75 80 85 90 95 100 105
Length (ern)
French data south of 48° N
Nurn ber 01 hauls : 124
20
18
16
14
12
%10
4
2
8
6
0
0 5 10 15 20 25 30 35 40 45 50 55 60 65 70 75 80 85 90 95 100 105
Length (ern)
Spanish data
Nurn ber 01 hauls : 120
%10
8
6
4
20
18
16
14
12
2
0
0 5 10 15 20 25 30 35 40 45 50 55 60 65 70 75 80 85 90 95 100 105
Length (ern)
Portuguese data
Nurn ber 01 hauls : 123
12
%10
8
20
18
16
14
6
4
2
0
0 5 10 15 20 25 30 35 40 45 50 55 60 65 70 75 80 85 90 95 100 105
Length (ern)
11
•
•
Figure 3.8 Horse mackerel distribution observed during the first quarter of 1990.
-+_-I._-I._---I_---lL.-_L-_.l-_.l-_.l-_...L._....L.._....L._....L._....L_-I._--+_61
60
59
5B
.......".. !
.
"
~
!
:
".". ··l"
..
• ~ •
..........
[
~
..
1
H _ "
~
•
..·..·..·•······..·..·..
.
~"·············;·······················i··
60
59
5B
....·..·+············1-57
•••••••••• ,j. •••••••••• •
56
55
57
,,··········· .. ·.... ·····f··
54 "
......."'."!'..,,
! .
I
!
:
;.
! •
······..·.."f····
.
53
52
51
•
49
50
4B
47
46
45
44
Trachurus trachurus
1st QUARTER 1990
•
Numberlhour o
1-1000
; 1001-2000
~ 2001-4000
• 4001-8000
8001-16000
• >16000
43
15 14 13 12 11 10 9 B 7 6 5 4 3 2 1
49
4B
47
46
45
44
51
50
52
53
56
55
54
12
,.-1
Figure 3.9 Horse mackerei distribution observed during the fourth quarter of 1992.
-t--L--...L---L---JI...--...L----L---JI...-_.1-_--L_-1_ _ o
+-_...L._-1_v-L-_+
60
59
58
57 Ji· ··•
/-57
56 -j , j············,·················f··········· i , ,
·········A
,l('I;,rldW::.....
55 -j· · ·..
o · ·..·
+ ·· · f··
'1
........· · ! - 5 5
54
53
..............+.
52
51
50
:
;
~
..
r .
1 i
• .. t •
1.
49
•
48
47
46
45
44
Trachurus trachurus
4th QUARTER 1992
•
•
Number/hour o
1-1000
; 1001-2000
~ 2001-4000
4001-8000
8001-16000
• >16000
...
• ~
,-
.
..
.. i
.:.
..............................., ....... t ..,
43-+--,---r--.,.--,,--+----''-r---,r--,---r--,--,---r--,---r--_+_
15 14 13 12 11 10 9 8 7 6 5 4 3 2 1
46
7
5
4
8
50
51
52
53
54
56
58
59
13
•
•
•
•
Figure 3.10 Length frequency distribution of horse mackerel during the first quarter of 1990.
German data
Num ber 01 hauls : 71
14
12
% 10
8
6
4
2
0
20
18
16
0 5
10 15 20
Length (ern)
25 30 35 40
Figure 3.11 Length frequency distribution of horse mackerel during the fourth quarter of
1992.
Scottish data
Numberolhauls :38
20
18
16
14
12
%10
8
6
4
2
0
0 5 10 15 20
Length (ern)
25 30
·~1
35 40
French data
Num ber 01 hauls :171
%10
8
6
4
2
0
20
18
16
14
12
0 5 10 15 20
Length (ern)
25 30 35 40
14
57
58
59
Figure 3.12 Mackerel distribution observed during the first quarter of 1990.
60 o
- + - - - l - - - + _.......
_-+--+--+---Io--"-~--"----'-~'----''----''--:t-6 J.
····f······ ..•..
·'j"···..····•·····•·..·
......
~ !
~~,t~i:~
60 i
!.~
....".... J...
t , •.•.••• : ' "
...................... .1..
·······;·············..r········
1
...... : ... ".... 1.
59
1.
58
..L
! .
'~
,~
······t················,·······
I·
57
5 6
-t +
56
55
54
53
52
5J.
50
49
48
47
1st QUARTER 1990
Numberlhour
46
45
44
0
1-50
51-150
•
~
~ 151-450
451-1350
1351-4050
>4050
43-+--,.--,.--,.--t--+-.L..y---r---r---,---,--:;--:;--;--r--t-
44
46
45
15 14 13 12 11 10 9 8 7 6 5 4 3 2 J.
47
48
49
50
5J.
52
53
54
55
15
•
•
•
•
59
Figure 3.13 Mackerel distribution observed during the fourth quarter of 1992.
o
- + - - - J L - - . . L - - - J - - - - - ' - - f - - . . . L . . . - - J . _ - J_ _.l-_...L-_.....L_--JL-_..L--cr-J-_-f_60
....•......
,
..
~
~
.
0-
' ...
~ ... .J
59
58
57
.........
, •...............
) .
·,······························1-57
56
55
. ..t"
+ .
........... l
~
···· .. ····t····.. ·······: i ; , ;
54
53
52
51
50
49
48
47
46
45
44
.@
............,1 ..
1
.""
······..····i
......
}
.. ..
..........
....
I
+
.. """" .. : ..." ....".1' ....
·······1
'''1
:.~~
·..·.. ····,1
...........!
. !
1
.i
."':
{ .
.10 •
• • ! • . . . .
~
~
... .....
t
; ,
·
...........1..
j ·..·
; " ;
1 •
.....j
j •
. ··············1..··
,
.
."."
....
~
....·.........,1·· i"
·l·..
..·· ....·,: ..··· ..·..
............ 1
..l .................•........i
Scomber scombros
4th QUARTER 1992
!
, .
Numberlhour o
Q
1-50
51-150
~ 151-450
• 451-1350
• 1351-4050
!.
• >4050
...
.............
'
.
:'.'
..............+
'.
.....
'@'
.
~.';
• Q
..............
'
..
~~~
..
~
.
..
·············1
: L :.~
4 3 - t - - - ; - - f - - ; - - + - - - , - - ' - r - - - r - - - , , - - y - - - , - -....
--r-.,.---,--i-
15 14 13 12 11 10 9 8 7 6 5 4 3 2
54
53
52
51
50
49
48
47
46
45
44
56
55
58
16
Figure 3.14 Length frequency distribution of mackerel during the first quarter of 1990.
German data
Number of hauls : 71
20
%
15
10
5
0
35
30
25
10 15 20 25 30
Length (ern)
35 40 45
Figure 3.15 Length frequency distributions of mackerel during the fourth quarter of 1992.
Scottish data
Num ber 01 hauls : 38
35
30
25
20
%
15
10
5
0
10 15 20
I!:
25 30
Length (ern)
'"
35 40 45
French data
Numberolhauls: 171
%
20
15
10
35
30
25
5
0
10 15 20 25 30
Length (ern)
35 40 45
•
•
17
•
Figure 3.16 Lepidorhombus whiffiagonis distribution observed during the first quarter of
1990.
o
-+_-1-_....L._--'-_-+--_-!-_..1.-_.1.-_1-_1------''------I_---L_---I._-1-_-+-61.
60
59
-I , ; · ..
1.. .. ..
1. .
.......................1.
.
~
..
.....; ......
..
~
+
!..
.
,
..·i..·..........· 59
58 58
57 57
56
55
56
55
54 54
53 53
52 52
51.
51.
50 50
49 49
48 -1· · ·,·..· ·..·
47
46
45
44
· , ·..·; · ·..
1 ,7"
..... Lepidorhombus whiffiagonis
1st QUARTER 1990
•
Numberlhour o
1·5 o 6-10
~ 11-20
• 21-40
• 41-80
>80
; ,; ""-
43
-1----r---i--...;--+---i--L,---..--,...-----,..----.---r---r---.--......--1-
1.5
14 1.3
1.2
1.1.
1.0
9 8 7 6 5 4 3 2 1.
48
47
6
45
44
18
Figure 3.17
Lepidorhombus whiffiagonis distribution observed during the fourth quarter of
1992.
59
- + _ - - l - _ - - l - _ - - l - _ - ' - _ - ! - _ - ' - _ - L - _ - L - _ . . . . L . - _ . . . . L . - _ . . . L - _ - ! - _ . . . L - . . " . . . . . . . L - _ + - 6 0
-+· ·..·
·f..•· · ,..· ·..·..,
........ 1
·f ·
•••••••••••.•• ,i. •••.• i .
, ..
~
.
..~
,
~
+ .
59
58
.........................
:, ...
,
58
57 -f ··........·..
·+..........···...
f ...·........·•..· ·,......··........, ......· ..·..·1-57
56 56
55 55
54 -f , ..
54
53
52
51
....................1
.~
....
···············i..··..·
+
~
:
53
52
51
50
49
48
47
46
45
44
...............+...
:
• 1 ............ : !
·············t
J
L
.............
····'t
"~
~
.
.
1
".!
:
I
.
~ j
"
........ !
:
...
i.~Pid~rhO~bUS wi,/ffiagonis
4th QUARTER 1992
I .....
t..··············..····
Numberlhour o
1-5
; 6-10
@ 11-20
!
• 21-40
• 41-80
. . >80
'@ •
, ,..•
43
-t----r--y---y----r--+-----''-r---.---.--,--,--,--,---,---,---+_
15 14 13 12 11 10 9 B 7 6 5 4 3 2 1
50
49
48
47
46
44
5
19
•
•
•
•
Figure 3.18
Lepidorhombus whiffiagonis distribution observed during the fourth quarter of
1990.
o
-t----l------'------,....L-----l.--=....a..,.".....--''--....L..-"?""'---....L.--"'------'----l----t-52
51
50
49 ..
48 --j j ;
+
, ••
!
···t·..··················;··············..·..···;········ ;...•
~
~
1;
;
•
.
.
47
-I ,
46
45
1..· ;
.................1.
;
;,: '
·
+ _ ;...
;
....................+ i i
,..........
• .
·····,',··"···1··'
• •
.
i i..............
····I·........·..······....,......·..··..·..·..
·T .. ··..··..·
·~r;··~·
··~
44 44
43
42
....................1.
.
...................
;
....
..................
\
···..······..··..·················....···t
~
41 i
...1
...................l
40
1.••.
1 l ,.~ ..
~.
39
..················1
38
......................,.,., ,,··,,········t
37
36
-f---+---..---l----+---r----i--+=--,c----r---;---+--+--__+_
13 12 11 10 9 8 7 6 5 4 3 2 1
43
4th QUARTER 1990
Numberlhour
1-10
..
11-20
~ 21-40
•
• 41-80
81-160
• >160
42
41
40
39
38
1 ....·........·..
+..·........·......
·11-37
49
48
47
46
45
50
51
20
Figure 3.19 Length frequency distributions of Lepidorhombus whiffiagonis during the fourth quarter of 1992.
Scottish data
Nurnber 01 hauls : 38
7
6
% 5
4
10
9
8
3
2
1 o
+-r...-r-r-r--t"'r-ol"'r-+".-'''''''''''''' o
5 10 15 20 25 30
Length (ern)
35 40 45 50 55
French data north of 48° N
Nurnberolhauls :62
6
% 5
4
3
2
10
9
8
7
7
6
% 5
4
3
2
10
9
8 o
5 10 15 20 25 30
Length (ern)
35 40 45 50 55
French data south of 48° N
Nurnberolhauls: 109 o
5 10 15 20 25 30
Length (ern)
35 40 45 50 55
21
•
22
Figure 3.21
Lepidorhombus boscii distribution observed during the fourth quarter of 1990.
12 7 6 o
-+__
-I-_ ___L_--.,..__--'_~--I.-,_-L----l--...,.....L---L.---I-----L--,..I--_1_52
51 -I , .
51
50
49
48 - , .
+
.'
+ + ,
47
4 6 -1 +
4 5
-I ;
, j !
j ; .
......
: ..
+ .; : ; .;.
; : ····:········..·..····, ,·········..
·····1
44
43
42
41
40
39
.. %
4th QUARTER 1990
Numberlhour
0
1-20
•
@ 21-40
~ 41-80
81-160
161-320
39
40
>320
38
41
38
37
36
13 12 11 10 9 8 7 6 5 4 3 2 1
43
42
37
44
47
46
45
50
49
48
23
•
•
•
•
%10
8
6
4
20
18
16
14
12
2
0
5
Figure 3.22 Length frequency distributions of Lepidorhombus boscii during the fourth quarter of 1990.
Spanish data
Number of hauls : 120
%10
8
6
4
2
0
20
18
16
14
12
5 10 15 20 25
Length (em)
30 35 40
Portuguese data
Numberofhauls: 123
10 15 20 25
Length(em)
30 35 40
24
Figure 3.23
Lophius piscatorius distribution observed during the first quarter of 1990.
o
-+_-L_-L_-L_--I-_--I-_-L_-L_-L_-L_-L_--+_---l_--!_---l_--1_6~
6 0
+
···i········ ., ~ , : .
59
5B
60
·······..··1···
!
·······..t·..·..
·········f···
.... ,!
.. j
!
59
5B
57 57
56
............ t i
············..<·······..
·····f·
.: ........ 1.
!.
+
.. i
;.
~ .
56
55 55
54 54
53 53
52 52
5~
5 0
-I
5~
··········r j•••••••••.•
~
v,...•..... • ; ;.........
.j
j.........
·If-
50
49 49
4B
47
46
45
44
Lophius piscatorius
1st QUARTER 1990
Number/hour o
@
1
2
~
3-4
• 5-8
9-16
>16
43
15 14 13 12 11 10 9 8 7 6 5 4 3 2 ~
48
47
46
45
44
25
•
•
•
Figure 3.24
Lophius piscatorius distribution observed during the fourth quarter of 1992.
o
- t - - - + - - - + - - - ' - - + - - i - _ - 1 - _ - 1 - _ . . . . L . - _ . . . . L . - - - + - - . . L - - + - - . . I - - V - . . I - - f - 6 0
59
1 .......; 1 ;
+ .
.....f
'"
.
59
58
...............i
j
, j..••..•.••.••...
+ , .
57
............ ··············"r····
\:J
·!················t··································t-58
.,.·································11-57
56
55
................j
·..·..··..·····t········
...............+ j.•••••.
"""...!.................•...............
11- 5 6
"1·····················11-55
54
................!.....
54
53
52
-I ,.........
i
+ +
~
51
50
•
49
48
47
46
45
44
········t·......·..······
.L········ j••·•·••·•·•·
........... 1 ......... .!
t ···········+··················
1.. .
Lophius piscatorius
4th QUARTER 1992
•
•
Numberlhour o
$
• 1
2
~ 3-4
• 5-8
9-16
>16
.
.··.·.. 1.
L···········.,.·······..···············..
·······i········· :.:
~;
j
.............. :
;
,....
i
!
:
;
43
-,.--,.--.,--.,--.---.---,--t-
15 14 13 12 11 10 9 B 7 6 5 4 3 2 ].
8
47
46
5
44
51
52
53
26
Figure 3.25 Lophius piscatorius distribution observed during the fourth quarter of 1990.
~2 o
-+---t----L---,....l.-----l--=~ ......--JI--_+-_z_l.----L--..L...----L--..l.--_t-5 2
5~ 5~
5 0
-1
+ -, ;.. ,
+....•..........;; ; ;
+ v·i···············:······················,············
+ ; .................•
-50
49 - ; !
.
49
46 -;····..
··············i·····················~··········· .
46
47
46
-I + +............ .., :
- j
+ +. "" : ;.
45
44
43
42
4th QUARTER 1990
4~
40
39
.
"
....
Numberlhour
0
• 1
~ 2
~
5-8
3-4
9-16
>16
36
37
36
~3 ~2 ~~ ~o 9 B 7 6 5 4 3 2 ~
37
44
43
42
4~
40
39
45
46
47
27
•
•
•
•
10
6
% 5
4
9
8
7
3
2
Figure 3.26 Length frequency distribution of Lophius piscatorius during the fourth quarter of
1992.
Scottish data
Number of hauls : 38
6
% 5
4
3
2
10
9
8
7 o
5 10 15 20 25 30 35 40 45 50 55 60 65 70 75 80 85 90 95 100 105
Length (cm)
Figure 3.27 Length frequency distribution of Lophius piscatorius during the fourth quarter of
1990.
Spanish data
Number of hauls : 120
5 10 15 20 25 30 35 40 45 50 55 60 65 70 75 80 85 90 95 100 105
Length (cm)
28
Figure 3.28
Lophius budegassa distribution observed during the fourth quarter of 1992.
o
+--I..---+--f---i-----!i---.JI--L--L--.L-_+-_...L-_-l-_+--.r....l-_+-
60
.....................
,
.
59
....; ··, · , ···········t
1 i
......
) .
!
..•...•....1.
59
58
~
..................+
I
+ ..1
+
<
..
.1.
57
56
....
.;..
i
..... ;J
55 r ......·....
-+· ..·......·....t ..·........·......·..........·..
!-58
,·· .........., ....·..·· ..·1:-57
"'-;;:.., · ·..·, ·.. 1-56
... ·..··+
1~55
54
53
52
51
50
49
.•.•••.••.••.•..1..• ......L.....
..............
~ ·············f j
.........L
..............
j
"' .
........... .i
I
~
.
...... ;
......:., \l\I
~
.......
:
..
." 41z
§
......, j.;.: :.
, ·..
'@
~
f»; ., ; :.
, ;
'
.i~
i i':"~
; j 0
.
·····..·t··· ..·..·..·..
i
,.,
'''i'
.............................
~
.
~,
.
··········1·..··············•
48
47
46
45
44
Lophius budegassa
4th QUARTER 1992
•
Number/hour o
§
..
1
2
~ 3-4
• 5-8
. . 9-16
>16
~
......
............., 1,.,.,.., ,..
~~,-!.
'
~~ f...
i
:.;
~
············t··..
43
-t---.---t---r--,---,r--L-r--r--,--.,--.,---.---,--....
- - , - - - f -
15 14 13 12 11 10 9 8 7 6 5 4 3 2 1
47
46
45
44
48
54
53
52
51
50
49
29
•
•
Figure 3.29 Lophius budegassa distribution observed during the fourth quarter of 1990.
.l--_--L_--....L-_--L_ _
-J..,__
- - - l ' - - - - l - - . . . , . . . . . L . - - - - ' - - - . L - - - . l . . - - . l - - - - + - 5 2
51
-+ , : + + ,
51
50
-I i ..,,,"", ,, , j
§!% "', , , ;
.v.i
; ·..· ·
·1-50
49
-I
+ :
,...........!
, : " :: !
, : +...
· ..
i ..
49
48
47
"
46
45
4 4
-1·....·..··........
+· ..........·......···......·........·..
+..··..·....··..··..
1....·..
43
•
41
42
40
39
38
4th QUARTER 1990
NumberJhour
0
• 1
~ 2
•
~
5-8
3-4
9-16
>16
37
36
1 3
....................1
1 2 11 1 0 9 a
7 6 5
..
3 2 1
42
41
40
39
38
37
44
43
48
47
46
45
30
Figure 3.30 Length frequency distribution of Lophius budegassa during the fourth quarter of
1990.
Spanish data
Number of hauls : 120
10
9
6
% 5
4
8
7
3
2
5 10 15 20 25 30 35 40 45
Length (ern)
50 55 60 65 70 75 80
•
•
31