.
,
•
•
International Council for
the Exploration of the Sea
C.M. 1994/G:3 Ref.H
Demersal Fish Committee
'.>~-"'"
PRELIMINARY REPORT OF TBE INTERNATIONAL O-GROOP rISS SORVEY
IN TBE BARENTS SEA AND ADJACENT WATERS IN AUGUST-SEPTEMBER 1993
The twenty-ninth annual International Q-group fish survey was
made during the period 16 August - 8 September 1993 in the
Barents Sea and adjacent waters. The following research vessels
participated in tpe survey:
~
State
Name of vessel Period
Norway
Norway
Russia
Russia
"Johan Hjort"
"G.O. Sars"
"Professor Marty"
"Pinro"
Research Institute
16.8 -
8.9
17.8 -
7.9
22.8 23.8 -
8.9
6.9
Institute of Marine
Research, Bergen
The Polar Research
Institute of Marine
Fisheries and Oceanography, Murmansk
Names of scientists and technicians who took part on the
different vessels are given in the Appendix.
Preliminary analysis of the survey data were made 5-7 October in
Murmansk. Observations concerning the geographical distribution
of Q-group fish and their abundance are given in this report
together with abrief description of the hydrographical
conditions in the area.
.•
..
"
2
MATERIAL AND METHODS
The geographical distribution of O~group fish were estimated
with a small mesh midwater trawl. The vessels which participated
in the survey in 1993, used the type of midwater trawl recommended by the meeting held after the survey in 1980 (Anon.,
1983). The trawling procedure was standardized in accordance
with the recommendation made at the same meeting. At about every
30 nautical miles sailed the trawl was towed in several depths
in one haul. The standard procedure consisted of towings of 0.5
nautical mile in each of 3 depths with the headline of the trawl
located at 0, 20 and 40m. An additional tow at 60 and 80m for
0.5 nautical mile was made when O-group fish layer was recorded
on the echosounder deeper than 60m.
A procedure starting with the trawl in the deepest layer would
probably be preferable. The trawl would then be "fishing" more
efficient troughout the whole trawl haul and maintain the
correct geometry. It is recommended that this is further
investigated.
Survey tracks and hydrographical stations are given
Trawl stations with and without catch are indicated
distribution charts in Figs. 16 - 27, as filled and
respectively. The density grading is based on catch
per 1.0 nautical mile trawling.
in Fig. 1.
on the
open symbols
in nuIDber
HYDROGRAPHY
Observations were made along all the survey tracs with 5 to 40
nautical miles between stations. Horizontal distributions of
temperatures and salinities are shown for 0, 50, 100, 200m and
bottom (Figs. 2-11). Figs. 12 - 15 show the temperature and
salinity conditions along the Bear Island - West, Bear Island North Cape, Kola and Cape Kanin sections. The mean temperatures
in the main parts of these sections are presented in Table 1.
According to temperature conditions, 1993 is the fifth "warm
year" in succession. However this year the mean sea temperatures
from 0-200m in the Barents Sea were only 0.1-0.2°C above the
long term average (1965-1993). Compared to 1992 a significant
fall in temperature (0.3-0.6°C) was observed. The reduced
positive anomalies in the hydrographical standard sections are
mainly due to lower heat content in the layer below the seasonal
thermoclyne. The temperature in 50-100m layer along the Kola
section was the same as observed in 1992. Upper mixed layer
temperature was significantly (0.5-1.5°C) higher than previous
year. This is associated with the favourable conditions of the
summer heating. In the West Spitsbergen Current (0-200m)
temperature was O.8°C above long term average and 0.4°C higher
than in 1992. Upper mixed layer temperature in this area was
•
3
also 1~0-1.5°C higher than previous year. Observed thermal
conditions were caused by decrease of Atlantic inflow into the
Barents Sea and increase into the area west of Bear Island and
West of Spitsbergen.
"
~..
'
,
,'"
DISTRIBUTION AND ABUNDANCE OF O-GROUP FISH AND GONATUS FABRICII
•
Geographical distribution of O-group fish are shown as shaded
areas in Figs; 16 - 26; arid of Gonatus fabricii in Fig. 27.
Double shading indicates dense concentrations. The criteria for
discriminatingbetween dense and scattered concentrations are
the same as used in earlier reports (Anon.,1980). Abundance
indices; estimated as the area of distribution with areas of
high densities weighed by 10, are given in Table 2. Another set
of abundance indices are given for O-group herring, cod and
haddock (Table 3) as described by Randa (1984). These are based
ori the nurnber caught during a standard trawl haul of one
nautical mile. Length frequency distributions of the main
species are given in Table 4.
.
.
The main distribution ofO-group fish were more westerly
orientated in 1993 than in previous years.
Berring
(Fig. 16)
Scattered concentrations with small dense patches were observed
more westerly than last year andapproched 80 0 N west of
Spitsbergeri~. The main concentrations were found in the central
part of the Barents Sea, and herring was distributed eastwards
to.38°E. The logarithrnic abundance index is astimated at 0.75,
which is,somewhat below the 1991 and 1992 indices, but still the
fourth highest in the time-series. This indicates that the 1993
yearclass is relatively strong.
•
capeiin
(Fig. 17)
As in last,year b-group capeiin was distributed mainl~ neir
Novaya Zemlya and in small quantities, and it seems to be the
second very poor yearclass in succession~ The length
distribution indicates that this O-group comes from summer
spawners~
C6d
(Fig. 18)
Along west off Spitsbergen dense concentrations were recorded in
a wider area than previous year. The largest areawith dense
concentrations was found in the central part of the Barnts Sea,
from about 20-40 o E and north to 76°N. In the southeastern part
of .the Barents Sea mainly scattered concentrations were
observed. Q-group cod was absent in the south east off the
4
Murman coast until Kolguev island. The logarithmic abundance
index for the 1993-yearclass (2.09) is the fifth highest in the
time-series, above that of 1983, and the yearclass must be
classified as strong.
(Fig. 19)
Haddock
To the west of Spitsbergen O-group haddock was distributed in a
smaller area than last year. In the western and central part of
the Barents Sea the distribution was similar to that of 1992,
but with few observations east of 35°E. The logarithmic index
was estimated at 0.64, which is the third highst since 1965,
indicating another good yearclass of haddock.
Saithe
(Fig. 20)
In the central part of the investigated area saithe was less
abundant than in 1992, while to the west of Spitsbergen the
species was observed over a larger area and in higher
quantities. In most years there are relatively few observations
of O-group saithe in the Barents Sea and therefore no index is
calculated.
Polar cod
(Fig. 21)
O-group polar cod is distributed in two seperate areas, with one
component west and southeast of Spitsbergen and a second
component in east along the western coast of Novaya Zemlya.
During the O-group survey the total area of distribution is not
completely covered.
In the north-western area, the distribution was similar to
previous year. The index was just a little lower than in 1992
and the 1993 yearclass of polar cod in this area seems to be of
average abundance.
Dense concentrations of the eastern component were distributed
over a wider area than in 1992. During the following pelagic
fish survey in September it was found that this component
covered a much wider area towards northeast, but since this area
normally is not investigated during the O-group survey, it was
not included in the calculations of the abundance index. Still
the index incre~sed by 32% and the 1993 yearclass in the eastern
area may be considered to be at least of average strength.
5
Redfish
(Fig. 22)
Similar to 1992, O-group redfish was only distributed in the
western areas, along the western edge of the shelf and off WestSpitsbergen. The area with dense concentrations was larger this
year, and the index slightly higher (8%), but still the
yearclass must be considered to be poor, as in the two previous
years.
Greenland halibut (Fig. 23)
Only scattered concentrations were recorded in limited patches
mainly to the west of Spitsbergen. The index increased with a
factor of more than 3 compared to previous year, but the
yearclass is still poor (the sixth in succession) .
•
Long rough dab
(Fig. 24)
The O-group was mainly distributed in the central part of the
Barents Sea over a wider area than previous year and with some
scattered patches in the eastern and southeastern areas. No
dense aggregations were observed. The index increased by over
40%, but still the yearclass is considered to be another poor
one.
Sandeel (Fig. 25)
Sandeel was mainly observed nearby Novaya Zemlya as in 1987, but
further to the north.
Catfish (Fig. 26) and blue whiting.
•
Only a few specimens of catfish and blue whiting were caught .
Gonatus
(Fig. 27)
O-group Gonatus fabricii was found in the western part of the
investigated area, but in smaller quantities than in 1992.
6
REFERENCES
Anon., 1980. Preliminary report of the International O-group
fish survey in the Barents Sea and adjacent waters in
August/September 1978. Annls bioi., Copenh., 35:273-280.
Anon., 1983. Preliminary report of the International O-group
fish survey in the Barents Sea and adjacent waters in
August/September 1980. Annls bioi., Copenh., 37:259-266.
Randa, K. 1984. Abundance and distribution of O-group ArctoNorwegian cod and haddock 1965-1982. Pp. 189-209 in
Godl2l, O. R. and Tilseth, H. (eds.): Reproduction and recruitment 01
Arctkco~ Proceedings of the first Soviet-Norwegian
symposium, Leningrad, 26-30 September 1983. Institute of
Marine Research, Bergen, Norway.
Tereshchenko, V.V. 1992. Some results from long-term oceanographie observations during O-group surveys in the
Barents Sea. ICRS CM 1992/C:18.
Toresen, R. 1985. Recruitment indices of Norwegian springspawning herring for the period 1965-1984 based on the
international O-group fish surveys. ICES CM 1985/H: 54 .
•
--------------------1
7
Table 1. Mean water temperature 1 in main parts of standard
sections in the Barents Sea and adjacent waters in
august-September 1965 - 1993.
Section 2 and layer (deep in meter)
Year
•
•
1
2
3
4
5
6
7
0-50
50-200
0-200
O-bot.
O-bot.
0-200
0-200
1965
1966
1967
1968
1969
1970
1971
1972
1973
1974
1975
1976
1977
1978
1979
1980
1981
1982
1983
1984
1985
1986
1987
1988
1989
1990
1991
1992
1993
6.7
6.7
7.5
6.4
6.7
7.8
7.1
8.7
7.7
8.1
7.0
8.1
6.9
6.6
6.5
7.4
6.6
7.1
8.1
7.7
7.1
7.5
6.2
7.0
8.6
8.1
7.7
7.5
7.5
3.9
2.6
4.0
3.7
3.1
3.7
3.2
4.0
4.5
3.9
4.6
4.0
3.4
2.5
2.9
3.5
2.7
4.0
4.8
4.1
3.5
3.5
3.3
3.7
4.8
4.4
- 4.5
4.6
4.0
4.6
3.6
4.9
4.4
4.0
4.7
4.2
5.2
5.3
4.9
5.2
5.0
4.3
3.6
3.8
4.5
3.7
4.8
5.6
5.0
4.4
4.5
4.0
4.5
5.8
5.3
5.3
5.3
4.9
4.6
1.9
6.1
4.7
2.6
4.0
4.0
5.1
5.7
4.6
5.6
4.9
4.1
2.4
2.0
3.3
2.7
4.5
5.1
4.5
3.4
3.9
2.7
3.8
6.5
5.0
4.8
5.0
4.4
3.7
2.2
3.4
2.8
2.0
3.3
3.2
4.1
4.2
3.5
3.6
4.4
2.9
1.7
1.4
3.0
2.2
2.8
4.2
3.6
3.4
3.2
2.5
2.9
4.3
3.9
4.2
4.0
3.4
5.1
5.5
5.6
5.4
6.0
6.1
5.7
6.3
5.9
6.1
5.7
5.6
4.9
5.0
5.3
5.7
5.3
5.8
6.3
5.9
5.3
5.8
5.2
5.5
6.9
6.3
6.0
6.1
5.8
Aver.
19651993
7.3
3.8
4.7
4.2
3.2
5.7
3.6
4.2
4.0
4.2
-
4.2
3.9
5.0
4.9
4.9
4.8
4.0
4.1
4.4
4.9
4.4
4.9
5.1
5.0
4.6
4.4
3.9
4.2
4.9
5.7
5.4
5.0
5.4
4.6
1)
Earlier presented temperatures have been slightly adjusted
(Tereshchenko, 1992).
2)
1-3:
4:
5:
6:
Murmansk Currenti Kola Section (70 0 30'N-72°30'N,33°30'E)
Cape Kanin section (68°45'N - 70 0 05'N, 43°15'E)
Cape Kanin section (71°00'N - 72°00'N, 43°15'E)
North Cape Currenti North Cape - Bear Island section
(71 0 33'N,25 0 02'E - 73°35'N,20046'E)
7: West Spitsbergen Currenti Bear Island - West section
(74 0 30'N, 06 0 34'E - 15°55'E)
8
Tabeil 2. Abundance indices of O-group fish in the Barents Sea
and adjacent waters in 1965 - 1993.
Year
1965
1966
1967
1968
1969
1970
1971
1972
1973
1974
1975
1976
1977
1978
1979
1980
1981
1982
1983
1984
1985
1986
1987
1988
1989
1990
1991
1992
1993
Cod
6
1
34
25
93
606
157
140
684
51
343
43
173
106
94
49
65
114
386
486
742
434
102
133
202
465
766
1159
910
Haddock
7
1
42
8
82
115
73
46
54
147
170
112
116
61
69
54
30
90
184
255
156
160
72
86
112
227
472
313
240
Polar cod
West
I
East
0
129
165
60
208
197
181
140
(26)
227
75
131
157
107
23
79
149
14
48
115
60
111
17
144
206
144
90
195
171
70
144
302
247
73
50
39
16
334
366
155
120
41
48
239
118
156
Redfish
159
236
44
21
295
247
172
177
385
468
315
447
472
460
980
651
861
694
851
732
795
702
631
949
698
670
200
150
162
Greenland
halibut
Long
rough
dab
1
1
8
3
13
21
16
9
35
22
12
38
17
16
40
36
55
41
8
5
2
1
3
11
66
97
73
17
26
12
81
65
67
83
113
96
72
76
69
108
95
150
80
70
86
755
174
72
92
35
28
32
55
•
•
9
Table 3. Estimated logarithmic indices with 90% confidence
limits of year class abundance for O-group herring,
cod and haddock in the Earents Sea and adjacent waters
1965 - 1993.
Herring 1
Year
1965
1966
1967
1968
1969
1970
1971
1972
1973
1974
1975
1976
1977
1978
1979
1980
1981
1982
1983
1984
1985
1986
1987
1988
1989
1990
1991
1992
1993
•
•
1)
Index
0.14
0.00
0.00
0.01
0.00
0.00
0.00
0.05
0.01
0.00
0.00
0.01
0.02
. 0.09
-
0.00
0.00
1.77
0.34
0.23
0.00
0.00
0.32
0.59
0.31
1.19
1.06
0.75
Confidence
limits
0.04
0.00
-
0.31
-
0.04
-
0.03
0.01
0.08
0.01
0.00
0.01
0.01
0.03
0.05
0.20
1.29
0.20
0.18
2.33
0.52
0.28
0.00
0.16
0.49
0.16
0.90
0.69
0.45
0.03
0.53
0.76
0.50
1.52
1.50
1.14
-
-
Haddock
Cod
-
-
-
Index
Confidence
limits
+
0.02
0.04
0.02
0.25
2.51
0.77
0.52
1.48
0.29
0.90
0.13
0.49
0.22
0.40
0.13
0.10
0.59
1. 69
1.55
2.46
1.37
0.17
0.33
0.38
1.23
2.30
2.94
2.09
0.01
0.02
0.01
0.17
2.02
0.57
0.35
1.18
0.18
0.66
0.06
0.36
0.14
0.25
0.08
0.06
0.43
1.34
1.18
2.22
1. 06
0.01
0.22
0.30
1. 04
1.97
2.53
1. 70
0.04
0.08
0.04
0.34
3.05
1.01
0.72
1. 82
0.42
1.17
0.22
0.65
0.32
0.59
0.18
0.18
0.77
2.08
1.98
2.71
1. 70
0.40
0.47
0.48
1.34
2.65
3.39
2.51
Index
Confidence
limits
0.01
0.08
0.00
0.29
0.64
0.26
0.16
0.26
0.51
0.60
0.38
0.33
0.12
0.20
0.15
0.03
0.38
0.62
0.78
0.27
0.39
0.10
0.13
0.14
0.61
1.17
0.87
0.64
0.00
0.03
0.00
0.20
0.42
0.18
0.09
0.15
0.39
0.40
0.24
0.21
0.07
0.12
0.10
0.00
0.30
0.48
0.60
0.23
0.28
0.00
0.05
0.10
0.48
0.98
0.71
0.48
Assessment for 1965-1984 made by Toresen (1985).
0.03
0.13
0.02
0.41
0.91
0.36
0.27
0.40
0.68
0.85
0.51
0.48
0.19
0.28
0.20
0.05
0.52
0.77
0.99
0.31
0.52
0.25
0.34
0.20
0.75
1.37
1.06
0.82
---~---~-----~-
--- - - - - - - - - - -
10
Table 4. Length distribution of O-group fish in percent in the Barents
Sea and adjacent waters in August - September 1993.
Length
(mm)
15-19
20-24
25-29
.30-34
35-39
40-44
45-49
50-54
55-59
60-64
65-69
70-74
75-79
80-84
85-89
90-94
95-99
100-104
105-109
110-114
115-119
120-124
125-129
130-134
135-139
140-144
Herring
Capelin
Cod
Haddock
Polar cod
East
West
1.9
3.7
10.8
13.3
12.6
18.6
22.6
9.7
6.4
0.3
0.2
4.9
29.6
40.5
22.0
2.7
0.1
Redfish
Green
land
halib
Long
roug
dab
Sandeel
1.1
0.1
0.6
3.8
3.8
8.0
12.1
16.7
25.5
16.8
8.8
3.0
0.5
0.1
0.1
0.1
0.4
3.3
10.0
18.8
21.4
14.2
3.0
5.0
3.3
5.8
4.6
5.0
2.3
1.9
0.9
+
0.1
0.2
0.5
1.3
2.0
3.1
5.9
10.7
15.5
20.2
17.1
13.5
6.9
2.3
0.6
+
+
+
+
+
0.1
0.1
0.2
0.3
0.4
0.5
0.8
1.1
2.1
2.9
6.3
15.1
14.2
15.8
14.0
13.4
6.2
2.3
1.8
1.2
0.6
0.5
5.3
24.4
45.4
21.0
3.8
0.1
+
+
+
-
4.3
-
8.7
-
-
5.7
44.3
38.6
10.2
8.7
30.4
30.4
13.0
4.3
+
0.8
12.3
19.9
17.3
9.1
9.2
10.8
10.
3._
3.2
2.3
1.4
0.1
Total
nurnbers
327193
702
134724
5810
278129
48055
34018
23
176
7431
Mean
length
74.6
54.1
81. 8
101.5
45.8
36.4
31.7
53.2
35.0
48.4
•
11
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10°
15°
20°
25°
40°
35°
30°
45°
50°
55°
60°
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Z Z
1
= iZ , -Z k
lr
Z
iZ
I
-z~L-zh-~'\j
~
f
1
i
\
"k)-+' ~~z~ZO;1 z~J
Z
~~~.
.
.~ ~
z-z
lJ(
H-Z-H-Z-Z-Z
nz_~t
\ l<
N~
~
.- :-- . -~1
r
.J"' -
67o.L-,.~~....-,_.........~:;::""-~-r-.-.-~--r-""-'~""""T~~~"""""-,-,-"",,,~""""-T-"':"""''''''''''~_'''''''''''-'~~_-''
Fig. 1. Survey tracks and hydrographie stations.
l
L
.
12
5·
10·
20·
25·
30·
35·
40·
&>
~
[
1993
t·e 0 m
18·
I
I
r
:iI
i
•i
i-
I
I
I•
•
•
· _.
,., Isotherms at 0
F19.
ID.
13
25·
30·
35·
45·
1993
S Om
•
73"
72·
70·
. 3. Isohalines at 0
Flg.
ffi.
14
2C.
lS·
30·
35·
40·
79·
1993
t·e 50 m
•
76·
.
L
7S·
I
7
)
F·19. 4 . Isothenns at 50
ffi.
15
79°
~
~
1993
•
S 50 m
78°
n°
76°
75°
74°
73°
350
~ '-~!i1
,
~!i2)
72°
71°
70°
69°
68°
67°
Fig. 5. Isohalines at 50
ffi.
16
50
100
150
20°
25°
30°
350
400
45°
550
60°
BOO
~.
1993
t"C 100 m
•
76°
74°
73°
72°
71°
70°
69°
68°
Fig. 6. Isotherms at 100 m.
17
1993
S 100 m
•
35,0
70'
F19. 7. Isohalines at 100 m.
18
5°
10°
15°
40°
1993
t"C 200 m
•
7'5°
74°
71°
69°
68°
67°
Fig. 8. Is otherms at 200
ffi.
19
10°
20°
15°
25°
35°
30°
40°
79°
1993
•
S 200 m
78°
I
r
i
77°
76°
75°
I
74°
34.9
72°
71°
/
70°
•
69°
68°
)
\J
73°
o
~
~1~-~
~.,. ~o
~ 34.~
•.
;Ja~~:
i{'
/;~
. ~
67°
Fig. 9. Isohalines at 200 m.
\
.
~~
,/
,;-34.9
20
30"
3S"
40"
79"
1993
tOC at bottom
78"
•
77"
76"
7S"
71"
71"
70"
•
69"
F·le>'
u 10. Isotherrns at the
bOttOffi.
21
10°
S°
20°
15°
30°
15°
35°
40°
45°
50°
SOO
~.
1993
S at bottom
•
77°
\
I
3~
>;01.. off!'
75°
I
\
'-
(3r
\~'~:~
73°
72°
71°
70°
•
I
34.9
34.9
76°
74°
.
69°
68°
67°
Fig. 11. Isohalines at the
bOttOffi.
35.0
'j'j0
22
Lat
74°~'
N
A
l.rg OSo 14' E
0~::;;==-~~;;;;;~~;;;~~~7""'""
SO
100
150
200
300
400
500
600
,
800
\
•
-os
'- ,
1000
Lat
74°30' N
B
l.rg 05° 14' E
Or--l----rT"""'T'"........,---r--.----v---,--,......,-r---r-~~..._r.."".,,..,_rr---,
SO
100
150
200
300
•
400
500
600
(
34.9
1300
\
' ' """349
\
1000
Fig. 12. Hydrographie seetion Bear Island - West.
Temperature (A) and salinity (B).
23
Lat 71.°11; N
Lng 19"20' E
A
o
50
6
100
\/
150
I.
200
250
300
•
400
500
Lat 71.°1' N
Lng 19°2O'E
0
71°26'
25°29'
B
50
100
•
150
(j,
35.0
200
250
300
1.00
500
Fig. 13. Hydrographie seetion Nonh Cape"- Bear Island.
Temperature (A) and salinity (B).
24
Lat 76°~' N
Lng 33"~' E
69"~'
O~==---\T"""--r------'---.-.-----r
50
i
i
i
.......
9:;;;
::.!..::::::====:::::::==:::::::===:::~
4
33"~'
i~
~
4_ _ _ _
100
150
200
250
300
•
69" 30
8
33° 30
8·
34.0 _ _-33-
_----:;=:::---
50
____-
----..........
- - - - - 34.5----....
~34o-
•
Fig. 14. Hydrographie seetion along the Kola meredian.
Temperature (A) and salinity (B).
25
La t
72" 30' N
Log
4'J> 15' E
68" 45'
43" 15'
A
o
50
100
150
~2
\
200
•
250
300
Lat
72°})' N
Log
4r1s' E
B
o
I~-'C-_
~_____
50
•
100
150
34.7
,-- ..... \
,
I
I
I
I
,
I
\
'\
34.95
200
"' "
"
'\
\
,
\
,
I
250
34.0
34.5 - - - ~
34.6
300
Fig. 15. Hydrographie seetion Cape Kanin - Nonh.
Temperarure (A) and salinity (B).
26
5-
10-
15-
2eT
25-
30-
35-
40"
45"
5eT
8eT
55"
":Li
79~
1993
O-Group Herring
~ 1-1000 fish pr. n mile
P-
m> 1000 fish
7ff'
pr. n. mile
A
77
76-
•
I
6
A
6
A
A
A
-.
.,
A
•
6
6
•
6
A
75"
74-
6
•
A
A
A
73"
6
A
A
A
6
A
A
A
72"
6
A
71"
A
A
A
A
A
A
AA
6
70"
6968-
6T
Fig. 16. Distribution of O-group herring.
A
A
66
6
6
6
A
A
A
A
A
A
A
6
6
A
•
27
5'
80'
78'
77
•
6
..
6
A
6
6
6
6
.. ..
6
6
..
6
..
6
6
6
6
6
6
6
6
A
6
& 6
6
6
..
6
6
6
6
6
..
6
6
6
6
..
6
6
6
~
6
6
6
6
...
6
6
6
...
6
..
6
6
6
6
6
6
66
6
6
A
6
6
A
6
A
6
73'
6
..
.~
6
6
6
72'
6
A
A
A
f}&
...
66
6
6
..
6
6
6
6
6
6
6
6
6
6
6
6
6
6
...
6
...
6
6
6
6
6
6
..
6
...
6
6
66
70'
69'
68'
67'
Fig. 17. Distribution of O-group capelin.
6
6
6
6
66
6
6
6
•
45'
6
6
71'
40'
50'
1993
O-Group capelin
~ 1-1050 fish pr. n. mile
;:>-
6
75'
74'
35'
A
.
A
6
30'
.' ..Ll
6
6
76'
25'
20'
6
6
A
79'
15'
10'
...
6
66
6
6
6
6
6 ...
6
6
6
6
6
55'
28
5·
15·
10"
20"
25·
30"
35·
40"
45·
50"
55·
":a
80·
79·
\
P-
1993
O-Group Cod
~ 1-85 fish pr. n. mile
m > 85
7fi
fish pr. n. mile
77
•
7ö
75·
A
74·
A
A ..
A
A
A
A
73·
72·
71·
6
6
70·
69"
68"
67
Fig. 18. Distribution of O-group eod.
e·
29
580"
10-
15-
20"
25-
30"
•
7ö
75-
P-
• • •
• • •
• • •
• •
1993
O-Group Haddock
~ 1-85 fish pr. n. mile
III >
85 fish pr. n. mile
~
•
.••
•
73"
727170"
69·
•
50"
. .• •
~
74-
45-
.. ~
~
77
40"
• •
79-
7fi
35-
68"
67
Fig. 19. Distribution of O-group haddock.
.
• • •••
" •
• • " •
" •
•
"
".
"". ••"
"
•
"
~.
• •
• "
.
• • •
• "
55-
30
5"
10"
20"
15"
25"
40"
35"
30"
45"
50"
55"
.,~
80"
79"
1993
O-Group saithe
~ 1-85 fish pr" n. mile
r:>-
m > 85
78"
fish pr. n. mile
77
6
76"
6
75"
6
6
74"
• • • •
• • .~ •
• •
• • • • ••
6
• • •
6
6
•
6
6
6
6
6
•
6
73"
Q
•
6
•
• •
6
6
• @
6
• •
6
72"
6
.6
•
71"
6
6•
6
6
6
6
•
6
6
•
6
• •
6
6
6
6
6
6
6
-6
• •
• •
• •
6
6
6
6
6
6
6
6
6
70"
69"
68"
67
Fig. 20. Distribution of O-group saithe.
6
6
6
•
•
6
6
6
6
6
6
•
•
6
6
•
•
.6
6
6
6
6
6
6
6
6
6
•
6
6
6
6
6
f;)
6
6
@
6.
6
•
6
•
6
6
6
6
6
6
6
6
6
6
•
6
6
6
.6
6
6
6
6
•
6
6
6
•
•
•
31
15"
30"
35"
45"
40"
50"
.. ~
80"
1993
79"
\
O-Group Polar cod
~ 1....;110 fish pr. n. mile
E > 110 fish pr. n. mile
~
7fi
6
6
77 ~
•
6
6
70
6
6
..
. .
.
..
.
.
. .. ..
..
.
. . ...
6
6
..
6
6
6
6
72"
71"
70"
. .. .. .. .. .
.. ..
..
.. .. . .. ... .. .
.. .. .. .. .. .. .. . .
.. . .. .
....
6
6
6
6
.. .. .... .. .. ..
..
..
..
..
69"
•
68'
67
Fig. .21. Distribution of O·group polar eod.
-;t
..
..
.. .. ..
. .. ..
.
. ..
.. ..
......
6
6
6
6
73"
6
6
6
75"
74"
. ..
6
..
.
.... ..
..
.
..
..
...... .
..
..
.. ..
..
......
..
..
.
. ....
..
..
55"
1----32
5"
10"
20"
15"
30"
25"
35"
40"
45"
50"
55"
... ~
80"
1,993
79"
O-Group Redfish
~ 1-85 fish pr. n. mile
po~
~ > 85 fish pr. n. mile
78"
77
..
76"
75"
74"
73"
72"
..
.. ..
@
I
A
.. ..
"
. . • ..
. .. ..
.. • . ....
•. ..
.. "
..
.. . .. ..• " ..
A
A
.. ,. ,.
A
71"
,.
70"
"
,.
.. ,.
,. ,. ,.
,. ,. ,.,.
,. ,. ..
,. ,.
...
,.
,.
..
..
.
.. .. ..
•
.. •
.. ..
.. ..
.. ..
.. ..
.
. ..
,.
,.
A
A
•
... ....
.. ..
.. .... ..
.
..
,.
,.
•
. • " ..
. ..
.. •,. ,. "
..
..
,. ,.,.
,." ,. ,.
,. ,.
,."
",. ,. ,."
.,. ,.• • ,. "
,. ,. •
,.
,.
A
69"
68"
67
Fig. 22. Distribution of O-group redfish.
•
33
5"
15"
10"
20"
30"
25"
35"
40"
45"
50"
.. ~
80"
79"
1993
O-Group Greenland halibut
~ 1-85 fish pr, n. mile
pr>-
111
78"
>
85 fish pr, n. mile
77
•
6
6
.A
6
6
6
6
6
6
6
~
•
6
6
76"
6
11
6
75"
73'
6
A
A
74"
A
•
A
•
•
6
6
A
A
••
•
6
A 6
• •Q
6
11
A
6
6
• •
A
A
• •
• • •
6
A
•
6
A
A
A
•
•
6
6
6
A
72"
6
A
6
6
•
A
A
A
A
A
6
6A
A
6
6
A
A
•
6
•
6
11
6
A
•
•
••
•
A
A
A
A
6
A
A
71"
•
A
A
6
•
A
11
A
A
•
A
A
6
6
6
A
A
A
A
A
A
6
6
6
6
6
A
6
•
A
A
A
6
6
•
A
-6
A
A
68'
67
Fig. 23. Distribution of O-group Greenland halibut.
AA
A
6
A
A
A
6
6
AA
A
A
A
A
A
•
•
6
A
A
6
6
A
•
• •
A
f&.
AA
A
A
•
A
69'
• •
•
A
A
70"
•
•
A
•
55"
A
A
6
A
A
34
10"
5"
80"
79"
77
•
•
•
p~
•
A
~ 1-85 fish pr. n. mile
• • •
•
• •
• •
A
A
•
•
•
•
•
•
A
A
6
•
••
• • • •
A
• • •
• • •
• ••
6
A
A
• •
• • • • • • • • •
• •• • • •
•
•
• ••
• •
•
• •
••
••
•
A
A
6
A
71"
55"
6
A
72"
50"
• •
6
73"
45"
40"
1993
O-Group Lang rough dcb
A
A
•
74"
•
• • •
75"
35"
,)~
6
•
•
76"
30"
25"
A
A
•
78"
20"
15"
A••
6
A
6
70"
• •
•
,.
A
•
•
••
• ~ •
•
•• •
@ •
•
• • • • •
6
A
69"
68"
67
Fig. 24. Distribution of O-group long rough dab.
•
35
5'
10'
80'
78'
77
•
.
A
A
A
A
A
A
A
A
•
A
A
A
75'
A
A
.
A
A
•
70'
A
A
A
• •
•
A ..
A
A
Q
A
A
• • ••
11.
..
.
A
•
11.
11.
..
•
11.
11.
• •••
11.
.6
6
11.
6
11.
• .."
A
A
11.
11.
11.
A
11.
A
11.
A
6
11.
11.
A
11.
6
11.
•
~
•
6
6
•
69'
•
50'
~
O-Group Sandeel
1-85 fish pr. n. mile
fli
> 85 fish pr. n. mile
..
..
..
A
-.
A
A
A
A
A
A
11.
A
11.
A
11.
A
•
A
A
A
11.
..
A
A
A
•
•
11.
A
11.
11.
11.
A
A
A
A
A
I
A
• • •
A
A
A
72'
..
A
.. ..
. .. . .. ..
.. .. .. ..
.. ..
.
A
45'
1993
P-
A
76'
73'
40'
35'
.,~
A
A
74'
30'
25'
20'
• •
•
79'
15'
68'
67'
Fig. 25. Distribution of O-group sandeel.
11.
11.
6
A
11.
11.
11.
®
6
11.
611.
A
11.
All.
.
11.
A
611.
~
A
55'
36
5"
15"
10"
20"
25"
•
30"
45"
40"
35"
55"
80"
1993
79"
O-Group Catfish
~ 1-85 fish pr. n. mile
pt>-
78"
77
•
• •
• •
• • •
• •
6
76"
6
75"
6
74"
•
•
• • ••
6
•
6
&
•
6
•
•
•
6
6
&
6
&
6
6
6
•
6
6
6
6
6
6
6
6
&
6
6
6
6
6
6
6
•
6
6
6
6
6
•
6
6
6
6
6
•
6
6
6
6
6
&.
6
6
6
&
6
6
6
6
6
-I.
•
• •
6
•
6
6 6
•
6
6
•
~
•
6
6
6°
6
&
&
71"
•
:
73"
72"
~.
• •
6
•
6
•
6
6
6
6
70"
6
&
6
6
6
6
&
6
6
6
6
6
6
6
6
~
6
6
6
&
6
66
&
6
6
6
6
6
6
6
6
6
6
6
&
6
6
6
6
6
6
6
6
6
66
6
6
6
66
•
6
6
6
6
6
&
69"
68"
67
""'h....,...-,,............,,...,.-......-.......,::..T""""'"""..-.-....-......................"'T""'T"....................- . - .............-.,............,.................,l-.........-~.,I....-+--r-...........~...............-l-
Fig. 26. Distribution of O-group catfish.
•
..
37
5"
10"
15'
25'
20"
30"
35"
40"
45"
50"
..
80"
~
1993
79"
O-Group Gonctus
p~
~ 1-85 fish pr. n. mile
78"
77"
f
•
• •
•
7S"
6
6
6
75"
74"
73"
72"
71"
• • •
• • • • •
.0
•
• Ci:) •
6~·
• 6
• ••
•
•
6
• • 6 @. •
• •
• 6 • ••
• • •
• 66
6
• • ~ ••
• • a.
•
6
70"
•
•
•
•
6
~.
&.
•
•
6
•
• •
•
6
6
6
6
6·
"
S9"
•
6
• •• • •
• • • •
•
• •• • • • • • • • •
• •
•
• • • •
• •
•
•
•
• •
•
• •
•
• • • • •
•• •
••
S8"
S7"
Fig. 27. Distribution of O-group Gonatus fabricii.
.6
•6
•
•
•
•
•
•
• • •
• • •
55"
APPENDIX
Research
vessel
Participants
"Professor
Marty"
S.Boychuk, V.Donetskov, Yu.Garbut, A.Gordov,
S.Ratushny, E.Shamray, V.Tretyak
"Pinro"
N.Chebotok, P.Duzin, E.Gusev, S.Hardin,
A.Krysov, F.Minyazov, G.Polyakin
"Johan Hjort"
B.Bergfll2ldt, I.Fjellstad, M.Fonn, Ä.Fotland,
H.Grcesdal, H.Larsen, K.Lauvas, S.Mehl,
M.Mjanger, A.Ncess, A.Romslo, L.Solbakken,
R.Toresen, B.Kvinge
"G.O. Sars"
M.Dahl, E.Holm, A.Hylen, S.Iversen,
L.Kalvenes, J.de Lange, M.Ml2lgster., 0. Ncevdal,
V.Ozhigin (PINRO) , R.Pettersen, A.Raknes,
N.Ushakov (PINRO) , E.0vretveit
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