• (j/o International Council for the Exploration of the Sea C.M. 1992/G:82 Ref.H Demersal Fish Committee PRELIMINARY REPORT OF THE INTERNATIONAL O-GROUP FISH SURVEY IN THE BARENTS SEA AND ADJACENT WATERS IN AUGUST-SEPTEMBER 1992 The twenty-eigth annual International Q-group fish survey was made during the period 12 August - 8 September 1992 in the Barents Sea and adjacent waters. The following research vessels participated in the survey: State Name of vessel Period Norway Norway Norway Russia Russia Russia "Johan Hjort" "G.O. Sars" "Michael Sars" "Professor Marty" "Fridtjof Nansen" "Akhill" 17.8 18.8 13.8 17.8 24.8 13.8 5.9 - " - - 3.9 7.9 7.9 28.8 - 5.9 15.8 - . 6. 9 - Research Institute Institute of Marine Research, Bergen --"-- --"-- The Polar Research Institute of Marine Fisheries and Oceanography, Murmansk Names of scientists and technicians who took part on the different vessels are given in the Appendix. ... ~. prlelimdina: y analysti~ of8thgeSSUtrvmbey da~a wHere PlfanntedS~o take pace urlng a mee lng ep e er ln ammer es. lnce none of the Russian vessels were able to call at Hammerfest data were analysed at IMR, Bergen and PINRO, Murmansk and the results exchanged by correspondence (telefax and telemail) . Observations concerning the geographical distribution of Q-group fish and their abundance are given in this report together with abrief description of the hydrographical conditions in the area. 2 MATERIAL AND METHODS The geographical distribution of O-group fish were estimated with a small mesh midwater trawl. The vessels which participated in the survey in 1992, used the type of midwater trawl recommended by the meeting. held after the survey in 1980 (Anon., 1983) .' The trawling procedure was standarized in accordance with the recommendation made at the same meeting. At about every 30 nautical miles sailed the trawl was towed in several depths in one haul. The standard procedure consisted of towings of 0.5 nautical mile in each of 3 depths with the headline of the trawl located at 0, 20 and 40m. An additional tow at 60 and 80m for 0.5 nautical mile was made when O-group fish layer was recorded on the echosounder deeper than 60m. Survey tracks and hydrographical stations are given Trawl stations with and without catch are indicated . distribution charts in Figs. 14 - 23, as filled and respectively. The density grading is based on catch per 1.0 nautical mile trawling. in Fig. 1. on the open symbols in number HYDROGRAPHY Observations were made along all the survey tracs with 5 to 40 nautical miles between stations. Horizontal distribution of temperatures and salinities is shown for 0, 50, 100 and 200m (Figs. 2-9). Figs. 10 - 13 show the temperature and salinity conditions along the Kola, Bear Island - North Cape, Bear Island - West and Cape Kanin sections. The mean temperatures in the main parts of these sections are presented in Table 1 . • , I It appears that mean sea temperatures from 0 to 200m were high and weIl above (0.4°C to 0.7°C) the long term average in all parts of the surveyed area, and 1992 is the fourth "warm year" in succession. The positive anomalies were mainly caused by large contents of heat in intermediate and deep layers and thus assosiated with watermasses of Atlantic origin flowing into the Barents Sea from west. Surface layer temperatures (0-50m) in the central and southeastern parts of the area were significantly lower in 1992 than in 1991 and just slightly above the longterm average. 3 DISTRIBUTION AND ABUNDANCE OF O-GROUP FISH AND GONATUS FABRICII Geographical distribution of Q-group fish are shown as shaded areas in Figs. 14 - 22, and of Gonatus fabricii in Fig. 23. Double shading indicates dense concentrations. The criteria for discriminating between dense and ~cattered concentrations are the same as used in earlier reports (Anon., 1980). Abundance indices, estimated as the area of distribution with areas of high densities weighed by 10, are given in Table 2. Another set of abundance indices are given for O-group herring, cod and haddock (Table 3) as described by Randa (1984) ~ These are based on the nurnber caught during a standard trawl haul of one nautical mile. Length frequency distributions of the main species are given in Table 4. _ \.. l!erring The distribution of herring was similar to that of 1991 inthe Barents Sea, but with fewer observations along West-Spitsbergen. The main concentrations were found in the Western Barents Sea along the edge of theshelf between the Norwegian coast and up to South Cape (76°N), and~eastwards as far as 50 o E. The logarithmic abundance index is estimated at 1.06, which is only slightly below the 1991 index (1.19) and indicates that the 1992 yearclass is relatively strong. Capelin • I (Fig.14) I (Fig.15) There were very few observations of Q-group capelin and dense consentrations were observed at one station only in the southeastern part of the Barents Sea. The overall catch in numbers is among the lowest ever recorded since the O-group investigations started in 1965 and the 1992 yearclass seems to be very poor. Cod .(Fig .16) O-group cod had one of the widest distributions ever observed, similar to that of 1991, extending from Norway and Murman coast to at least 80 0 30'N off West-Spitsbergen (into the drift ice) and to 77°N in central parts. In the east it was recorded north to 73°N and east to Novaya Zemlya (52°E) . The abundance indices for the 1992 yearclass are the highest in the time~series and weIl above both the 1983, 1985 and 1991 yearclasses. At this stage the 1992 yearclass must be classified as a very strong one. 4 (Fig.17) Haddock Haddock was, as in 1991, mainly distributed in western areas from the Norwegian coast to 80° off West-Spitsbergen. The eastern limit in the Barents Sea was ar about 42°E. Dense consentrations were found in a smaller area than in 1991, mainly north of Cape North to 74°N. The abundance indices is the second highest ever observed, and the 1992 yearclass may be classified as rich. Saithe (Fig.18) Unlike in previous years, O-group saithe occurrded in many hauls in most of the survey area. This may indicate good saithe recruitement in 1992. • Polar cod (Fig .19) O-group polar cod is distributed in two separate areas, with one cornponent west and southeast of Spitsbergen and one component in east along the western coast of Novaya Zemlya. During the 0group survey the total area of distribution is not cornpletely covered. In the north-western area, the distribution was wider than in 1991, mainly off West-Spitsbergen, and the index about twice that of 1991. The 1992 yearclass of polar cod in this area seems to be at least of average abundance. • In the southeastern area along Novaya Zemlya the distribution was similar to that of 1991, but the area with dense concentrations was smaller and the index only half of that in 1991. The 1992 yearclass in this cornponent of the polar cod stock may therefore be considered to be of average strength or slightly below average . I Redfish (Fig.20) The rnain distribution of redfish was along the western edge of the shelf, from 73°N to 80 0 30'N off West-Spitsbergen. The area of both scattered and dense concentrations weresmaller than in 1991 and the index 25% lower, the lowest since 1968. The 1992 yearclass of redfish may therefore be considered as poor. Greenland halibut (Fig.21) Only single fish occurred in hauls west and east of Spitsbergen and, as in the previous four years, the abundance index indicates that the yearclass is very poor. 5 Lang rough dab (Fig.22) This species occurred mainly souteast of Spitsbergen. The abundance index is the lowest recorded since 1970 and the 1992 yearclass is considered to be poor Blue whiting, sandeel and catfish. Only a few specimens of these species were caught this year. Gonatus (Fig.23) Specimens of O-group Gonatus fabricii were found over a larger area than in 1991. The distribution is westerly, from the Norwegian coast at 28°E and west into the Norwegian Ocean and north to about 80 0 N off Western Spitsbergen. REFERENCES Anon., 1980. Preliminary report of the International O-group fish survey in the Barents Sea and adjacent waters in August/September 1978. Annls biol., Copenh., 35:273-280. Anon., 1983. Preliminary report of the International O-group fish survey in the Barents Sea and adjacent waters in August/September 1980. Annls biol., Copenh., 37: 259-266. Randa, K. 1984. Abundance and distribution of O-group ArctoNorwegian cod and haddock 1965-1982. Pp. 189-209 in God0,O.R. and Tilseth,H. (eds.): Reproduction and recruitment ofArctic cod. Proceedings of the first Soviet-Norwegian symposium, Leningrad, 26-30 September 1983. Institute of Marine Research, Bergen, Norway. Tereshchenko, V.V. 1992. Some results from long-term oceanographie observations during O-group surveys in the Barents Sea. ICRS CM 1992/C:18. Toresen, R. 1985. Recruitment indices of Norwegian springspawning herring for the period 1965-1984 based on the international O-group fish surveys. ICES CM 1985/H: 54. 6 Table 1. Mean water temperature 1 in main parts of standard sections in the Barents Sea and adjacent waters in august-September 1965 - 1992. " ,. >" -,.' ". , Year " ....',., 1965 1966 1967 1968 1969 1970 1971 1972 1973 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988 1989 1990 1991 1992 It I Aver. 19651992 . . 1) 2) ,'," 2 ,'" 0-50 50-200 6.7 6.7 7.5 6.4 6.7 7.8 7.1 8.7 7.7 . 8.1 7.0 8.1 6;9 6.6 3.9 2.6 4~0 3.7 3.1 3.7 3.2 4.0 4.5 3.9 4.6 4.0 3.4 2.5 2.9 3.5 2.7 4.0 4.8 4.1 3.5 3.5 6~5 7.4 6.6 7.1 8.1 7~7 7;1 7.5 6.2 7.0 8.6 8.1 7.7 7.5 .. "'.' . ." C o · · • • • •, .,. 3 • A' , " , < , ,. I 0-200 O-bot~ 4.6 3.6 4.9 4~6 "., ,""" 4~4 4~0 4~7 4~2 4~9 3.7 4.8 4.4 4.5 4.6 3.7 4.6 4.1 " 3.7 2.2 3.4 2.8 2.0 3.3 3.2 4.1 4.2 3.5 3.6 4.4 2.9 1.7 1.4 3.0 2.2 2;8 4.2 3.6 3.4 3.2 2.5 2.9 4.3 3.9 4.2 4.0 5.1 5.5 5.6 5.4 6.0 3.6 4.2 4.0 4.2 5.3 5.8 6.3 5.9 5.3 5.8 5.2 5.5 6.9 6.3 6.0 .6.1 . .,. 3.2 5.7 2~0 3~3 4~5 "" ..... . 7 , 0-200 5.7 4.6 5.6 4.9 4.1 2.4 5.2 5.0 4.3 3.6 3.8 4.5 3.7 4.8 5.6 5.0 4.4 . .-, 0-200 5~1 5.2 5.3 '. 6 •., .O-:-bot ~ L9 6.1 4.7 2.6 4.0 4.0 4.0 4.5 5.8 5.3 5.3 5.3." ,.. "" 5 ".4. , 2.7 4.5 5.1 4.5 3.4 3.9 2.7 3.8 6.5 5.0 4.8 5.0 3~3 7.3 .......... , Section 2 .' and layer,:.(deep .in.meter). .... . . ,1 '. , ".,.-,"', .... . - 6~1 - 5.7 4.2 3.9 5;0 4.9 4.9 4.8 4.0 4.1 6~3 5;9 6~1 5.7 5.6 4.9 5.0 5.3 4~4 4.9 4.4 4.9 5.1 5.0 4.6 4.4 3.9 4.2 4;9 5.7 5.4 5~7 ",,"" I···· 5~0 4.5 ., ", ... " Earlier presented temperatures have been slightly adjusted (Tereshchenko, 1992). Murmansk Currenti Kola Section (70 0 30'N-72°30'N,33°30'E) Cape Kanin section (68°45~N - 70 0 05'N,' 43°15'E) Cape Kanin section (71°00'N - 72°00'N, 43°15'E) North Cape Currenti.North Cape - BearIsland section (71°33'N,25°02'E - 73°35'N,20046'E) 7: West Spitsbergen Currenti Bear Island.- West section (74°30'N, 06°34'E - 15°55'E) 1-3: 4: 5: 6: 7 Tabell 2. Abundance indices of O-group fish in the Barents Sea and adjacent waters in 1965 - 1992. Year • , 1965 1966 1967 1968 1969 1970 1971 1972 1973 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988 1989 1990 1991 1992 Cod 6 1 34 25 93 606 157 140 684 51 343 43 173 106 94 49 65 114 386 486 742 434 102 133 202 465 766 1159 Haddock 7 1 42 8 82 115 73 46 54 147 170 112 116 61 69 54 30 90 184 255 156 160 72 86 ·112 227 472 313 Polar cod West I East 0 129 165 60 208 197 181 140 (26) 227 75 131 157 107 23 79 149 14 48 115 60 111 17 144 206 144 90 195 70 144 302 247 73 50 39 16 334 366 155 120 41 48 239 118 Redfish 159 236 44 21 295 247 172 177 385 468 315 447 472 460 980 651 861 694 851 732 795 702 631 949 698 670 200 150 Greenland halibut Long rough dab 1 1 8 3 13 21 16 9 35 22 12 38 17 16 40 36 55 41 8 5 2 1 3 66 97 73 17 26 12 81 65 67 83 113 96 72 76 69 108 95 150 80 70 86 755 174 72 92 35 28 32 8 Table 3. Estimated logarithmic indices with 90% confidence limits of year class abundance for O-group herring, cod and haddock in the Barents Sea and adjacent waters 1965 - 1992. .. .' ,', Herring 1 Year . • I 1965 1966 1967 1968 1969 1970 1971 1972 1973 1974 1975 1976 1977 1978 1979 1980 1981 1982 1983 1984 1985 1986 1987 1988 1989 1990 1991 . 1992 1) Index " 0.14 0.00 0.00 0.01 0.00 0.00 0.00 0.05 0.01 0.00 0.00 0.01 0.02 0.09 - 0.00 0.00 1. 77 0.34 0.23 0.00 0.00 0.32 0.59 0.31 1.19 1.06 Confidence .limits 0.04 0.31 0.00 0.04 - - 0.03 0.01 - - - 0.08 0.01 - 0.03 0.00 0.01 0.01 0.05 0.20 0.20 0.18 2.33 0.52 0.28 0.00 0.16 0.49 0.16 0.90 0.69 0.03 0.53 0.76 0.50 1.52 1.50 1.29 - Haddock Cod - - Index + 0.02 0.04 0.02 0.25 2.15 0.77 0.52 1.48 0.29 0.90 0.13 0.49 0.22 0.40 0.13 0.10 0.59 1. 69 1.55 2.46 1.37 0.17 0.33 0.38 1.23 2.30 2.94 Confidence limits Index 0.04 0.08 0.04 0.34 3.05 1. 01 0.72 1. 82 0.42 1.17 0.01 0.08 0.00 0.29 0.64 0.26 0.16 0.26 0.51 0.60 0.38 0.33 0.12 0.20 0.15 0.03 0.38 0.62 0.78 0.27 0.39 0.10 0.13 0.14 0.61 1.17 0.87 0.01 0.02 0.01 0.17 2.02 0.57 0.35 1.18 0.18 0.66 0.06 0.36 0.14 0.25 0.08 0.06 0.43 1.34 1.18 2.22 1.06 0.01 0.22 0.30 1. 04 1.97 2.53 0~22 0.65 0.32 0.59 0.18 0.18 0.77 2.08 1. 98 2.71 1. 70 0.40 0.47 0.48 1.34 2.65 3.39. ',' Assessment for 1965-1984 made by Toresen (1985). Confidence limits 0.00 0.03 0.00 0.20 0.42 0.18 0.09 0.15 0.39 0.40 0.24 0.21 0.07 0.12 0.10 0.00 0.30 0.48 0.60 0.23 0.28 0.00 0.05 0.10 0.48 0.98 0.71, 0.03 0.13 0.02 0.41 0.91 0.36 0.27 0.40 0.68 0.85 0.51 0.48 0~19 0.28 0.20 0.05 0.52 0.77 0.99 0.31 0.52 0.25 0.34 0.20 0.75 1.37 1. 06 9 Table 4. Length distribution of O-group fish 1 in percent in the Barents Sea and adjacent waters in August - September 1992. Length (mm) 15-19 20-24 25-29 30-34 35-39 40-44 45-49 50-54 55-59 60-64 .65-69 70-74 75-79 80-84 85-89 90-94 95-99 100-104 105-109 110-114 115-119 120-124 125-129 130-134 135-139 140-144 • 1) Herring Capelin Cod Haddock + + + 0.1 0.7 2.4 6.1 12.0 14.5 14.8 13.7 12.5 10.7 6.7 3.7 2.0 0.2 0.1 33.3 45.5 9.1 3.0 3.0 3.0 3.0 + 0.1 0.7 2.0 5.1 8.1 11.3 16.8 20.4 18.0 9.8 4.4 2.0 0.8 0.1 0.2 + + + 0.1 0.4 1.0 2.0 5.2 8.0 12.1 13.0 11.5 10.6 9.8 10.1 5.8 4.4 2.5 1.7 1.1 0.6 0.1 Polar cod East 6.3 6.3 22.6 25.0 31.3 7.8 0.9 .West 0.1 0.8 6.3 16.2 31.5 32.0 8.8 3.7 0.6 0.2 0.1 Redfish Green land halib Long roug dab 7.3 7.3 12.5 0.6 1.7 14.9 40.5 39.3 3.1 + 0.3 0.9 12.2 53.1 30.1 3.1 0.3 Sandeel 2.3 29.3 36.1 27.1 4.5 2.1 35.4 17.7 17.7 0.8 Total numbers 162002 33 179354 3795 116 68621 80474 8 169 Mean length 90.0 44.0 80.2 94.3 46.8 38.7 43.1 68.5 38.3 Based on Norwegian observations only 132 47.5 10 10° 5° 15° 40° 35° 30° 25° 20° 45° 55° 50° ~z-.z-i 80° \ Z 79· j_z. z-z \ r z~ 1 Z I Z 12 AUG - 8 SEP ..1992 p.,. • Z CTO Sonde station X NANSEN BOTTlE 7" I z - z - z2-/ 1 I \ Z z-z-z~z..\. 77"1 Z ,z-z-z-z z . -z-~z-z-z-z-z-z 76· . z-z,> I z z-z z.......... / z - z - ! z I z z I I I z-z-z-z-J z-z-z-z-z \ z z Z-Z-z-z--J z-z-z-z-z \ \ z z-z-z-z-z Z-Z-'I-I-1 \ I z z-z-z-z z-N-Z-Z-Z-Z-1-Z-~~Z-l-1' HJ Q \ " i,~ - z - z - z \ . Z- 1 - Z-Z-Z \ 1. 1 " '1 <z Z.., Z-Z z" 2-Z-Z-1--1Z Z I "Z Z I \ I Z-z l-Z-Z z-i-Z-z I " \. \ I \ Z-Z I IJ 1 ",. r ,,"z 1,,\ Z\,Z-l Z-I-~\ ~-z 71· 70· :.,M I I r z z 1I z z z z 1 I 1I I ~ z, z, Z Z z f I 1I 1-Z-Z--Z-Z Z 1 Z I 1 Z-Z-Z-Z I I I l Z-Z-Z \ Z-Z-Z-Z I I1 ZI Iz J I "I I I z IZ I Z I Z Z I 1 f f Z,,, Z-Z-Z-Z Z Z I z Z " Z I I \, -Z I I I,Z, ~-z-Z' Z lZ--;:::a..-l "" 1 I 1 Z ZZ \....Z-Z Z-, ~ 'Z cl 'z . ' ~ \ z_~.\4; 72· z-z Z Irr I 75° • I I I z I II I 1 ZX U IZ Z ZI ~ 1I z iJ;(1 1 I I 2X I I 1 Z Z L ~ I Z~f ~f 69" 68· Fig. 1. Survcy tracks and hydrographie stations. . I Z f I I z-Z l-fl' . 2 I Z I Z I Z I z I nt t1f I l-Z I ,-Z 1 1 Z + / I I Z--~Z r Z- z Z-Z-Z 1 1 Z-Z, \ '0 X 1-2+-Z-Z-Z L r-l z-+Z-Z...J Z-Z-Z-Z-Z-Z Z -Z-Z~ I Z-z-z ,Z-f l-Z Z Z Z Z - - - l i rl ZI",,-Z_.l 0U 11 50 20" 25 0 35 0 'lO0 55 0 400 .c;;9 1992 toC 0 m ~ 75 0 ,:'~ ,, 74 0 5.../ 6 73" 72 0 • 71 0 \S 70 0 69" 68 0 670 Fig. 2. Isothenns at 0 m. 600 12 1992 SOm • 67 0 ..... ~--,-~. Fig. 3. Isohalines at 0 m. i i-----r--1' ~-I~ I i • i i .l, i i I 13 So 10° 1So 20" 2So 30° 35° Sso 40° ~ 1992 ~ 73" 71° • 71° 70° 69" 68° 67° Fig. 4. Isothenns at 50 m. tor 50 m 60° 14 5° 10° 15° r 25° 30° 35° /000 45° ,c;;9 eo- 79° 1992 S 50 m p.., e 350/ 75° 1" 14° 730 72° e 71° 70· 690 68· 67· Fig. 5. Isohalines at 50 m. ')00 55° 15 'j0 10° l'.i0 200 2'j° 35° 30° 40° t,'j0 SOO 'j'j0 ~\) d9 ~ Fig. 6. Isothenns at 100m. ~ .9..(\ 1992 tOC 100 m . 60° üg~~ ~Q 16 1992 S 100 m ~ 35.0 73" 69"L~~~~ 68° 67° P·19. 7• Isohalines at 100m. 17 1992 tO( F19. · 8. Isothenns at 200 m. 200 m 18 SO lSo 10° 20" 2So 3S 0 30° 40° 4So SOG Sso 60° . ()g~O[j?; ß~ (, 80° - .c:;9 79° Ii> ~ .9.(} . Q ~ 1992 S 200 m 78° .~ , I 35.0 76° 7So 74° :> Q 73° 35.0 ~ U ( 72° 34.9 I e 9 71° 10° 69° 68° 67° Fig. 9. Isohalines at 200 m. ~ r . L 19 A 7 so -----S__----- 100 . 8~_""," 6~--------7-----S~ --6_ _ _ _ _ 150 200 250 300 Fig. 10. Hydrographie seetion along the Kola meredian. Temperamre (A) and salinity (B). 20 500 zoo 300 400 500 Fig. 11. Hydrographie seetion North Cape - Bear Island. Temperature (A) and salinity (B). 21 A lat 74°3ON lng06°34E SO 100 150 200 300 400 SOO 600 e 800 1000 B lat 74°30 N l"a 06°34E SO 100 150 200 300 400 SOO \ 34.9 J 600 800 1 34.9 1000 Fig. 12. Hydrographie seetion Bear Island - West. Temperature (A) and salinity (B). 74°30 18"30 22 A Lat 7t30N L"rP4115E 50 100 150 200 250 SO 100 '-----34. , ------... ......, I I ISO I 34.fs / I I ZOO I / / 250 Fig. 13. Hydrographie seetion Cape Kanin - North. Temperature (A) and salinity (B). 68"45 4145 0 ~ ,..., :},~ , , I I', , ....J...-_ , • , 9t , , l: , I , , , , , 0 <p q q Oö'"'t ........ q ~-r Ci a: ..... ~ :;> 0 S~ <1 q q q q q q q A ~ q q ~~ ~~ q q q = . 0 ~ <l ~q ~ ~ '"'" ~i q ... n. ~ q q :I: VI :I: ... I ci n. q <l q z l::l Z q q q <I q q :i M N :\? -Vy~ 0.' Q "" r!J4 ~ ~-l \.., . 't'~'~'1 <:... ' ~.,7 q q <l q~ q <I <I q q q <l q q :~tY <I <I . ~ q q iG~" ~ q 'q <I <I <I q <I q <I f q q ~ <I _ <I I :.~ - ~'\.~ ~'\.'\.~'\.,,*,,';''i0,90 l t bh 'e V ..cl Po. ::J ~ 0 I ca c:: ~~ -~ q - ~r <I - q q q <I q ..... <I q <I .q <I <I <I q q ~ ® <I ~ q q q :},~ ..... ~~ll<~ ~.~ 'CI • ~-\ \V~ f~ :~~:w ~<l .... ~q q 0 0 2 q <I <1 <I ...... ... 0 0 CD e ...... 0 ....., 0 ...'" 0 - q 0 ;! 'S 0 -.:i q ....bh <J q q ::J ,~ <J <I q <l '::I .c ... 0 m ....... 0 0 ;:: ... 0 C> • 0 .., <>- 0 .., CD ...... 0 ~ 24 5- 10A 10- '~ A p:;>., • A 77- A " " A 4 76- 6 756 '" 4 A 6 6 4 A ~ 6 6 6 6 A 6 6 6 6 6 6 A All. 6 6 i:>. i:>. 6 A 6 '" 6 A A A 6 6 6 ... 6 6 6 6 6 6 A 41> 6 6 6 6 6 A 4 A A A ...... 4 A A A 6 A A A A A A • 6 6 6 A 6 A 6 A A 4 A A 6 4 6 64 6 6 6 6 A ® 6 60 A A A 41> All. A -- A A 6 A '" .... " ... '" /@ 4 A ... A FISH PR. N.HILE ,. 1050 --11-- A ... ... 1 -1050 111 A A A 71° ~ A A 72° 0 li ROUP CAPflIN ~ "" 6 1992 A A A 73" JS- e::;9 79- 74- • 25- A A 7'- A '" A • • !S- A A 4 ·4 4 4 A A A 6 41> 6 6 6 A 6 6 '" 6 6 "... 4 • 4 4 441>41> ~~ 41> A 41> 41> A 6 A 4A 41> A A A A 4 A 6 6 A A A A A 4 A A 6 A A 't 70° A A A A~ A A 6 A A 69° 68° 67° Fig. 15. Distribution of Q-group capelin. 6 41> A A " A A A A 25 so 10° ISO 20" 2ljo 35° 30° SOO 45° 40" ,Q9 1992 • O·liROUP (00 p." • 79° ~ ~ ~ 78° /> 1 ·85 ,. 85 FISH PR. N.t1ILE -lI- 6 6 n° 76° 7'j0 666 />66 74° 6 73" 6 6 72° e 71° 70° 690 68° 67° Fig. 16. Distribution of O-group cod. 55° 60° O){:x>pp-eq dno.I2-0 JO uopnQ!l1sIO oLl o~H 089 r-, " .1J -" ' V vv v v vv 069 oOL v@ oLL v v v v e oZL v v V v v v V v v v oU V V v v o'L v V oSL vvv v v v v v v Vv v Vv v V v V v v ~o9L v v v V v v tou V v V V - 11- 31lW'N 'lid HS I~ S9 < S9· I ~ ~ .~ ll)OaaVH dnOll!l 0 Z661 f • ~ o6L (. ~(JV~~ • c:;9 I .09 I • Ir",...." i ~u oSS ( ( i .OS " >i i .S'I .0'1 i .S{ 9(: v ~ 0.1")1\.,,,_ i .O€ i .SZ i .Ol i .SI ~ v v v i .01 v , .S 27 10' S' 20" lS' 30' 2S' W 3'!)' 4S' SO' '::H> 6 4 ,69 80' 79' ~ ~ 6 .:. 7S' 6 A /1 A 0 \. 0 (j RO UP SAITHE ~o A 4 ():g~?ol]: ""Q 1992 6 6 • <I.(} O. . 6 6 60' SS' 1 - SS FI SH PR. N.MI LE > SS -,,- 4 n' /1 A A A /1 4 A A /1 A A 76' 75' A 6 74' 73" 6 A A '" e 71' A A 6 /1 A A A 4 6 A A A 72' A A 6 A· A 4 A .:> 6 A A 6 ~: 70' A A :~ A@A A 6 A A A 4 A .. A ~6 A 6 6A AA 6 6 0~ ~.. -, \ 69' '-,.I 6S' 67' Fig. 18. Distribution of Q-group saithe. 'po:) JUlod dno~-o JO uopnqy.nslO '61 ''sB oL9 089 069 ~.~ e v v v v v v v v .~. . "V V v v~ V V V 9 V 9 9 V V V 9 9 V V V V V V V 9 V V -9 V V V V V V V V V V V V V V VVV V V ~ I V V 9 V V 9 9 V 9 9 V V V V V V V V V V 9 V 9 V V V V V V V l-olL 99 V V 9 V f-o€L V V V V V " V V 9 V V V V •. V ~~ V V '9 ~oSL V 9 1-o9L , V V vV e V OLL< nil-rN 'lid HSI~ Oll· L ~ ~ l:}J ~{\'V ~ ,~ COJ lIV10d dOOll!l 0 & Z66L 009 .55 .os .s, , , oO't .S( 8Z ~lj I .GE I \\ '" .wß V !-oLL V V J08L WA06L ~~(JV~.~ ! &? V V 9 _ _ li _ _ l-o'lL V V V .~ .~. V V ~ oLL V V V 9 v 9 V V v V v v VV V I 9 V V V 9 9 V V V . v , i .Sl ~.~ ' , oOZ i .Sl .Cl oS e 29 SO ISO 10° ~O 20" 35· 30° # 80· 1992 e 79° O-GROUP REDFISH ~. ~ ~ ia 78° 76° I>. I>. I>. I>. I>. '" ~ I> I>. 1>.1>. I>. 74· I>. I>. I>. I>. I>. 6 I>. "'I>. I>. 6 "'I> '" 6 '" "'6 6 '" I>. 6 I>. I>. 6 I> '" '" '" I>. I>. 6 6 I>. I> 7'j. I>. '" '" '" '" '" '" '" '" '" '" '" '" '" '" '" '" '" '" 6 6 72° '" I>. '" I>. 66 6 I>. ~ 1>.'" I>. I>. I>. e 71° I>. I>. I>. '" • I>. 6 6 I>. -11- I>. '" 6 6 ~ I>. I>. I>. '" 6 6 '" A 70° 6 '" '" I>. A Fig. 20. Distribution of O-group redfish. 6 6 I>. 6 6 I>. I>. I>. .Il. 6 6 "0 I>. A A 68° '" '" I>. 6 .Il. 69" '" '" 6 6 I>. 6 I>. '" '" '" '" I>. I>. '" '" 1>.6 I>. I>. 6 > 8S I>. I>. I FISH PR. N.HILE I>. I>. 77° 7JO 1 • 8S I>. I>. I>. I>. '" '" A I>. 6 'lnqnuq PtmllIQQ.IQ dno~-o JO uopnq!JlsIO 'IZ 'Bld .L9 ~. ~ cl .~' V v v V "1"1 v v v V v V v v v v ., v v v v v v v v v '1 v v v v v v v v v v v v ."W;' 1 0 WP v V V V v v ;, ;f!. ."1 V v v v v v V v V v v v v v v v v v V V vvv v v v v V v v v v v v V v v v v v v v v v vvv v v v v v v HSI~ 58 • I ~ V V v v v V V v V V V V v v v V V v v v v V V V V v v V v v v v V v v Vv v v v v v V v .IL v v v .OL V v v v vv v v v v vv v v v v v Iv~ v v v V v V v v v v f-o€L v v V V v V V V V V v v v v v v -v v v v v v v v v v v v V V f-oSL h9L v v f-o'L v V V .~:. : ~ v f-oLL v v v v v V , olL 1 ·~.~-~~l· ~h.. · v Z66L v • ~~ J~ ,,'2 er' .55 .os 05' 00'1 oS( OE .oE 051 oOl v v v v 05L e ~.lL V V V V (;;"1 Vv .~ lnlll1VH lINV1Nllll!l dnOIl!l-O V v v l1'W·N 'lid v v V v v t v V V v. v V V l7 V v v V V V v f-.69 v v V v t.99 oOL ............... •5 roOl e 31 5° 10° 4 09 eo- 79° ~ e I:> t. ~ I:> n° '" '" '" '" " " '" I:> 75° ll. ll. I:> I:> I:> I:> ~ I:> ll. ll. ll. ll. I:> ~ I:> I:> I:> ll. I:> .. I:> I:> ll. I:> ll. I:> ll. I:> ll. I:> I:> I:> ll. ll. ll. ll. ~ 4 ll. I:> ll. 71° ~ 1:>4 I:> I:> I:> e ll. ll. ll. ll. ll. I:> I:> I:> I:> ll. I:> ll. ll. ll. ll. I:> ll. 4 I:> I:> 1:>1:> I:> I:> '" --" -- ll. 4 I:> '" '" ll. l. ll.ll. ll. I:> ll. ll. I:> I:> '" 1:>'" ll. '" ~ ll. I:> '" I:> ll. I:> I:> ll. ll. I:> .1:> I:> I:> ll. I:> ll. ll. ll. I:> 4 I:> 4 I:> '" 68° 67° Fig. 22. Distribution of O-group long rough dab. ll. ll. '" ll. ll. .. '" I:> I:> I:> ll. 69° ll. ll. ll. I:> ll. I:> I:> I:> ll. ll. ll. I:> I:> I:> I:> I:> '" " 70° FISH PR. N.HILE > 85 ll. ll. 6 6 ll. I:> I:> 71° ll. ll. 6 ll. '" ll. I:> I:> I:> l' 85 I:> 4 I:> I:> I:> 73" ~ I:> oll. 76° 0 GROUP LONG ROUGH OAB ~. I:> I:> 74° 1992 I:> I:> 78° I:> I:> I:> e )50 )00 I:> I:> I:> ]So 2lJO 15° I:> I:> I:> ll. 4 I:> ll. ll. I:> 4 I:> " 32 S° 10° 15° 15° 20° 4 80° A ~O 30° SOG 45° 40° 55° 60° ~ A 1992 - 79° ~ ~ ~ ~ 7So A 76° 75° A A 74° 73" 71° 71° 70° A A A A AA A A A A A A A A A A A A A A A A A A AA A; " A AAA A A A A A A A A A A ,, ,, ,, rr A A A A @" A A A A " 4 A ,, ,, " . " , A '" A ••.,~ A A A A A " " "I>. A " 4" A " 6So " A A " I>. 6 A A " " ,," " A A A " A A A " 4" A 69° A A A A A r r -11- ~ A I FI SH PR. N.MI LE > 85 A A A A 1 • 85 A A n° e 0 GROUP GONATUS A A A A A A " " A "" t " "" " A f< ....... ~ .~-~ ~ 67° ~ Fig. 23. Distribution of D-group Gonatus fabricii. -+ APPENDIX Research vessel Participants Professor Marty N.Bolshakova,S.Boychuk, V.Chizhikov, I.Dolgoleko, A.Gordov, T.Jusupov, L.Kuzmin, S.Ratushny, V.Ryazantsev, N.Ushakov Fridtjof Nansen S.Baturin, S.Boranov, A.Dorchenkov, V.Kapralov, O.Leontovich, S.Ostrovsky, A. Pedchenko, T.Shamray, Y.Shamray Akhill A.L.Karsakov, A.G.Nikiforov, V.K.Ozhigin, V.A.Tataurov, V.P.Vorontsov, N.S.Vovchuc Johan Hjort I.M.Beck, M.Dahl, H.Grresdal, S.Lygren, S.Mehl, E.Meland, E.Ona, L.Solbakken, I.Svellingen G.O. Sars J.Erices, H.Hammer, A.Hylen, R.Johannessen, L.Kalvenes, H.Larsen, K.Lauvas, M.M0gster, O.Nakken, A.Raknes, S.Wilhelmsen, V.Serebryakov Michael Sars O.Gullaksen, T.I.Halland, K.Hansen, G.Iversen, K.Korsbrekke, B.Kvinge, M.Mjanger, 0.Nrevdal