INTERNATIONAL COUNCIL FOR THE EXPLORAnON OF THE SEA
CONSEIL INTERNATIONAL POUR L'EXPLORATION DE LA MER
C.M. 1992/G: 10
REPORT OF THE leES STUDY GROUP ON TAGGING
EXPERIMENTS FOR JUVENILE PLAICE
IJmuiden, Netherlands, 16-22 March 1992
A TYPICAL SPi\WNING GROUND
FOR PLAICE
This document is areport of a Study Group of the International Council for the Exporations of lhe ea
and does not necessarily represent the views of the Council. Therefore it should not be quoted
witllOut consultatioß with:
General Secretary
lCES
Palcagade 2-4
DK-1261 Copenhagen K
DENMARK
PLAICE TAGGING STUDY GROUP
Contents of the report
1. Introduction
1.1. List of participants
1.2. Tenns of reference
1.3. Background
2. Material and Methods
2.1. Tagging methods
2.2. Data considered by the Study Group
2.2.1. general
2.2.2. UK Data
2.2.3. French Data
2.2.4. Dutch Data
2.3. Description ofthe tagged plaice release and recaptured allocation areas
2.3.1. Nursery release areas, juvenile plaice production
2.3.2. Recapture areas: Spawning grounds, Management areas
2.4. Distribution of fishing effort
2.5. Computer programs
2.5.1. general
2.5.2. dispersion parameters
2.6. Other fonnulas used by the Study Group
3. Results and Conclusions
3.1 Migration and dispersion
3.1.1 Recapture patterns by release area
3.1.2 Dispersion in relation to release area
3.1.3 Seasonal and sexual differences in dispersion
3.2 Relations between nursery grounds and spawning areas
3.2.1 Origin ofplaice in spawning areas
3.2.2 The contribution of a nursery 10 different spawning areas
3.3 Movement of plaice between different management areas
3.3.1 Origin ofplaice in Management areas
3.3.2 Contribution of nurseries 10 management areas
4 Management considerations
5 Recommendations
6 References
7 Appendices
Juvenile plaice tagging
1
INTERNATIONAL COUNCIL FOR THE EXPLORATION OF THE SEA
CONSEIL INTERNATIONAL POUR L'EXPLORATION DE LA MER
C.M. 1992/0: 10
REPORT OF THE ICES STUDY GROUP ON TAGGING EXPERIMENTS
FOR JUVENILE PLAICE
1.
INTRODUCTION
The ICES Study Oroup on Tagging Experiments for Juvenile Plaice met at the
Netherlands Institute for Fisheries Research (RIVO), IJmuiden, The Netherlands on
16-20 March 1992.
1.1.
List of Participants
P. van Leeuwen
M. Giret
E.Visser
J. Riley
R. Millner
A. Rijnsdorp
F. van Beek (chairman)
1.2.
Netherlands
France
Netherlands
UK (England)
UK (England)
Netherlands
Netherlands
Terms of reference
The terms of reference were to:
a) analyse the relation between nursery grounds and spawning areas for all stocks
from which data are available
b) provide information on the movement of plaice between different management
areas
c) estimate dispersion rates of plaice for the purpose of evaluating the effect of
c10sed areas
1.3.
Background
In the late 1970's the ICES Flatfish Working Group was developing an interest in
additional technical measures for the proteetion of juvenile flatfish and instigated
national programmes of tagging juvenile fish (overalllength < 25 cm) chiefly in the
period 1979-1982. The recapture data for plaice from these experiments together with
that from historie data sets plus later releases designed to fill gaps in the geographie
eover of all signifieant plaice nursery areas, were analysed at the Study Oroup
Juvenile plaiee tagging
2
\..,
principally to predict the impact geographically, of locally applied technical
protection methods. The estimation of the impact of the plaice boxes proposed in
1987 (Anon 1987) had to make some assumptions on the patterns of movement of
fish within the nursery area and from the nurseries to the fisheries. A more
sophisticated analysis of the effects of closed areas is only possible if data on the
patterns of movement and dispersion rates are available. The Working Group
studying the returns from juvenile sole tagging (Anon, 1989) proposed that similar
experiments for plaice should be identified.
•
The progress made here was seen as further steps down the trail initiated by Borley
(1916) who, being concerned about damage to plaice stocks by fishing, first looked
systematically at tagged plaice recaptures and concluded, "it is clear, that restrictive
legislation applied to the fishing off one coast, is not confined in its effect to the
waters in, or adjacent to, those in which they are enforced." The group intended to
provide quantitative detail to add to Borley's generalisation and to the results of
earlier tagging programs (Borley 1916; Carruthers 1924; Blegvad 1935; Hickling
1938; Bagge 1970; de Veen 1970, 1978; Lockwood and Lucassen 1984). Experience
gained in the analysis of the juvenile sole data (Anon 1989) was applied to the plaice
data in that the basic plaice recapture distribution from the nursery release sites were
amended to take account of the O-group production of each area and recapture rates
were corrected for fishing effects as appropriate.
2.
MATERIAL AND METHODS
2.1.
Tagging methods
As Bagge (1970) and more recent trials have confirmed that the Petersen button
(Petersen 1893) is both a practical and effective tag, it was used in all the releases.
The disc size, either 16 of 14 mm diameter and the metal of the fixing, titanium or
stainless steel, varied between releases. No valid comparisons of these alternatives
was possible from the data available.
Every attempt was made to release apparently undamaged and lively fish. An analysis
of recapture of fish released with varying degrees of catching damage by MAFF
Lowestoft staffbetween 1980-82 was made (Table 2.1.1). Skin damage was scored on
a 3 point scale and liveliness on a 2 point scale. This indicated that slight skin damage
has no effect on the chances of recapture but fish sluggish on release suffer increased
tagging mortality panicularly if they also have moderate skins damage. No allowance
have been in this report for the condition of the fish on release nor for regional
variations in the proportion of tagged plaice recaptured when for a variety of reasons,
the fish was not subsequemly reported or for which inadequate data were provided.
2.2.
Data considered by the Study Group
2.2.1. General
Data on tagged fish are kept in various laboratories in various formats. For the
purpose of this Study Group, the tagging experiments relevant to this group were
made available in a specified exchange format, in order to be able to use the same
analysis prograrns for all experiments. This exchange format of the data is in ASCII
file in which on each line release and recapture information of a tagged fish is stored
on specified positions (fixed format). The format is shown in Figure 2.2.1.1.
Juvenile plaice tagging
3
Tagged plaice data were available from releases made in the North Sea and in the
eastern and western English Channel since 1962. The length distribution of all tagged
and recaptured plaice is shown in Table 2.2.1.1.
Exclllsion by length grOllp: In order to follow the dispersion of immature fish and to
ensure that the fish tagged on the nursery grounds had not come from other areas
only fish < 25 cm were used in further analyses. The number of plaice < 25 cm
released in each area and percentages recaptured are summarised in Table 2.2.1.2.
Exclllsion by time of recnpture: Table 2.2.1.3 gives the annual recapture rate as a
percent of a11 recaptures. In many areas, there is a high percent recapture in the first
year before the individual plaice have had a chance to disperse away from the nursery
area. In order to avoid weighting the dispersion pattern by the early recaptures only
fish that were recaptured more than 12 months after the date of release were used. The
only exception was the analysis of dispersion parameters which examined data from
the first year as we11 as subsequent years.
2.2.2. UK Data
Releases were made between 1978 and 1982 in 2 main areas in the North sea centred
around the Wash-Humber and the Thames and 2 areas in the Channel centred around
Rye bay in the east of VIId and Poole in the west of VIId.
2.2.3. French Data
Releases were made between 1976 and 1980 at 4 sites from the North Sea 10 the
western Channel: the French-Delgian border, Daie de Somme, Baie de Seine and Daie
du Mont St MicheL
2.2.4. Dutch Data
Since the early sixties around 200 000 plaice were tagged by the Netherlands for
various purposes, mostly in the North Sea. The release and recapture information of
these tagging experiments are being brought into a computerized data base. This work
has not been completed yet. For the purpose of this Study Group priority has been
given to tagging experiments carried out in nursery areas. The Dutch sub-set of data
considered by this group is thus only a part of the potentially available data. The
Dutch contribution to this Study Group consists of releases made between 1962 and
1987 in estuaries and sites along the continental coast of the North Sea extending
from Hvide Sande on the Danish coast to the Belgian coast.
For a number of experiments fmal recapture percentages are underestimated, because
not a11 recaptures are included in the database or in the case of recent experiments
tagged fish are still in the sea.
2.3.
Description of the tagged plaice release and recaptured allocation areas
The demonstration of the relation between nursery and spawning areas required in the
terms of reference, necessitated that these areas be delimited. This was achieved by
portioning the coastal nursery areas on the basis of ICES rectangles or part of
rectangles where this was realistic. The open sea spawning areas was divided using
leES Divisions und rectungles.
Juvenile plaice tagging
4
.. ,r·'
2.3.1. Nursery release areas, juvenile plake production
Fifteen nursery release areas covering most of the coastline of IVb and c, and VUd
and e were selected on the basis of geographical features and the presence of
significant numbers of juvenile plaice identified in annual pre-recruit surveys in these
areas. Most of the information on the distribution of juvenile plaice in its nurseries
originates from the Demersal Young Fish Surveys (DYFS), carried out by Delgium,
The Netherlands and Germany (Anon, 1985; Van Deek et al, 1989) along the
continental North Sea coast and in its estuaries and English pre-recruit surveys along
the UK coast in the North Sea and the English Channel (Anon, 1985; Millner et al,
1988). In the English Channel, information from the International Deam Trawl
Surveys (lDTS) carried out by England, Franee and Delgium (Anon, 1990; Millner &
Whiting, 1990) and French pre-reeruit surveys was used. The release areas, together
with their names and identifieati?n numbers are shown in Figure 2.3.1.1.
•
The release areas show large differenees in the abundanee of juvenile plaice that ean
be assumed to reflect differences in reeruit produetion. The level of recruit production
of the various nursery areas (nursery power) was estimated from the available prereeruit survey data.
For the continental side of the North Sea the nursery power was estimated from the
mean l-group plaice density in the auturnn surveys carried out between 1970-1990,
and the surface area of the nurseries (Table 2.3.1.1). The l-group density is
considered to give a better representation than the O-group density because a part of
the O-groups will be still distributed on the tidal flats outside the survey area.
For the UK side of the North Sea and eastern English Channel, the nursery power was
estimated from O-group density in auturnn surveys carried out between 1981-1990
(Table 2.3.1.2).
In order to link the French side of the ChanneI with the UK side, 2-group plaice
abundance estimates were used from three beam trawl surveys earried out in August
1989 - 1991 (Table 2.3.1.3).
The three data sets were converted to the same seale in order to calculate a relative
contribution of each nursery to the total recruit production of the total North Sea
(Division IV) and English Channel (Division VIId and e). Table 2.3.1.4. illustrates
this procedure. In the first step, the estimates of the continental and UK side of the
North Sea (Table 2.3.1.1. and 2.3.1.2.) were combined, assuming that the UK side
contributed 10% and the continental side 90% of the North Sea recruitment (Anon
1985). The observations in column A were converted to a relative contribution
summing to 100% for the total North Sea (eolurnn D). For the Hvide Sande nursery a
recruit production of 50% of the Sylt nursery was assumed, based on a limited
number of survey data. The contribution of the UK northeast coast (north of
Flamborough Head) was equally estimated on a limited number of survey data. In the
seeond step, the observed plaice production values in the nurseries on the UK coast of
the Channel (Rye and Pool Day, column A), obtained simultaneously with the North
Sea nurseries Wash and Thames, were sealed to column D. In the third step, the
Channel nurseries on the UK and French coasts were scaled using observed plaice
densities of colurnn C (DTS surveys 1989-1991; Table 2.3.1.3). The final contribution
is given in colurnn D ofTable 2.3.1.4., assuming a production value for Daie Mont St
Michel.
The nursery power values indicate that about 87% of the production of plaice recruits
oeeurs on the contincntal eoasts of the North Sea, 10% on the UK east eoast and just
under 3% only on the Channel nursery grounds (Table 2.3.1.4).
Juvenile plaiee tugging
5
2.3.2. Recapture arcas: Spawning grounds, Management areas
Recaptures in the North Sea were allocated to sea areas, designed as much as
possible, to subdivide geographically, plaice spawning grounds. The information used
for this subdivision were the centres of spawning described by plankton surveys by
many nations (Harding et al, 1978; Heessen & Rijnsdorp, 1989; Simpson, 1959).
Other criteria used in this subdivision were the RIVO plaice biological sampling
areas. The recapture areas are shown in Figure 2.3.2.1
In the analysis four management recapture areas were considered: the North Sea (IV),
the Eastern English Channel (VII d), the Western English Channel (VII e) and other
areas combined where recaptures were reported from (IIIa, VII f, g and VII j).
2.4.
Distribution of fishing efTort
The pattern of recaptures will both depend on the distribution of the tagged fish and
the distribution of fishing effort. In 1987 the North Sea Flatfish Working Group
collated effort information by statistical rectangle and by quarter in order to estimate
discard mortality rates in plaice (Anon, 1987). Quarterly fishing hours were available
from the Dutch beam trawl fleet in 1974-1977 and from the UK otter trawl fleet in
1974-1977. It was assumed that the effort distribution ofthe Danish and Gerrilan fleet
was similar to that of the Netherlands, and that of the Belgium fleet to the UK. The
UK effort (EUK) was representative for 30% of the plaice landings and the
Netherlands effort (EN) for the remaining 70%. A combined effort index (ETOT) was
then calculated by according to Ecombincd 70% EN + 30% EUK.
•
=
The pattern 'of effort distribution calculated as such is not strictly representative for
the nonhern Nonh Sea, since this area is the main fishing area of the Scottish
demersal fleets. Scottish plaice landings is about 10% of the total international
landings in the 1980s. The average quarterly fishing effort over the period 1987-1991
of the three main Scottish demersal fleets (seine, otter trawl, pair trawl) were made
available during the working group by counesy of Dr R. Cook (Aberdeen) and were
combined with the already available effort (Ecombined) using the same procedure. It
was thus assumed that the three demersal fleets in Scotland had equal catchability for
plaice. The result is given in Appendix IV. The relative fishing effort in the six subareas distinguished in the North Sea is given in Table 2.4.1 and clearly shows that the
demersal fishing effort is mainly concentrated in the southern and southeastern North
Sea throughout the year. During summer the effort shows a slight shift to the nOrth
and a small increase in relative effort in the central Nonh Sea around in the areas of
Fisher Bank and Flamborough.
2.5.
Computer Programs
2.5.1. General
Two computer programmes were developed for the analysis of the data. Programme
TAGANA2 reads the exchange format files. It caIculates for each recapture the
individual migration parameters, time in sea, distance travelled and direction of
dislocation relative to the Nonh Pole The formulae used by this programme are
described in the section below. In addition it caIculates from release- and recapture
positions the corresponding leES rectangles. This information is written in an
outputfile together with length, sex and maturation parameters. This output file is
Juvenile plaice tagging
6
used as input for programme TAGANA3. TAGANA2 also produces administrative
output on the contents of the file such as, number of releases and recaptures, periods
in which the experiments were carried out, missing information and a table with the
release length distribution of the releases and recaptures.
Programme TAGANA3 produces tables with the migration parameters by
experiments, such as the mean direction of displacement, velocity, mean time in sea,
average distance travelled by the individual fish, distance of displacement of the
population and coefficients of dispersion (formulae used are also given in the section
below). The tables for the experiments considered are given in Appendix I. The units
used in the calculations were months and nautical mHes. The parameters are presented
for the release quarter and the subsequent 15 quarters after release..
TAGANA3 allows for a number of options with regard 10 the selection of data to be
analysed. There options are :
a) selection by sex
b) selection by release quarter
c) selection of release length range
d) selection of release year range and
. e) selection of recaptures caught after a certain period in sea
Except for option d) all other options have been used by the Study Group.
Other output from T AGANA3 is a table, presenting number of recaptures by
rectangle and quarters for 4 years after the release quarter. These tables are presented
in Appendix II. This information is also produced in maps, which were used for
visual inspection.
All computer programmes were written in PASCAL and run on a VAX at RIVO
Umuiden.
2.5.2. Dispersion parameters
This section gives an overview of the migration parameters used in the calculations of
the Study Group.
Parameters calculated for a single fish (reference)
•
The distance of displacement of a single fish: r, which is the distance between release
and recapture position in nautical mHes is ealculated as
r
.
= 2 * M * arcsm
{2-2*cOS(a)*cOS(b)*COS(C-d)--*sin(a)*Sin(b)
2
2.5.1
where M =mHes per radian =3437.746771
a: release latitude in radians
b: recapture latitude in radians
e: release longitude in radians
d: recapture longitude in radians
The direction of displacement of a single fish (0) in degrees compass points is
calculated using the following procedures and formulae. In eases where release and
recapture latitude are the same, the direction of displacement is East or West (900,
270°). Similar release and recapture longitudes indicate displacement in North or
Juvenile plaice tagging
7
south direction (0°, 180°). In all other cases where release and recapture positions
were not the same (most cases) e was calculated as:
1t
. b-a,.
er =2- arcsm(x)
2.5.2a
or if d<c
1t
b-a,.
er = 21t - 2- arcsin(x)
2.5.2b
ar expressed in degrees
8d
where x = 2
=
360 * er
21t
* · {2-2*cOS(a)*cOS(b)*COS(C-d)--*sin(a)*Sin(b)
arcsm
2
Parameters calculated far a group of fish (experiment)
The migration parameters for the various experiments were calculated according to
formulae given by Jones (1965) and Anon (1971).
The distance of displacement of a single fish: r, can be broken down in a North/South
component: r cose and EastlWest component: r sine, The mean direction of
displacement 'I' of a group of fish then becomes:
tan 'I' =
Irsine
~rcose
2.5.3
The velocity,V, with which the centre of the population is moving in direction '1',
expressed in miles per month is given by:
(Lr sin8)2 + (~r cos8)2
V =_.l-
_
Lt
•
2.5.4
where t is the time in sea in months after release for each fish
the mean number of days' absence T is given by
T
= Lt
n
2.5.5
The distance of displacement of the population (R) than becomes:
R=VT
2.5.6
The average distance travelled by a group of fish (D) is the mean of the distances
travelled by all individual fish (n) given by:
Juvenile plaice tagging
8
D
= Lr
2.5.7
Ln
The monthly coefficient of dispersion of fish about their centre of density (a 2 ) is
given by:
L
(Lr cos8)2 + (Lr sin8)2
r
: ----------
•
2.5.8
whereas the coefficient of dispersion of fish about their centre of density after a mean
time, T, becomes:
2.5.9
2.6.
Other formuIas used by the Study Group
The recaptures in a certain area can originate from different nurseries. The chance to
recapture a fish, tagged in a particular nursery, depends on the number of fish tagged
in that nursery. In order to take account of different levels of tagging in the various
nurseries the recaptures can be expressed as a fraction of the number released:
2.6.1
where nj is the number of recaptures from nursery j and Nj the number of releases in
than nursery.
In a particular recapture area, i, this fraction becomes:
•
2.6.2
The contribution of the recruitment from a nursery area to a recapture area also
depends on the importance of the nursery to the total population. In order to estimate
the relative contribution of several nurseries to the recruitment or population in a
selected area these fractions should be weighted by the "power" of the nursery, Pj.
The power of the nursery area gives the proportion of recruits from a nursery area
relative to the total production of all nursery areas considered, based on abundance
estimates of juvenile plaice from pre-recruit surveys and the surface area of the
nursery ground (see section 2.2.3.). The recruitment from nursery j to recapture area i
can than be compared by:
n··
rij
=W
* Pj
J
Ri
=L
2.6.3
15
and
rij
2.6.4
j=l
Juvenile plaice tagging
9
where Ri is the total in the selected recapture area i when there are 15 nursery areas.
As fishennen or anglers, who return the recaptures will not discriminate between the
origin of the recaptures in aselected area, in other words, the effort or fishing
mortality is the same for all recaptures, the ratios Tij * Rr l can be considered
representative for the composition of the origin of the population in the recapture
area. In fact the ratios calculated in this way would be similar to the ratios of the
straight recaptures from the different nurseries, if the level tagging intensity in
nurseries would have been proportional to the importance of the nursery to the total
recruitment.
A problem occurs when not all nurseries are covered by tagging experiments. Fish
originating from these areas will not be recognised and considered. This will lead to
overestimates of the importance of the other nurseries.
It should be pointed out that in the caIculations carried out by the Study Group the
actual release numbers, Nj, were replaced by an estimate of "remaining releases in sea
(Table 2.2.1.2). The Study Group considered this appropriate
after one year":
because it analysed only recaptures over 12 months in the sea while a great part ofthe
recaptures is made in the first year elose to the release sites and because the
percentage of recaptures in the 1st year differs considerable between release areas.
The estimated remaining releases after 1 year were calculated using the fonnulae:
Nt
•
n~+'
N~+ =N.*-JJ
J
nj
where nt is the number of recaptures made after the first year.
A further amendment was made to the release numbers in the Hvide Sande, Sylt and
Norderney nurseries. The recapture data from the experiments carried out in these
areas were not all included in the data base. Also a number of tagging experiments in
these areas were carried out recently and a significant number of recaptures of these
experiments can be expected in the future. The low recapture percentages in these
areas are artificial resulting in an underestimate of the contribution of these areas
compared to other areas. In order to take account of this deficiency, the release
numbers were set at a lower level producing a recapture percentage of 25% equivalent
to the converged experiments in the adjacent areas (Waddensea E, Waddensea W and
Dutch coast).
•
The fonnulae discussed above estimate the origin of the fish in a selected recapture
ground, which may represent a spawning area or management area.
Another question is the dispersion of the fish tagged in a particular nursery area to
various recapture grounds. The number of recaptures in different recapture areas
depends now also on the amount of effort, which usually varies considerably between
recapture areas. In order to take account for different levels of effort in different
recapture areas fonnula 2.6.3 should be extended 10:
n"
r"IJ -- Nj
~
* p.J * E·1
2.6.5
where Ei is the effort in recapture area i. The ratios of these corrected fractions should
now be compared within a nursery area, rij * Rr 1 , where the sum of these ratios,
when there are 3 recapture areas is given by:
Juvenile plaice tagging
10
.
,
3
Rj
=L
rij
2.6.6
i=l
The absence of effon in a potential recapture area makes it impossible to estimate the
contribution of a nursery to that area. This will not affect the ratios between the other
areas but will bias the absolute estimates of contribution of a nursery to the other
areas.
3.
RESULTS AND CONCLUSIONS
3.1.
Migration and dispersion
The nonnal pattern of recaptures indicates that the plaice tagged as 1 to 2 year olds
passed through two phases. Firstly a dispersion phase lasting between one and two
years at the end of which the area of distribution of recaptures was fully described. In
the second phase additional recaptures accumulated in this area and within which
relatively minor seasonal movements could be identified.
The number of recaptures in each experiment usually fell progressively with time (see
Table 2.2.1.3 and Appendix I) but this was not always the case. Norderney and the
Wash area were exceptions. Most recaptures, however, took place during the
dispersion phase by the end of which most fish would have achieved sexual maturity.
The summary maps of recaptures for each experiment by quarter (see Appendix III)
exc1ude plaice caught within one year of release but must still inc1ude a proportion of
immature and/or fish in the dispersion phase and this could partially mask any
seasonal movements.
3.1.1. Recapture patterns by release area
•
The quarterly recapture summary maps are reviewed by release nursery area with
particular reference to the Figure 3.1.1.1. showing the centres of distribution of
recaptures after Jones (Jones 1965). In the figure the points are numbered by quarter
years (Le. 1 represents recaptures in Jan-March inclusive) and where feasible the four
quarter points have been joined up in sequence and would therefore be a summary
measure of seasonal movements. The changes in position over the year appear 10 be
minor and mostly add up to about 100 - 200 mHes. They cover therefore, a small
proportion of the total sea area described in the dispersion phase. The recapture
pattern of plaice (free in excess of one year) are reviewed below to identify points
additional to the general patterns described above.
Hvide Sande The centres of recapture are situated between 2 and 3° to the west of the
release position. Recaptures in the spawning quarter (Q1) tend 10 the south and west.
Sylt Recaptures in Q1 indicate the fish from this area participate in three spawning
areas all 10 the east of 2°E. Recaptures in Q3 and Q4 tended 10 be more offshore than
in Q1 and Q2.
Norderney Recaptures in Q1 are in three spawning areas with the centre of
distribution just to the east of 2°E in the transitional area. Most recaptures in Q3 and
Q4 were in the Gennan ßight.
Juvenile plaice tagging
11
Eastern Waddensea Recaptures in Ql indicate that most recruits from this areas
spawn in the transitional area or in the Southern ßight. Subsequent recaptures
predominate N of 54°N and E of 2°E.
Western Waddensea Significant recaptures in the Channel in Ql with fish possibly
participating in six spawning centres from Start Point in VIIe to the German ßight.
Recaptures in Q3 and Q4 indicated the return ofmost plaice 10 the North Sea (IVc).
Dutch Coast Generally similar to the Western Waddensea. 71 % recaptures in the 1st
year reduces data available here.
Zeeland Estuary Recaptures in Ql indicate significant participation of recruits in
spawning in VIId and e, the Southern ßight and the southern edge of the transitional
area with some recaptures in the Channel persisting throughout the year indicating
permanent residence there.
French ßelgian Coast nVc) Limited numbers of recaptures show patterns similar to
those from the Zeeland Estuary except that only one recapture was made in the
Channel in Q4.
•
Wash Area Recaptures in Ql predominately from the Flamborough spawning area
centred 2°N of the release rectangles but stretching coastwise north to the Firth of
Forth. Recapture patterns in Q2-3 similar to Ql but with some extreme additional
movements north the Moray Firth and slight offshore movements eastwards. Unique
recapture pattern with time, only 8% recaught in the 1st year.
Thames Area Recaptures in Ql predominate in the centres of spawning (Lee and
Ramster 1981) to the east of the Southern ßight and the southern part of the
Flamborough spawning area. Penetration into the Channel is minimal in Q2 and Q3
and there are many recaptures from further north up to the Moray Firth, east to the
Dutch coast and south and west chiefly from the eastern Channel.
Rye Area 65% recaptures in the 1st year severely reduced the number of subsequent
recaptures. Those in Ql indicate most spawning in VIId. Recaptures in Q2-4 indicate
seasonal movement into the North Sen. Movement to the south and west minimal.
roole Area Recaptures from rectangle of release predominate in all quarters. Some
recaptures in Q4 and Ql in VIIe suggest spawning in that Division.
•
Haie de Somme Recaptures in Ql chiefly from the French side of the Channel
spawning areas. One from southern Ireland, none from the North Sea. Recaptures in
Q2-4 indicate exclusively a westerly distribution and a permanent occupation of VIId
and e only.
ßaie de Seine A high first year recapture rate (67%) reduced subsequent recapture but
those in Ql indicate exclusively a spawning locally on the southwestern edge of the
VIId spawning concentration. Recaptures in other quarters suggest that any
permanent movement out of the eastern Channel is to VIIe or further west.
Haie du Mt St Michel With 83% of recaptures made within the 1st year the
identification of seasonal patterns is frustrated. In total one recapture is recorded from
outside Division VIIe, in de ßaie de Somme, indicating that this small nursery area
probably produces recruits only in the Channel
Juvenile plaice tagging
12
3.1.2. Dispersion in relation to release area
This section will discuss the dispersion of the juvenile plaice. Three parameters
describing the dispersion were distinguished: 1) the average distance between release
and recapture position (R in miles); 2) the dispersion rate (a2 in miles2.month- 1) and
3) surface area encompassed by the recaptures (a2 *t in miles 2 ). The parameter
estimates for the different release areas are given in Appendix 11. A first inspection of
the dispersion parameters of the different experiments did show a large variability
within each experiment and it was therefore decided to calculate average parameter
values for different release areas. For the three parameters distinguished, a weighted
average was calculated for each area by weighting over the number of observations.
Based on the depth distribution and the nature of the sea bed five areas were
distinguished:
1) North Sea continental coast between 51°N and 58°N (release areas 1-8);
2) North Sea UK coast (release areas 9 and 10);
3) French coast ofthe Channel (areas 13-15);
4) Rye bay (area 11) and
5) Pool Bay (area 12).
Relationships between the parameters and the time after release is shown in Figures
3.1.2.1 a-c. The x-axis is expressed in relative quarters after release and thus do not
match with the absolute quarters of the year.
Figure 3.1.2. La shows that the average distance R from the release position increases
continuously with time since release, although in some areas there is a tendency to
level off after about 2 years. The average distance differs substantially between areas,
being larger in the North Sea compared to the Channel, with Pool ßay on the UK side
showing a panicularly low distance between release and recapture position.
•
No clear trend in the rate of dispersion was apparent with time in sea after release
(Figure 3.1.2.1.b). However, a more detailed inspection of the pattern within the
different release areas show some similarity. In the continentaI North Sea release the
dispersion rate slightly increases up to 3 years after release and then gradually
decreases. This pattern of an increase and then a decrease is also observed in the UK
coast of the North Sea and the French coast of VII d. An increase in the rate of
dispersion is shown in the Rye Bay release. The overall level of dispersion rate differs
substantially between the areas. Now the UK side of the North Sea and Channel show
higher dispersion rates than the continental coasts, with Pool Bay as an exception. In
this area the dispersion rate is very low. de Veen (1970) gave an estimate of
dispersion rates in juvenile plaice of about 20 miles2 .day-l, which roughly compared
with the dispersion rates observed in plaice released on the UK coast of the North
Sea.
The size of the distribution area of the recaptures shows an increase with the time in
the sea (Figure 3.1.2.1.c). The rate of increase is largest in the UK North Sea and Rye
ßay release. The continental North Sea and VII d are intermediate, while Pool Bay
again shows an exceptional result and a very small distribution area. The increase in
the distribution area seems to increase up to 2-3 years after release and seerns then 10
level off. The level reached in the UK North Sea and Rye Bay releases appears to be
larger than that of the continental release.
Juvenile plaice tagging
13
3.1.3. Seasonal and sexual differences in dispersion
In the above section dispersion parameters were compared without taking account of
possible seasonal and sexual differences. The extensive number of recaptures of the
tagging experiments carried out along the continental coast of the North Sea did allow
the investigation of seasonal and sexual patterns in dispersion. For an exploratory
exercise the recapture data of the western Wadden Sea release was chosen. Tbe sire
range of released fish was further restricted to fish <20 cm, because we were
interested in the dispersion of juvenile plaice from the nursery grounds and among the
males >=20 cm a substantial proportion may already be mature.
We will first focus on the migration parameters obtained from releases in different
quarters of tbe year (Figure 3.1.3.1). In contrast to the figures in tbe previous section,
tbe x-axis now denotes the quarter of the year in tbe year of release (1-4) and tbe
following year (5-8). Tbe average distance between the recapture position and tbe
release position increases with tbe time after release, but the pattern of increase differs
between the releases in the 1st and 2nd quarter compared to tbose in the 3rd and 4th
quarter. Tbe latter fish are recaptured elose (20 mHes) 10 the release area up to the
autumn of the following year, whereas those released in the 1st and 2nd quarter are
already recaptured at an average distance of 80 miles from the release area. This
pattern is apparent in botb male and female plaice.
Tbe dispersion rates estimated from the quarterly recapture patterns indicate a wide
variability between tbe different quarterly releases. Tbere appears to be a slight
tendency for the dispersion rate to peak in quarter 5 corresponding to the spawning
period. Tbe trend over time of the total distribution area of the recaptures shows a
similar pattern as the distance between recapture and release position.
Comparison of the releases in the different quarters is hampered by the unbalanced
number of recaptures in the different experiments. In order to explore seasonal
patterns in migration parameters further, the recaptures of the different experiments
were pooled by the quarter of the year. The results are plotted in Figure 3.1.3.2 and
show that the seasonal pattern already vaguely apparent in the disaggregated data, is
revealed more elearly in the pooled data. It also allows us to compare the differences
between males and females. The average distance between recapture and release
position tends to peak in the 1st quarter in tbe first and second year after tagging and
does not indicate a difference between the sexes. The average dispersion rate of male
and female shows a surprisingly consistent pattern with a peak in the 1st quarter and a
generally lower level 2nd to 4th quarter. No dear difference between male and female
emerges, except that the male shows a bigher dispersion rate in the year after release.
The increase in tbe size of the distribution area does not appear to be different
between male and females. Tbe increase in the size of the distribution area is
relatively small in tbe year of release and becomes only substantial just prior to the
spawning season in the first year after release.
3.2.
•
Relations between nursery grounds arid spawning areas
3.2.1. Origin of plaice in spawning areas
Table 3.2.1.1. gives the number of recaptures nij in tbe different spawning areas i
from nursery area j in the first quarter. The contribution of the different nursery
grounds to the recaptures on a spawning area in the first quarter was calculated
according to fOTmulae 2.6.3 and 2.6.4.
Juvenile plaice tagging
14
Table 3.2.1.2. presents the results (in percentages) and show that for instance the
spawning area in the German ßight receives in the first quarter its recruits mainly
from the Sylt nursery area (80%), with a minor contribution of Hvide Sande (10%),
Nordemey (5%) and the Waddensea 5%. The Southem Bight spawning ground
receives 58% of its recruits from the Wadden Sea. However, also the nursery grounds
in the eastem North Sea contribute 24%. Recruitment to the Southem Bight spawning
from the UK coast is negligible (4%). The spawning grounds in the western North
Sea (Flamborough) are supplied for 78% of recruits by the nursery grounds along the
UK coast (Wash, Thames estuary and Rye Bay), while only 22% originales from the
continental coast, in particular from the nursery areas in the German Bight.
(Norderney).
The spawning areas in the English Channel receives a substantial proportion of its
recruits from the nursery grounds in the North Sea and only a minor proportion from
nursery areas in the Channel itself. Division VII d receives 57% and division VII e
receives 56% from the North Sea, in particular from the western Waddensea and the
Scheldt estuary.
•
The North Sea nurseries also indirectly feed the UK North Sea coast. Since eggs and
larvae spawned from these North Sea recruits in the eastern Channel, subsequently
drift onto the English east coast nurseries and juveniles also move from the channel to
the English east coast. Consequently the continental North Sea nurseries can be seen
to have an important influence directly on the Channel but also indirectly on the UK
North Sea coastal areas.
For completeness the information for the other quarters is also given in Table 3.2.1.2.
3.2.2. Thc contribution of a nurscry area to different spawning areas
In the above section the recaptures made in a spawning area were analysed far the
contribution by the different nursery areas. To answer the question how the
recruitment from a particular nursery ground is partitioned over the different
spawning areas, we have to take account of the differences in fishing effort.
Information on the distribution of fishing effort was available for the North Sea but
not for the English Channel.
•
In order to estimate the contribution of a nursery area to the different spawning areas
the observed numbers of recaptures were weighted over the fishing effort by
spawning area according to formulae 2.6.5 and 2.6.6, and standardized to a column
total of 100. Table 3.2.2.1 presents this information and shows for instance that 11 %
of the plaice tagged in the eastern Wadden Sea recruit to the German Bight, 23% to
the Transition Area und 57% to the Southern Bight. The plaice tagged in the western
Wadden Sea provide only 2% of the recruits to the German ßight, 15% to the
Transition Area and 80% to the Southern Bight.
Data for the other quarters are also given in Table 3.2.2.1.
3.3. l\Io\'ement of pIaice between different management areas
Three different plaice management areas were considered in this report: the North Sea
(IV), the Eastem English Channel (VIId) and the Western English Channel (VIIe).
The juvenile tagging results indicate that to some extent exchange occurs to adjacent
areas as Skagerak!Kattegat (IIIa), VIIh,j,k und the ßristol ChannellCeltic Sea (VIIf,g).
However since no information on juvenile plaice tagging experiments from these
Juvenile plaice tagging
15
areas were available to the Study Group, the relationships between these areas and
those considered by the Study Group could not be quantified. Since significant
nursery areas are known in lIla and there are no natural physical barriers between the
Skagerak area and the North Sea, the lack of information from lIla is the most
prominent.
3.3.1 Origin of p)aice in Management areas
The origin of the recaptures by management area was analysed for each quarter of the
year separately based on the results presented in Table 3.2.2.1 and discussed in
section 2.2.1. Table 3.3.1.1 shows the summed contribution ofthe nursery areas in the
different management areas to the recaptures made in these management areas. This
table gives the results for the total year and the separate quarters. The data for the
total year indicate that the recruitment from the nursery areas in the North Sea provide
almost all recruitment to this area, but also a significant part to the management areas
in the English Channel. Inspection of the contributions in the separate quarters does
not show a clear difference between the quarters.
•
3.3.2. Contribution of nurseries to management areas
In order to estimate the contribution of a nursery area to the different management
areas information is needed on the relative level of fishing effort in these management
areas. No information is available on the demersal fishing hours in the Divisions VIId
and e, but the population assessments carried out for the plaice stock in these areas
indicate that the level of fishing mortality is not much different and similar to the
North Sea. The proportions of recaptures in the North Sea and Divisions VIId and e
from juvenile tagging experiments carried out in these areas are therefore direcdy
comparable. These proportions are corrected for the different levels of tagging in the
various nursery areas and weighted by the power of the nursery areas (see section
2.6). The results are shown in Table 3.3.2.1.
This table shows for instance that 1.4 and 0.4 % of the juvenile plaice tagged in North
Sea nurseries is recaptured in Divisions VIId and e respectively. Under the
assumptions made, these percentages can be considered as representative for the part
of the North Sea plaice production which leaves the North Sea and contributes to the
stocks in the English Channel. From section 3.3.1 it appears that in the English
Channel this North Sea contribution represents between 38 and 53% to the stocks
present in Divisions VIId and e respectively (Table 3.3.1.1).
•
Assessments, carried out by the ICES North Sea Flatfish \Vorking Group (Anon.
1992) indicate an average recruitment level ofplaice in the North Sea of 500 million
l-year-old recruits compared to 30 and 5 million in VII d and VII e respectively. The
indication by the tagging experiments that 1.4% and 0.4% of North Sea plaice
migrates to VII d and e which constitute 38% and 53% of the local stocks correspond
with estimates of recruitment in VII d and e of 18,4 and 3.5 million respectively,
which is less than a factor 2 different from the Flatfish Working Group estimates.
This result must be considered as remarkably good given the uncertainties in the
assessments by the Flatfish Working Group, especially in the Channel. Further it must
be considered that the recent time periods over which the FWG calculate average
recruitment only partially overlap the period over which the tagging experiments were
carried out. Also thc assumptions made by this Study group, concerning factors
influencing the estimates, such as fishing mortality in the three areas and nursery
power, are rather cmde, incomplete and preliminary.
Juvenile plaice tagging
16
4. Management considerations
The results of the tagging experiments have some important implications for the
management of the plaice in the North Sea and the Channel.
1 - the recruitment from the nursery grounds of the plaice box (continental coast of
the North Sea) will enhance the plaice stock in the southern and eastern North Sea ,
but not directly in the western North Sea.
2.- the plaice population in VIId and VIIe receive a substantial proportion of their
recruitment from the North Sea nursery areas and the plaice box, therefore, is
expected to enhance the level of recruitment in the Channel.
5. Recommendations
•
The covering of nursery areas by tagging experiments in on juvenile plaice shows
gaps in Danish coastal areas (Jammer Bight, Skagerak and Kattegat) and eastern
Scottish coastal waters. Tagging experiments together with the analysis of historic
tagging data in these areas should be carried out in order to provide information on
the importance of these areas to the recruitment in the North Sea.
The preliminary analysis on plaice tagging data in nursery areas is based on a subset
of historical data, which is available in a computerized data base. More historical
experiments can be made available in future.
Estimates of the "nursery power" of nursery areas, presently not available from
certain areas as Jammer Bight, Skagerak and Kattegat should be made available from
survey data.
Adequate effort distribution data of the commercial fleets are required, especially in
the English Channel, in order to be able to calculate the contribution of recruits from
a particular nursery area to several spavming populations and management areas.
•
Effort distribution data are also required to estimate the bias in the dispersion
parameters caused by the pattern of effort distribution.
6. REFERENCES
Anon. 1971. Report by the Roundfish Working Group on North Sea Cod. ICES
C.M.1971/F:5.
Anon. 1985. Report ofthe O-group Flatfish Working Group. ICES C.M.1985/G:2
Anon. 1987. Report of the Ad Hoc Meeting of the North Sea Flatfish Working
Group, ICES CM 1987/Assess:14.
Anon. 1989. Report of the Ad Hoc Study Group on Juvenile Sole Tagging. ICES
C.M.1989/G:21
Anon. 1990. Report ofthe Study Group on Beam Trawl Surveys in the North Sea and
Eastern Channel C.M.1990/G:59
Anon. 1992. Report of the North Sea Flatfish Working Group, ICES CM
1992/Assess:6.
Juvenile plaice tagging
17
Bagge, O. 1970. Tbe reaction of plaice to transplantation and taggings. Meddelelser
fra Danmarks Fiskeri- og Havundersogelser Ny Serie: 6(5). 1-331 p.
Beek, F.A. van, Rijnsdorp, A.D. and Clerck, R. de 1989. Monitoring juvenile stocks
of flatfish in the Wadden Sea and the coastal areas of the southeastem North
Sea. Helgoländer Meeresunters. 43: 461-477.
Blegvad, H. 1935. Transplantations of plaice from the North Sea to the Belt Sea
1928-1933. The scientific survey Report of the Danish Biological Station 39:
9-84.
Borley, J.O. 1916. An analysis and review ofthe English plaice marking experiments
in the North Sea. Fishery Invest. Ser 2,3(3), 126 pp.
Carruthers, J.N. 1924. Report on the English post-war plaice marking and
transplantation experiments. Fishery Invest. Ser 2, 6(5), 22 pp.
Harding, D., NichoIs, J.H. and Tungate, D.S. 1978. The spawning of the plaice
(Pleuronectes platessa L.) in the southem North Sea and English Channe!.
Rapp. P. -v. Reun. Cons. int. Explor. Mer, 172: 102-113.
Heessen, HJ.L. and Rijnsdorp, A.D. 1989. Investigations on egg production and
mortality of cod (Gadus morhua L.) and plaice (Pleuronectes platessa L.) in the
southem and eastem North Sea in 1987 and 1988. Rapp. P.-v. Reun. Cons. int.
Explor. Mer 191: 15-20.
Hickling, C.F. 1938. The English plaice marking experiments 1929-1932. Fishery
Invest., Lond., Sero 2, VoI16(1).
Jones, R. 1965. FAO/UN, FAO/GFCM seminar on sardine tagging. Split,
Yugoslavia, 2-14 November 1964, Report and Iectures. FAO Fish. EPTA Rep.,
(1995)
Lee, AJ. and J.W. Ramster 1981. Atlas of the sea around the British Isles. MAFF
Lowestoft
Lockwood,S.J. and Lucassen, W. 1984. The recruitment of juvenile plaice
(Pleuronectes platessa L.) to their parent spawning stock. J. Cons. int. Explor.
Mer, 41: 268-275.
Millner, R.S., J.D. Riley and C.L. Whiting 1988. Variability in abundance of 0- and
l-group plaice and sole on the east coast ofEngland. ICES CM 1988/G:14
Millner, R.S. and C.L. Whiting 1990. Distribution and abundance of juvenile sole
and plaice in the eastem English Channel from young fish surveys. ICES CM
1990/G:38
Petersen, J.C.G. 1883. On the Iabeling of living plaice. Rep. Danish bio!. Stat. 4.
Riley, J.D., Symonds, DJ. and Woolner, L. 1981. On the factors influencing the
distribution of O-group demersal fish in coastal waters. Rapp. P.-v. Reun. Cons.
int. Explor. Mer 178: 223-228.
Simpson, A.C. 1959a. The spawning of the plaice (Pleuronectes platessa ) in the
North Sea. Fish. Invest. Sero 2, 22(7), 111 pp.
Veen, J.F.de 1970. On the orientation of the plaice I. Evidence for orientating factors
derived from the ICES transplantation experiments in the years 1904-1909. J.
Cons. int. Explor. Mer, 33: 192-227.
Veen, J.F.de 1978. On the selective tidal transport in the migration of North Sea
plaice (Pleuronectes platessa) and other flatfish species. Neth. J. Sea Res. 12:
115-147.
Juvenile plaice tagging
•
•
18
Table 2.1.1.
Fish conditions on release in relation to recapture based on fished
released at <25 cm in the UK experiments in the period 1980-1982.
Plaice numbers
Symtom
Score
Skin damage:
nil
1
slight
2
•
Numbers
Numbers
released
recaptured
% Recaptured
1314
1748
502
664
38.2
38.0
3
307
84
27.4
Skin damage:
lively
1
sluggish
2
3110
259
1175
75
37.8
29.0
moderate
Skin damage x Liveliness
1
1
1285
494
38.4
2
1
1559
38.5
3
1
1
2
266
29
600
81
2
2
2
189
37
3
8
64
3
30.5
37.6
33.8
8.1
•
19
Table 2.:1.1.1 Length distribution at tagging and recapture
A: L.nglh dislribution al lagging
Release area:
I
length cm
to
11
I
12
10
13
18
14
26
15
85
16
156
17
247
18
369
518
19
20
675
21
815
22
722
23
515
24
265
25
243
26
205
27
219
28
139
142
29
30
81
31
31
32
11
33
7
34
6
35
36
37
38
39
40
41
42
43
44
45
46
47
48
49
50+
unknown
5512
<25 cm re1.4422
Total
2
3
20
94
264
373
385
477
586
655
448
309
179
137
lOS
121
91
70
60
51
23
21
14
9
14
5
5
7
2
4
I
3
1
I
1
4544
3794
3
5
29
105
148
169
140
59
27
44
26
31
15
10
I
1
4
4
42
140
610
1272
1541
1347
1017
744
308
151
45
6
5
2
2
5
6
5
37
206
538
852
1449
2089
2659
2525
2082
1816
1450
1106
871
650
369
132
80
52
23
15
2
5
7
37
148
395
528
665
820
871
875
852
838
814
736
673
550
472
384
I
7237
7233
I
12
59
313
589
779
760
723
733
672
578
507
428
363
292
224
158
97
64
38
22
13
6
3
2
7
20
29
54
to8
158
200
187
187
170
128
88
62
48
26
23
16
14
11
3
2
3
2
2
I
1
9
10
11
12
0
0
22
25
40
81
126
143
126
78
48
19
13
2
4
17
103
182
199
219
242
282
361
416
394
424
404
441
1
I
18
28
31
53
6S
69
84
60
64
t
400
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
263
177
152
100
83
55
46
33
17
16
14
7
5
3
4
I
I
I
0
0
0
I
0
0
0
0
123
138
157
87
87
26
28
17
8
14
5
5
I
2
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
2
11
50
76
111
78
71
68
43
16
16
to
18
9
20
22
17
13
9
9
5
9
3
I
3
0
0
0
I
0
1
0
0
0
0
0
0
0
0
13
4
18
19037
18353
2
4
20
to887
8829
7519
6599
45
1598
1339
724
724
5067
4088
1252
971
692
570
7
31
62
83
87
93
146
166
214
261
276
296
265
324
228
190
124
66
52
40
28
27
35
30
17
16
11
12
10
3
5
7
5
6
3
1
3
9
15
3256
1724
5
6
7
8
9
10
11
12
13
4
22
78
155
304
424
598
636
645
556
472
407
326
230
118
75
51
26
9
10
1
2
3
11
45
1
1
4
48
117
183
178
175
188
198
180
159
178
156
121
to2
67
50
25
17
9
6
3
2
3
813
729
7
304
297
187
136
81
50
33
21
26
16
9
9
5
1
I
4
2
4
6
3
I
81
79
14
2
14
21
64
71
77
94
68
89
89
71
60
54
33
27
30
15
19
17
17
11
8
4
8
3
2
I
15
I
7
4
13
13
20
27
19
29
29
32
23
25
15
14
16
21
to
9
11
8
7
6
3
4
5
4
3
1
I
I
975
500
383
162
14
15
•
ß: Lenglh dislribullon or aU «<aplures
Release area:
1
length cm
to
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
4
I
6
30
50
76
112
69
20
4
6
5
6
16
6
2
2
2
34
35
36
37
38
39
40
41
42
43
44
45
2
2
2
8
16
to
36
24
21
12
12
13
14
10
14
13
7
I
3
2
2
I
I
2
3
2
9
13
39
22
7
3
9
4
I
3
1
4
5
23
120
309
385
355
285
207
101
49
16
3
2
68
127
149
181
203
213
195
192
195
168
137
132
109
65
77
37
35
14
9
7
5
9
3
3
1
6
8
10
25
36
38
40
36
37
31
21
13
13
6
6
5
3
4
10
18
31
43
44
26
15
6
4
I
I
2
24
36
46
57
83
95
136
168
147
170
183
184
197
122
118
90
66
48
25
28
19
11
11
4
3
4
1
4
3
2
2
5
13
8
22
37
32
37
26
33
36
57
60
81
45
51
58
44
24
11
17
11
4
5
2
1
1
8
16
27
28
22
21
11
8
6
8
6
4
16
8
16
6
2
6
8
4
2
2
4
7
6
7
19
16
19
28
37
29
31
42
30
22
12
9
8
5
5
5
8
6
3
1
1
5
12
13
17
18
9
12
20
10
10
14
7
5
3
2
2
5
I
2
2
1
2
2
I
I
•
1
1
3
4
6
7
7
11
6
6
7
4
3
8
2
3
1
2
2
I
I
1
2
1
1
2
46
47
48
49
50+
WlknOwn
Total
419
230
113
1860
1
8
5159
19
2418
82
2251
45
382
205
2085
720
236
15
380
171
91
20
Table 2.2.1.2. Number of plaice <25cm released by nursery area and recapture rates
in the first year and total in all years combined.
--------------------------------------------------------------------------------------------------------Total
Number
Recaptures 1st year Total recaptures
( n. - n.1+)
(n.)
remaining
releases after
Released
(Nj)
---------------------------------------------
1 year in sea
(N~+)
Release areas
#
name
J
J
J
No
%
No
%
J
---------------------------------------------------------------------------------------------------------
•
4422
1 Hvidesande
2 Sylt
3794
3 Norderney
4 WaddenSeaE
729
7233
5 WaddenSea W
6 Dutch coast
18353
8829
130
44
2.9
1.2
365
125
8.3
3.3
940 4 )
324 4 )
31
4.3
13.2
14.0
24.4
284 4 )
953
102
1765
3372
18.4
15.4
4724
1806
25.7
20.5
5253
2185
18.3
15.7
1741
253
26.4
18.9
2028
228
203
1455
28.0
628
433
35.6
44.6
1351
244
3328
7 Scheldt estuary
8 French-Belgian coast
6599
1339
1359
1206
210
9 Wash
10 Thames
724
4088
27
974
11 Rye Bay
971
324
3.7
23.8
33.4
570
1724
112
19.6
158
27.7
166
5.7
13.8
168
500
99
69
98
9.7
19.6
708
148
162
24
14.8
29
17.9
28
12 Poole Bay
13 Baie de Somme
14 Baie de Seine
15 Baie du Mt St Michel
--------------------------------------------------------------------------------------------------------1)
3)
2)
1) corresponds with release numbers in table 2.1.1.1
2) corresponds with recapture numbers in Appendix I
3) recaptures in the 1st year differ from recaptures in the first 4 quarters in Appendix I
since they are calculated separately as recaptures >12 months in sea
4) numbers have been changed in order to match with 25% total recaptures
21
Table 2.2.1.3. Percentage recapture values for each release area of plaice «25 cm at
release) by years after release.
-------------------------------------------------------------------------------------% recaptures in quarters after
release quarter
-------------------------------------------------name
#
0-3
4-7
8-11
11-15 16+
-------------------------------------------------------------------------------------Hvidesande
31
32
28
1
7
0
22
2
Sylt
23
37
10
7
28
21
16
Norderney
33
3
3
2
45
14
4
WaddenSeaE
35
3
1
WaddenSea W
64
28
1
5
5
22
4
1
+
Dutch coast
71
6
52
1
1
38
8
Scheldt estuary
7
French-Belgian coast
15
2
1
8
79
3
22
15
21
Wash
8
33
9
29
10 Thames
58
8
3
3
24
2
11 Rye Bay
65
6
3
12 Poole Bay
61
4
1
1
34
4
53
9
3
13 Baie de Somme
30
4
14 Baie de Seine
18
67
7
3
15 Baie Mont 5t Michel
83
0
13
3
0
-------------------------------------------------------------------------------------Release areas
•
•
22
Table 2.3.1.1. Production of l-group plaice along the continental coast ofthe North
Sea (from van Beek et al. 1990).
(area codes refer to RIVO otolith sampling areas)
---------------------------------------------------------------------------------------------------Area
#
Surface area
Density
Production
Total
%
production
---------------------------------------------------------------------------------------------------French/Belgium coast
400
Scheldt estuary
401
•
634
637
Dutch coast
402
403
Wadden Sea west
404a
610
612
616
617
Wadden Sea east
0.466
1732
3.162
5477
220
6.781
304
0.699
1492
212
2323
1.061
2465
1050
3.615
3796
1254
571
8.751
6.826
10973
3898
47
366
91
7.650
27.598
360
10101
292
3.205
618
619
60
50
0.566
73
28
620
268
1.836
492
Norderney
405
1254
1253
9.279
4513
2118
3.873
24.490
7181
5.4
6261
4.7
25623
19.3
11567
8.7
11626
8.7
69348
52.3
10973
11626
Sylt
406
407
0.7
959
8.751
1.214
404a
•
2058
959
17479
51870
---------------------------------------------------------------------------------------------------a #404 was allocated for 50% to the western and 50% to the eastern Wadden Sea.
23
Table 2.3.1.2. Production ofO-group plaice along the UK coast ofthe North Sea and
VIId (revised from Millner et ai. 1988; Millner and Whiting, 1990).
Area
#
N ECoast+
Scotland
HumberWash
Thames
Rye Bay
Poole
Surface area
Production Index
%
> 1000
50
1.0
4247
2748
2968
1536
57.3
29.7
2005
1499
475
145
9.2
2.8
•
Table 2.3.1.3. Production of2-group plaice in VIId from beam trawl survey data.
Catch reates are mean numbers per 30 minutes per 4m beam trawl
averaged over all stations in a rectangle over years 1988-91
Recatangle
Mean catch
rate
Allocation for
nursery area
Production
index
---------------------------------------------------------------------------------------------------30E8
30E9
0.83
1.96
0.83
1.96
Poole Bay
30FO
8.48
8.48
Rye Bay
30Fl
4.90
2.79
10.48
{ 2.0
•
{
{ 2.9
29Fl
29FO
5.9
0.6
28E9
28E8
1.10
1.36
1.10
1.36
27E9
1.13
1.13
27E8
no data
28E7
no data
5.9
{ 0.3
{
{ 0.3
est 1.50
Baie de Somme
9.11
Baie de Seine
3.89
Baie du
1.50
Mt St Michel
24
Table 2.3.1.4. Calculation ofthe power ofthe nursery areas in the North Sea and
English Channel from the surface area and the density of juvenile
plaice recorded during different pre-recruit surveys. Column A gives
production figures for l-group along the continental coast from Table
2.3.1.1. and for O-group along the UK coast. Column B gives the
relative nursery power within the North Sea. Collumn C gives
abundance estimates of 2-group plaice in the nursery areas in the
Channel which, are standardized to the North Sea level by the
production values for the two nursery areas common to both series:
Rye Bay and Poole Bay. Column D gives the final estimate of the
relative power ofthe different nursery areas. For Hvide Sande and
Baie de Mont Saint Michel production was assumed )*.
===================================================
A
B
c
D
Release areas
===================================================
•
1
2
Hvide Sande)*
Sylt
34674
69348
3
Nordemey
11626
4
5
6
7
8
Sum
WaddenSeaE
11567
WaddenSea W
25623
6261
Dutch coast
Scheldt estuary
7181
French-Belgian coast 959
9
10
Wash
Thames
Sum
•
18.15
36.30
6.09
6.05
13.41
3.28
052
3.76
0.50
50
90.0
0.11
}
2968
1536
652
3.37
6.45}
3.28
4554
10.0
167239
UKNE coast
18.66
3732
6.26
6.22
13.79
3.37
3.86
97.27
100.0
Sum North Sea
===================================================
1.01
1.04 10.48
11 Rye Bay
475
0.31
0.32
2.79
12 Poole Bay
145
0.88
9.11
0.91
13 Baie de Somme
0.38
0.39
3.89
14 Baie de Seine
0.15
15 Baie Mont St Michel )*
0.15
1.50
Sum Channel
2.80
2.73
===================================================
Sum North Sea + Channel
100
===================================================
25
Table 2.4.1.
Relative index of fishing effort (%) by quarter and sub-area in the
North Sea.
-------------------------------------------------------------------------------------------------------------------Q1
Q2
Q3
Q4
Total year
-------------------------------------------------------------------------------------------------------------------Moray Firth
0.16
0.14
Flamborough
3.15
Fisher Bank
0.15
4.43
0.13
3.94
3.49
0.58
15.01
1.97
4.42
4.87
2.97
14.23
German Bight
7.65
7.04
7.80
6.12
28.61
Transition Area
5.54
5.57
5.48
4.70
21.29
Southem Bight
4.76
6.69
5.38
3.44
20.27
-------------------------------------------------------------------------------------------------------------------Total
23.23
28.30
27.60
20.87
100.00
•
--------------------------------------------------------------------------------------------------------------------
•
26
Table 3.1.2.1 Average dispersion rate (a2 in miles 2. month- 1) during the first year
after release.
N
Release Area
North Sea
continental coast
169
UK coast
278
English Channel
Rye Bay (VIId)
•
6203
804
275
Pool Bay (VIId)
133
21
French coast VIId
French coast VIIe
60
198
178
94
32
•
27
Table 3.2.1.1 Nwnber of recaptures > 12 months in sea by subarea and release area
1
Hvide
Sande
recaptures
2
Sylt
3
4
0Id-
Wad
5
6
7
Wad Outch Scheldl
emey
east
west
1st quarter (spawning period)
SkaglKat
I
Fish.B
2
15
Ger.Bighl
7
10
3
Transit
I
1
Flam
2
4
4
Saum B
VIId
VIIe
VIIj
total
no position
28
4
2nd quarter
SkaglKal
3
52
Fish.B
Ger.Bight
14
Transil
6
array
fhm
Saum B
Vlld
VIIe
total
76
no position
17
3rd quarter
SkagIKat
Fish.B
Ger.Bight
Transit
Marray
Flam
Saum B
Vlld
Vlle
VIIh
total
no position
4th quarter
SkaglKal
Fish.B
Ger.Bighl
Transit
Marray
Flam
Saum B
39
14
1
3
3
25
38
3
80
2
10
11
12
13
14
15
Wash Thames
70
70
70
70
7E
Dover
estuary
Rye Poole
BlSomme
8
I
46
5
9
2
42
5
206
10
4
3
3
6
81
6
6
0
3
1
8
2
I
154
7
280
25
13
11
1
12
3
4
98
48
5
39
120
5
35
2
2
8
60
1
2
117
4
1
164
1
25
4
20
6
219
36
8
7
6
1
10
3
6
139
39
4
26
81
2
19
I
56
14
3
10
4
4
293
12
3
476
35
3
4
60
157
12
8
191
II
6
3
9
0
10
17
14
1
14
8
1
3
5
2
23
19
1
27
1
30
14
9
I
I
41
36
11
3
2
2
1
27
41
18
3
97
39
6
2
2
4
2
27
31
3
5
71
2
1
4
90
2
4
8
4
2
209
27
284
34
lJ8
117
14
15
0
4
66
25
1
16
46
4
11
2
8
7
39
5
114
4
1
65
1
2
84
4
2
1
10
1
21
186
31
82
13
0
22
80
Jl
102
13
19
18
15
89
5
11
30
2
4
11
7
30
4
2
2
0
99
2
1
106
5
135
43
4
76
36
10
2
6
7
35
14
3
2
11
1
33
58
14
5
1
4
33
1
127
42
41
3
19
1
20
2
9
159
3
1
2
1
32
5
23
57
108
48
410
62
2
1
13
0
4
II
4
0
1
0
1
730
118
1
2
3
0
3
81
183
223
4
73
679
120
2
1394
168
17
6
23
0
6
4
10
0
1
0
0
63
208
190
3
93
407
81
23
1
1069
160
0
0
1
27
112
106
0
69
338
51
9
0
1
714
136
2
8
9
8
8
0
17
0
6
1
7
0
2
3
10
31
23
9
1
7
7
3
14
2
4
1
8
6
10
0
16
0
5
0
1
33
13
Total year
SkagIKal
4
fish.B
123
Ger.Bighl
49
Transit
9
Marray
Flam
Saum B
6
12
1
4
5
141
19
3
1
2
26
8
17
10
328
150
89
293
1
26
33
8
1
5
26
3
7
196
3
VIId
23
772
29
8
2
2
1
VITe
VlIh
Vllf,g
10
7
1
295
12
3
387
32
27
11
17
2
50
18
20
5
1
5
1
1
VIIi
total
no position
10
191
72
44
9
48
23
723
89
1226
125
424
25
491
44
43
lJO
0
66
345
135
91
18
42
4
Sum af
BISeine BlMichel
3
5
34
8
17
22
1
I
I
7
15
13
2
2
5
10
1
54
9
3
13
3
VIId
VIIe
VIIh
Vllf,g
total
no position
coast estuary
8
4C
69
26
0
0
5
0
')
9
194
560
626
7
283
1834
304
75
1
1
1
3894
582
•
•
Table 3.2.1.2. Origin of plaice in spawning areas in percentages
1
Hvide
2
Syll
Sande
3
4
5
6
7
8
9
NordEasl
Wesl OUlch Scheldl FRJBelg Wash
emey Wadsea Wadsea coasl estuary
Coasl
1st quarter (spawning period)
SkagIKal
70.7
Fish.Bighl 55.0
42.6
Ger.Bighl
9.7
8004
4.6
Transil
6.8
7.5
Moray Fyrth
Aamborough
14.2
South Bighl 6.6
9.6
7.4
VIId
20.0
1.0
3.3
24.2
1.8
12.5
2.5
VlIe
VIlh
Vllf,g
VlIa
VlIj
e
2nd quarter
SkagIKal 100.0
Fish.Bighl 39.9
Ger. Bighl
13.1
Transil
17.8
Morray
Aamborough
South Bighl 1.2
VlId
VlIe
VIlh
Vllf,g
VlIa
VlIj
Jrd quarter
SkagIKal
Fish.Bighl 44.2
Ger.Bighl
16.8
Transit
1.7
Morray
Aamborough
South Bighl
VlId
VlIe
VIIh
Vllf,g
VlIa
Vllj
•
4th quarter
SkagIKal
Fish.Bighl 48.3
Ger.Bighl
26.0
Transil
3.9
Morray
Aamborough
South Bighl 2.3
10
Thames
eslUary
11
70
Rye
12
70
Poole
13
14
15
70
70
7E
B/Somme BISeine B/St. 11
9.3
1.5
41.1
0.1
4.7
0.4
2.6
4.2
45.3
17.2
28.8
1.0
5.4
5.1
4.2
0.6
8.9
17.5
15.7
0.6
4.4
1.4
100
100
7.2
5.9
JOO
JOO
64.7
0.9
6.9
12.1
2.7
3.3
6.9
JOO
1.4
100
25.2
11.7
2.5
10.5
8.4
7.0
JOO
6.9
15.1
100
JOO
100.0
100
57.9
59.8
6.9
52.6
48.7
57.9
0.9
12.5
9.9
10.1
2.6
1.3
13.3
5.5
0.3
8.6
13.4
3.4
6.7
0.6
0.6
15.7
6.1
2.5
3.5
19.4
0.5
4.8
47.1
2.4
46.1
10.2
12.4
0.6
4.1
17.8
16.7
4.0
45.9
3.3
5.4
0.4
8.1
0.7
10.8
2.0
2.4
0.5
4.4
1.3
12.0
1.3
3.2
0.2
0.1
2.3
22.6
18.0
7.4
24.1
0.2
0.6
1.6
0.0
1.3
18.9
1.6
15.6
0.5
0.8
2.2
0.2
62.6
65.1
0.6
1.8
67.3
56.9
2.0
3.7
9.2
0.3
0.2
0.7
14.8
15.4
6.1
14.6
11.8
0.4
0.3
2.3
15.9
13.9
15.9
14.4
25.6
JOO
JOO
JOO
JOO
0.6
100
2.9
37.2
11.8
3.0
7.0
12.1
5.1
16.7
1.8
17.4
2.1
4.4
11.5
9.3
13.2
6.2
2.5
3.7
8.4
l.l
VIId
0.4
3.9
24.0
0.6
3.4
0.4
3.0
3.7
34.4
9.0
0.4
11.5
1.3
l.l
18.4
6.6
22.1
VlIe
VIIh
Vllf,g
1.0
0.4
0.6
2.6
1.9
62.5
0.7
0.7
5.0
21.8
6.6
19.4
14.5
100
100
JOO
JOO
0.7
0.7
1.9
58.1
8.7
6.4
4.8
21.0
100
100
100
100
6.5
11.3
100.0
100.0
49.5
53.9
45.8
JOO
JOO
JOO
JOO
100
100
0.8
JOO
JOO
25.7
JOO
JOO
JOO
11.6
33.4
15.4
14.8
9.1
15.3
100
100.0
100
0.5
0.3
2.7
15.3
15.6
7.3
13.4
15.1
100
100
100
100
100
100
VlIa
VlIj
Total year
Skag/Kal
39.1
Fish.Bighl 43.8
Ger.Bighl
15.5
Transil
6.7
Moray Fyrth
Aamborough
South Bighl 2.6
VlId
Vlle
VIIh
Vlli,g
VlIa
Vllj
56.8
53.7
60.6
34.7
0.8
9.1
6.6
6.8
7.4
6.5
3.3
2.8
0.6
9.8
10.6
2.9
7.9
0.7
1.3
0.5
3.7
28.9
8.1
3.6
43.7
9.5
12.7
JOO
0.4
5.2
0.9
9.8
2.3
2.8
0.2
0.3
2.2
11.7
0.8
15.9
7.6
19.5
0.2
0.1
1.3
3.1
2.7
0.3
1.6
64.9
61.6
0.5
1.3
0.0
0.6
1.3
0.4
40.5
5.1
6.2
6.9
.0
13.9
0.7
13.3
6.6
8.0
100.0
JOD
JOO
JOO
100
100.0
100
100.0
?Q
Tab1e 3.2.2.1
Contribution of a p1aice nursery area to different spawning areas in the North
Sea
Wad
east
5
Wad
west
1st quarter (spawning period)
Fish.B
79.7
40.1
0.0
Ger.Bight
9.6
51.6
19.1
Transit
1.9
0.0
8.8
Morray
0.0
0.0
0.0
Flarn
0.0
0.0
31.0
41.0
South B
8.8
8.3
100
Sum
100
100
5.2
11.1
23.3
0.0
3.2
57.1
0.0
1.9
15.3
0.0
2.9
79.8
100
2nd quarter
Fish.B
78.5
Ger.Bight
13.3
Transit
7.2
Morray
0.0
F1am
0.0
South B
1.0
Sum
100
63.2
33.6
0.0
0.0
0.0
3.2
7.0
52.9
16.7
0.0
14.0
9.3
100
3rd quarter
Fish.B
Ger.Bight
Transit
Morray
Flarn
South B
Sum
80.2
18.0
1.8
0.0
0.0
0.0
1
Hvide
Sande
2
Sy1t
4
8
7
6
Dutch Sche1dt
coast estuary
4C
Dover
0.0
1.2
14.5
0.0
5.7
78.7
0.0
3.0
5.5
0.0
2.4
89.2
0.0
0.0
5.5
0.0
91.4
3.2
100
14.8
0.0
12.3
0.0
46.3
26.6
0.0
0.0
0.0
0.0
100.0
0.0
100
100
100
100
100
2.6
40.7
25.2
0.0
5.3
26.2
1.6
7.6
29.5
0.0
1.2
60.1
0.0
2.8
25.2
0.0
1.8
70.1
1.7
0.4
3.6
35.7
0.0
58.7
0.0
1.0
0.0
52.8
45.0
1.1
3.3
1.4
1.7
34.5
29.5
29.6
0.0
0.0
21.0
0.0
79.0
0.0
100
100
100
100
100
100
100
100
42.2
23.1
28.2
0.0
6.5
0.0
10.1
63.0
26.9
0.0
0.0
0.0
3.9
56.8
22.7
0.0
6.5
10.1
1.4
5.7
25.3
13.2
3.9
50.5
0.0
3.0
29.1
0.0
6.0
61.9
2.0
3.1
8.8
0.0
4.9
81.1
0.0
0.0
2.3
47.5
50.2
0.0
2.1
0.9
6.8
25.9
28.2
36.3
100
100
100
100
0.0
0.0
26.8
0.0
18.6
54.6
100
100
100
100
100
100
4th quartel'
Fish.B
Ger.Bight
Transit
Morray
Flarn
South B
Sum
67.2
26.9
2.5
0.0
0.0
3.4
39.7
32.1
16.7
0.0
0.0
11.4
0.0
17.1
22.3
0.0
30.1
30.5
4.4
35.0
17.3
0.0
6.5
36.8
0.7
5.5
20.7
0.0
3.0
70.0
0.0
2.9
10.6
0.0
1.3
85.2
0.0
1.2
6.3
0.0
2.1
90.4
0.0
2.6
0.0
0.0
97.4
0.0
3.6
2.6
11.3
0.0
47.1
35.5
0.0
0.0
100.0
0.0
0.0
0.0
100
100
100
100
100
100
100
100
100
100
total year
Fish.B
Ger.Bight
Transit
Morray
Flarn
South B
Sum
78.0
15.5
3.8
0.0
0.0
2.7
51.2
32.3
10.5
0.0
1.9
4.1
6.7
43.2
17.9
0.0
15.8
16.4
3.8
36.9
22.6
0.0
5.6
31.1
1.2
5.3
23.4
2.9
2.6
64.6
0.0
2.6
21.0
0.0
3.3
73.0
1.4
1.5
5.6
13.7
1.9
75.9
0.0
0.7
1.8
33.2
63.4
0.9
3.9
1.2
7.2
17.6
36.0
34.0
0.0
0.0
26.2
0.0
46.4
27.5
100
100
3
Norderney
100
100
100
100
100
100
10
9
Wash Tharnes
estuary
100
30
100
•
•
Table 3.3.1.1. Origin ofplaice in management areas.
Management areas (A) receive their recruits from nurseries (B)
B
A
management
areas
receives recruits originating from
nurseries in management area:
VIId
VIIe
99.7
0.3
38.0
61.3
33.9
1.9
0.0
0.7
13.3
North Sea
Total year
North Sea
VIId
VIIe
others
52.8
98.1
1st quarter
North Sea
VIId
VIIe
others
99.9
57.0
50.6
95.6
2nd quarter
North Sea
VIId
VIIe
others
3rd quarter
North Sea
VIId
VIIe
others
4th quarter
North Sea
VIId
VIIe
others
100
100
0.1
0.0
43.0
0.0
15.1
100
100
100
100
29.3
4.4
0.0
0.0
100.0
99.5
24.3
59.0
75.7
29.7
0.0
100.0
0.0
0.0
11.3
0.0
99.9
38.2
36.5
100.0
0.1
62.8
63.5
0.0
0.0
0.0
0.0
50.8
100
100
0.0
0.2
61.3
31.8
0.0
99.8
37.0
total
0.5
100
100
1.8
17.4
100
0.0
100
100
100
100
100
0.0
100
100
100
100
31
Table 3.3.2.1. Contribution ofnurseries to management areas.
Nurseries (B) send recruits to management areas (A)
A
B
nurseries in management area
to:
management
area
North Sea
VIId
send recruits
VIIe
Total year
North Sea
VIId
VIIe
others
total
97.2
1.4
0.4
1.0
100.0
9.1
80.9
9.3
0.7
100.0
0.0
20.0
80.0
0.0
100.0
1st quarter
North Sea
VIId
VIIe
others
total
96.6
2.2
0.5
0.7
100.0
5.2
80.2
13.0
1.6
100.0
0.0
0.0
100.0
0.0
100.0
2nd quarter
North Sea
VIId
VIIe
others
total
97.3
1.5
0.4
0.8
100.0
7.5
83.6
8.9
0.0
100.0
0.0
33.3
66.7
0.0
100.0
3rd quarter
North Sea
VIId
VIIe
others
total
98.6
0.9
0.5
0.0
100.0
12.9
79.6
6.3
1.2
100.0
0.0
0.0
100.0
0.0
100.0
4th quarter
North Sea
VIId
VIIe
others
total
95.4
1.2
0.2
3.2
100.0
4.9
82.5
12.6
0.0
100.0
0.0
0.0
0.0
0.0
0.0
•
32
Figure 2.2.1.1 Plaice tagging exchange format
::\:)))f
alfanumerjc
,,)': ':)(2,
')}'{{';j~
axp number
}:
relaasa day
ralaaSa
month
:{':::::\iJ!
({{::t:s
ralaasa yaar
}}:)}4
:::::::::t$
tag serjes
ral lat dagr
ral lat mln
ral Ion dagr
rel Ion mln
ral EW coda
treatment
tag numbar
ralaasa langth
rel
maturlty
ral condition
rel plgm
raC langth
recapture sex
rac maturity
birth yaar
raC day
raC month
raC yaar
raC lat dagr
rac lai mln
raC Ion dagr
rac Ion min
rac EWcode
~
natiooaljty
~
otolith sl2a
L1
33
E3 E4
46
45
44
43
-
9
E5 'E6 E7 ES E9
~
f7
Y
.' l"
r
40
39
38
'A
~7 )'l~
-
37 _~,j<J
36 ~
35
\
1,5
~
~
/
5
I
Hvide
Sande
r:
I
~
I
\
"
II
,
,
~
IJ
,
\
i
~
,I
-
y~~
Sylt
0
" .
I~
;
i
,
.I
I
~~
(
At
I .~
!
\
-' 5
I~
I
~~
~
fJ}~ ~-
-
~rJYT
e-
.~
I
J
j
~
'0;7
~(j
'/~
I
)
r
'42 -~~/..
41 ~.)~/f
Fa F9
F5 F6 F7
,
-fl.>.
rI
F2' F3 F4
/
~......r1,F
r..
FO Fl
I
~ ~?
~
.. ··(~\~
"
- Eostern
.~~
~
~
crr<?
26
25
n
~I
I
I
c
\
.......
M
Boy de Seine
~ Boie du mt,St Michael
I
I
I
I
I
5
Woddens ee
~I 'y!estern Waddensee -
Wosh orf
().
-
~prq~ney
~
CI("
34 -- (
I".
33 ~
j
coasl
Thomes orec
32
~~
.
_ ,..rl
~ Zeelond estucry
v c- '"\.
J-~
.~
31
I
French I Belgion coost
[)
POOle~_~e [7
30
. r~
I
j -' .Boy de Somme
~
29
rLJ.
.-,» •
"
28
.. ~
27
~
I
- 5
'-
,
-
I
I
I
I
, .. I
·t
Fig. 2.3. 1.1. Division of Ploice nursery areos.
34
E3 E4
46
45 ~.
44
43
ES .E6 E7 E8 E9
.L
].,yr. /
~r
Ü
j
r,,, '~
J
'\
()
j
\
~ambOrOUgh
r
28
27
26
VII j l~]
'i I
vmJ
~
c'
~
TRANSITIONAL
AREA
~
r
-
-
.•.
~
-
-
r(--'-'-'-~
,~
,.,)
-
~
~~OUf~ern
/
Bight
.
I ~~
-
$
.~
-"'\
Yr'-r- .~,-y
.~
." S"'"l
•\
."
/)'
./V'~
I
-
. j.r Ii"',\
--
VII
Vlle
~I
~~
iI
'J
....
1-
,ö
I
)
-zr-~
~,
~
I
~~
y~~
,...,
I
\
-
.1\
i
-
~ -
~
I
~
~
(J}~( ~
Fisher Bight
..
-~~
-
.L-
I
lr-5
I
'~
?
SkageracK
I
Kalle
~,~y
VlIg
f
'\
37 ~ ~;v
36 ~\
35 ~ ~
34 ~ (
31
30
I
.-.
~ 'I~ß~
39 n1.:l
32
I
~
Moray Firth
~t
42 . . ~~b
41 ~~/t
~~. ~
40
33
F8 F9
F2' F3 F4 F5 F6 F7
~f
;::7}:
'8 :
FO Fl
r-'''/
. I'
-
d
Ir
-
),'---v r -
-
.~
-v
I
I
I'
I
I
I
I
I
I
I
I
I
Fig. 2.3,2.1. Division of plaice spawning area
35
5t
w
4
2
o
2
4
6
8 E
57
55
49
Figure 3.1.1.1.
Quarterly centres of distribution of recaptures of plaice tagged in
different nursery release areas (recaptures >12 months in sea
only)
36
--- ------ - - - - - - 1
3pt running mean
160
140
'--IVb,c continent
120
OoIVb,c UK
100
miles
80
···Vlld France
60
~Vlld Rye
40
Bay
••- Vlld Pool Bay
20
0
0
2
4
6
810121416
Time after release (quarter)
3pt running mean
700 , . . . - - - - - - - - - - - ,
B
600
··-IVb,c continent
500
Q-IVb,c UK
milesl\2
/month
400
"'Vlld France
300
~
Vlld Rye Bay
200
-.- Vlld Pool Bay
100
o
o
2
4
6
8
10 12 14 16
Time after release (quarter)
3pt running mean
16000 . - - - - - - - - - - - ,
c
12000
··-IVb,c continent
a-IVb,c UK
milesl\2
8000
"'Vlld France
~Vlld
4000
Rye Bay
-.- Vlld Pool Bay
o
Time after release (quarter)
Fig.3.1.2.1. Migration parameters (3pt running mean) in relation with the number of
quarters after release. A. distance from position of release (miles); B - dispersion rate
(miles2.month- 1); C- surface area over which recaptures are distributed.
37
Males
140 . , . - - - - - - - - - - - - - - . . . . . . ,
,'-01
miles
·'·03
Fig. 3.1.3.1a. Migration parameters of males in relation time after release in absolute
quarters far separate experiments carried out in Ql, Q2, Q3 and Q4.
A. distance from position ofrelease (mHes); B - dispersion rate (miles2.month- 1);
c-
surface area over which recaptures are distributed.
38
Females
90
-r----------------,
A
80
70
·'-01
60
miles
50
40
•.• 03
30
20
4
3
5
7
6
8
Time after release (quarter)
Females
450 - . - - - - - - - - _ - - - - - ,
B
400
350
'·-01
300
miles"2 250
Imonth 200
·"03
150
+04
100 _ / -
5
~ 1_-+_-+_-+_-+_-+_-+__
1
2
4
3
5
6
7
8
Time after release (quarter)
Females
4000
,.--------------n
c
3500
3000
·'-01
2500
~Q2
miles"2 2000
1500
/-
1000
•
500
./
. /I
o I
1
2
+04
..
~
3
5
6
7
8
Time after release (quarter)
Fig. 3.1.3.1 b. Migration parameters of females in relation time after release in absolute
quarters for separate experiments carried out in Q1, Q2, Q3 and Q4.
A. distance from position ofrelease (miles); B - dispersion rate (miles2.month- 1);
c-
surface area over which recaptures are distributed.
39
100 " T ' " " - - - - - - - - - - - : - - - - - . ,
A
90
80
70
60
miles
·--Males
50
D-Females
40
30
20
10
o+-+--+--+-++-+-+-+-+--.-t-1
o
1
2
3
4
5
6
7
8
9 101112
Time after release (quarter)
3 5 0 "T'""------:------,r-----.,
300
250
miles"2
/month
200
·--Males
150
D-Females
100
50
0+-+--+-++-+--+--+--+-+-+-+--1
o
1
2
3
4
5
6
7 8
9 101112
Time after release (quarter)
6000
-r-----"""":""-----.-----,
5000
4000
·--Males
miles"2 3000
D-Females
2000
1000
o -f-IF--+--+--+-+-+--+--+--+--+--+--I
o
1 2
3
4
5 6
7 8 9 101112
Time after release (quarter)
Fig. 3.1.3.2. Migration parameters of male and female plaice in relation time after release
in absolute quarters averaged over the separate experiments carried out in Ql, Q2, Q3 and
Q4. A. distance from position of release (mHes); B - dispersion rate (miles2.month- 1);
surface area over which recaptures are distributed.
c-
40
APPENDIX I
Recaptures by quarter and rectangle
REC43F8
Release Rectangles:
0
2
year:
quarter:
Rectangle
REC45Gl
REC44F7
REC43F3
REC43F4
REC43F7
REC43F8
REC43F9
REC42F6
REC41F5
REC41F6
REC41F7
REC41F8
REC40F3
REC40F4
REC40F5
REC40F7
REC40F8
REC39F2
REC39F4
REC39F6
REC39F7
REC39F8
REC38F2
REC38FS
REC38F7
REC37F4
REC37F6
REC37F7
REC36F2
REC36FS
REC3SF4
REC34F3
Total
No position
Not used:
APPENDIX I
3
4
1
1
REC40F7
2
3
(Hvide Sande, 2nd quarter)
4
2
1
2
3
4
3
1
2
3
4
4
1
4+
Total
1
1
1
1
1
1
1
1
1
1
1
1
1
3
1
1
1
1
3
1
1
1
1
1
1
1
1
2
1
4
1
1
1
1
1
1
1
1
3
1
1
1
1
1
1
1
1
1
1
1
1
1
0
0
1
0
1
0
4
0
8
2
3
0
1
2
8
0
12
1
9
0
3
1
1
2
1
9
1
2
1
1
0
0
0
0
0
2
1
1
1
1
1
1
3
1
2
3
5
1
1
1
2
2
11
3
1
3
1
1
5
1
1
1
2
2
1
1
1
1
1
63
10
REC40F7 (Hvide Sande, 3rd quarter)
Release Rectangles:
year:
q;uarter:
Rectangle
REC45E6
REC44F2
REC44F4
REC44F8
REC44F9
REC43F3
REC43F5
REC43F7
REC43F8
REC43F9
REC42F5
REC42F6
REC42F7
REC42F8
REC41F3
REC41F5
REC41F6
REC41F7
REC41F8
REC40F2
REC40F4
REC40F5
REC40F6
REC40F7
REC40F8
REC39F2
REC39F3
REC39F4
REC39F6
REC39F7
REC39F8
REC38F2
REC38F6
REC38F7
REC38F8
REC37F4
REC37F5
REC37F6
REC37F7
REC36F5
REC35F2
REC35F4
REC34F3
REC34F4
REC33F3
Total
No position
Not used:
APPENDIX I
0
3
1
4
]
2
2
3
4
1
3
2
3
4
1
2
3
4
4
4+
1
2
Total
1
1
1
1
1
1
1
1
1
4
2
4
4
4
6
7
1
6
7
6
39
1
6
1
6
48
29
2
2
4
10
7
7
1
1
2
1
1
2
1
5
1
1
3
2
1
7
1
2
244
48
1
1
1
1
1
1
2
1
2
2
1
1
1
1
1
2
2
2
2
1
1
1
1
2
1
1
1
4
1
1
2
1
1
1
1
1
2
1
1
17
3
1
8
1
3
2
2
1
1
2
1
2
3
1
1
1
1
1
1
1
4
16
25
7
2
8
2
2
2
1
1
4
1
1
1
1
1
2
1
3
2
2
2
4
1
2
2
1
2
1
1
1
1
1
2
1
2
1
1
1
1
1
1
1
1
1
1
1
2
1
1
1
1
1
1
1
1
7
0
3
0
12
1
74
11
16
3
12
3
1
12
1
43
4
19
4
14
7
1
12
11
11
2
2
0
0
0
0
0
0
0
1
2
REC38F7
Release Rectangles:
0
2
year:
quarter:
Rectangle
REC42F7
REC41F4
REC41Gl
REC40F4
REC40F7
REC39F6
REC38F4
REC38F6
REC38F7
REC38F8
REC37F2
REC37F4
REC37F7
REC37F8
REC36FS
REC36F7
REC35F2
REC35F4
REC3SFS
REC34F4
REC33F3
Total
No position
Not used:
APPENDIX I
(Sylt, 2nd quarter)
4
1
2
3
4
1
4
3
2
1
3
2
3
4
1
2
3
4
4+
Total
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
2
1
2
1
1
2
1
2
4
2
1
4
1
1
1
1
1
1
1
2
1
1
0
0
9
1
3
0
2
0
11
0
5
0
1
3
0
1
1
4
0
3
0
1
0
1
0
0
0
0
0
0
0
0
0
0
1
6
3
5
2
2
1
11
4
2
2
1
2
1
1
1
43
3
Release Rectangles:
0
3
year:
q:uarter:
Rectangle
REC44F4
REC43F6
REC43F7
REC43F8
REC41F6
REC41F7
REC40F2
REC40F3
REC40F4
REC40F5
REC40F6
REC40F7
REC40F8
REC39Fl
REC39F6
REC39F7
REC39F8
REC38F6
REC38F7
REC37F4
REC37F5
REC37F6
REC37F7
REC36F2
REC36F4
REC36F5
REC36F6
REC35F5
REC34F2
Total
No position
Not used:
REC39F8
4
3
4
2
1
2
3
4
3
1
2
3
4
4
4+
1
2
Total
1
1
1
1
1
2
1
1
1
1
1
1
1
1
1
1
1
3
1
1
1
1
1
1
1
1
1
3
1
1
1
1
1
1
3
1
1
1
1
1
1
1
1
2
1
1
1
1
1
2
1
1
1
2
0
0
0
1
1
7
1
5
0
4
1
2
1
10
0
2
1
2
0
5
0
7
2
7
0
0
0
1
3
1
0
0
3
4
4
1
2
3
7
1
1
1
1
1
1
1
2
1
1
1
6
9
1
1
6
7
2
6
1
1
2
1
3
1
2
1
1
1
1
64
9
0
year:
q:uarterRectangle
REC43F4
REC41F7
REC40F8
REC39F6
REC39F7
REC37F8
Total
No position
APPENDIX I
2
1
Release Rectangles:
Not used:
1
1
REC38F7 (Sylt, 3rd quarter)
0
4
REC39F7
1
1
2
3
REC38F7 (Sylt, 4th quarter)
4
2
1
2
3
4
3
1
2
4+
Total
1
1
1
1
1
1
0
0
0
0
1
0
1
0
0
0
0
0
2
0
0
0
0
0
0
0
0
0
0
0
1
0
0
0
1
0
0
0
0
0
0
4
1
1
1
1
1
1
6
0
REC37F8
Release Rectangles:
0
3
year:
quarter:
Rectangle
REC37F5
REC37F7
REC36F3
Total
No position
Not used:
4
2
0
3
4
1
3
2
3
4
1
2
3
4
4
4+
1
2
Total
1
1
1
0
0
0
0
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
2
3
4
4
1
2
3
0
0
3
0
1
year:
quarter:
Rectangle
REC43F7
REC40FO
REC40F4
REC40FS
REC39E8
REC39F4
REC39F8
REC38F4
REC38F5
REC38F6
REC38F7
REC37Fl
REC37F2
REC37F4
REC37F5
REC37F6
REC37F7
REC37F8
REC36FO
REC36F2
REC36F3
REC36F4
REC36F5
REC36F6
REC36F7
REC35F3
REC3SF4
REC34F2
REC34F3
REC34F4
REC33F3
REC32F2
REC30E9
Total
No position
APPENDIX I
2
2
1
1
REC37F7 (Norderney, 4th quarter)
Release Rectangles:
Not used:
1
1
(Norderney 3rd quarter)
0
4
1
1
2
3
4
2
1
3
2
3
4
1
4+
Total
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
2
4
1
1
1
1
1
1
3
1
1
1
1
1
1
1
2
1
14
2
2
1
1
3
1
3
2
3
2
1
7
2
4
1
1
1
1
2
1
4
1
1
2
2
1
1
1
1
1
1
1
1
1
1
1
1
2
1
1
1
1
1
1
1
2
1
1
1
1
1
1
3
1
1
1
1
0
7
0
6
3
7
2
2
0
4
3
12
2
6
4
3
2
3
1
4
1
5
2
0
3
2
0
4
3
2
2
3
0
1
5
1
1
71
28
Release Rectangles:
year:
quarter:
Rectangle
REC43F9
REC42F6
REC42F8
REC41F3
REC41F6
REC41F7
REC41F8
REC40F2
REC40F4
REC40F5
REC40F6
REC39Fl
REC39F2
REC39F3
REC39F4
REC39F5
REC39F6
REC39F7
REC39F8
REC38FO
REC38F1
REC38F2
REC38F3
REC38F4
REC38F5
REC38F6
REC38F7
REC38F8
REC37FO
REC37Fl
REC37F2
REC37F3
REC37F4
REC37F5
REC37F6
REC37F7
REC37F8
REC36FO
REC36F2
REC36F3
REC36F4
REC36F5
REC36F6
REC36F7
REC36F8
REC35F2
REC35F3
REC35F4
REC35F5
REC35F6
REC35F7
REC34F2
REC34F3
REC34F4
REC34F5
REC33F2
REC33F3
REC33F4
REC32F1
REC32F2
0
3
REC35F6
REC35F7 (Eastern part of Dutch Waddensea 3rd quarter>
1
4
1
2
3
4
1
2
1
2
3
4
3
1
2
3
4
4
4+
1
2
Total
2
1
1
1
3
1
1
1
1
1
1
2
2
2
1
6
4
1
3
2
3
5
1
2
1
1
1
1
1
1
1
1
1
1
2
1
1
1
1
1
2
1
1
2
1
1
1
3
1
1
2
1
1
2
1
1
1
1
1
1
1
1
2
1
1
2
2
1
2
1
4
2
3
2
1
1
4
8
5
11
8
1
1
1
1
2
1
3
5
5
1
1
58
3
145
16
1
20
4
2
1
15
4
1
5
2
1
2
2
1
8
3
2
2
1
1
1
2
1
34
30
4
1
2
1
1
10
4
2
2
2
1
3
7
3
1
5
1
4
5
23
16
1
1
5
9
12
11
2
1
3
13
14
1
1
1
1
1
1
1
1
2
2
1
1
1
2
1
1
3
1
1
2
2
1
1
1
1
1
2
2
1
3
1
1
1
1
1
1
1
2
2
3
1
2
1
1
1
2
10
11
89
3
8
18
19
2
6
1
1
2
1
3
3
5
1
2
REC32F3
REC32F4
REC31Fl
REC31F2
REC30FO
REC30Fl
Total
222
No position
8
Not used:
8
APPENDIX I
1
1
2
13
29
9
1
4
9
15
11
3
1
1
1
1
1
10
3
1
3
4
1
2
1
5
2
1
2
4
17
2
3
1
1
5
1
2
1
2
1
1
1
1
1
1
1
12
1
1
1
4
1
14
2
1
7
6
1
1
1
2
2
3
5
6
2
1
2
1
1
1
5
3
2
1
1
1
1
1
1
1
1
1
25
9
2
1
1
5
3
1
1178
78
1
1
4
1
2
48
0
36 224 181
7
7
4
1
45
5
1
72 117
2 10
94
8
34
5
18
1
24
7
16
4
6
3
8
0
2
3
4
9
15
25
39
38
11
2
10
12
17
111
182
27
2
3
25
52
35
283
25
5
29
20
2
4
45
9
2
14
24
5
6
Release Rectangles:
year:
quarter:
Rectangle
REC44F4
REC44F9
REC43F8
REC42F3
REC42F8
REC41FS
REC40E9
REC40F4
REC40F6
REC40F7
REC39Fl
REC39F2
REC39F4
REC39F5
REC39F6
REC39F7
REC38E9
REC38F1
REC38F2
REC38F4
REC38F5
REC38F6
REC38F7
REC37Fl
REC37F2
REC37F3
REC37F4
REC37F5
REC37F6
REC37F7
REC37F8
REC36F1
REC36F2
REC36F3
REC36F4
REC36F5
REC36F6
REC36F7
REC35F2
REC35F3
REC35F4
REC3SF5
REC35F6
REC35F7
REC34F2
REC34F3
REC34F4
REC34F5
REC33F2
REC33F3
REC33F4
REC32F2
REC32F3
REC31FO
REC31Fl
REC31F2
REC31F3
REC30FO
REC29Fl
Total
No position
Not used:
APPENDIX I
0
4
REC35F6 (Eastern part of Dutch Waddensea 4th quarter)
1
1
2
3
4
2
1
2
3
4
3
1
2
3
4
4
4+
1
2
3
Total
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
2
1
1
1
1
2
1
2
2
1
1
1
1
1
1
1
1
7
3
8
1
1
4
3
1
1
1
1
15
6
1
1
1
1
1
3
2
1
1
2
2
1
1
1
3
4
4
11
1
6
33
6
6
1
6
2
2
1
2
5
4
3
1
8
8
1
2
2
1
9
2
1
3
2
2
1
2
1
1
11
4
2
2
1
1
1
2
5
2
1
4
1
1
3
1
7
1
1
2
12
2
1
2
3
1
1
5
6
1
1
2
4
1
2
1
1
1
1
1
2
1
1
1
2
1
2
1
2
1
1
1
2
1
2
1
1
2
33
6
19
3
10
1
14
2
1
1
2
3
3
1
3
3
1
4
9
8
1
2
1
1
3
3
1
1
1
3
1
1
2
1
1
2
1
1
20
0
1
1
1
31
2
1
1
71 120
5
3
41
1
32
1
49
13
5
3
0
1
4
1
2
0
3
0
8
5
14
7
1
1
1
2
1
1
1
1
1
1
1
1
1
2
4
2
1
1
1
3
5
6
4
6
1
4
9
8
6
17
3
1
5
2
12
58
28
11
3
19
16
12
28
8
6
19
27
2
3
42
8
10
31
1
1
6
2
2
2
462
47
Release Rectangles:
year:
Quarter:
Rectangle
REC44E7
REC41F3
REC40Fl
REC40F2
REC40F6
REC40F7
REC39F1
REC39F2
REC39F3
REC39F4
REC39FS
REC39F7
REC38F1
REC38F2
REC38F4
REC38FS
REC38F6
REC38F7
REC37Fl
REC37F2
REC37F3
REC37F4
REC37F5
REC37F6
REC37F7
REC37F8
REC36FO
REC36F2
REC36F3
REC36F4
REC36F5
REC36F6
REC36F8
REC35Fl
REC35F2
REC35F3
REC35F4
REC35F5
REC34F2
REC34F3
REC34F4
REC34F5
REC33F2
REC33F3
REC33F4
REC32F2
REC32F3
REC31Fl
REC31F2
REC31F4
REC30E6
REC30E9
REC30FO
REC30F1
REC29E5
REC29Fl
REC28E9
Total
No position
Not used:
APPENDIX I
0
1
REC35F5
2
3
4
REC34F5 (Western part of Dutch Waddensea, 1st quarter)
1
1
2
3
4
2
1
2
3
4
3
1
4+
2
3
4
Total
1
1
1
1
1
1
1
3
1
1
1
1
1
1
1
1
2
1
1
1
1
1
1
2
1
1
1
1
1
1
1
1
2
1
1
1
3
2
1
1
1
2
1
2
1
1
1
1
2
2
1
2
2
1
1
1
3
1
1
1
1
8
5
27
2
1
10
7
18
2
1
1
2
4
5
1
2
1
1
2
1
3
6
2
1
1
1 20
16 99 103
112 181 51
9
25
5
4 22
80 101
33
6
1
12 21
31 12
13
12
10
6
2
2
1
3
9
1
15
9
4
5
10
8
9
23
8
1
5
30
5
8
3
3
1
1
1
1
1
1
3
30
11
3
12
36
1
3
6
13
1
3
10
8
1
18
9
3
7
10
1
1
4
5
2
1
9
1
1
3
2
2
1
1
3
1
3
1
3
2
2
2
3
2
1
2
1
1
2
3
1
1
2
3
4
2
1
1
1
1
71
8
1
2
3
51
302
370
16
100
259
46
11
117
74
14
38
1
12
1
1
1
1
1
1
1
1
2
164 491 399 113 131 164
0
5
6
2
4
8
1
1
1
2
2
3
1
3
1
1
1
4
5
2
1
1
1
4
3
5
1
7
6
9
7
1
1
6
16
21
76
8
29
2
1
14
2
18
1
5
1
2
0
3
1
6
2
3
1
0
1
8
2
41
8
3
2
1
2
1
1626
46
Release Rectangles:
year:
<;juarter:
Rectangle
REC40F7
REC39F1
REC38Fl
REC38F2
REC37F2
REC37F4
REC37F6
REC36F1
REC36F2
REC36F3
REC36F4
REC36F5
REC36F6
REC35Fl
REC35F2
REC35F3
REC35F4
REC35F5
REC34F2
REC34F3
REC34F4
REC34F5
REC33F2
REC33F3
REC33F4
REC32F2
REC32F3
REC32F4
REC31Fl
REC31F2
REC31F3
REC30FO
REC30Fl
Total
No position
Not used:
APPENDIX I
0
2
REC36F5
3
4
1
1
REC35F5 (Nestern part of Dutch \'laddensea, 2nd quarter)
2
3
4
2
1
2
3
4
3
1
2
3
4
4
1
4+
Total
1
1
1
1
1
1
1
1
1
1
1
2
3
1
1
1
1
1
1
1
5
29
3
14
4
1
2
2
2
1
1
1
1
4
1
4
1
8
15
1
6
12
2
1
1
2
1
2
7
2
4
1
2
4
2
1
2
12
1
5
2
4
2
2
1
4
1
1
2
3
1
2
1
2
1
2
1
2
2
1
3
3
1
1
1
1
1
1
2
2
1
1
3
4
3
1
1
2
0
0
43
2
25
0
24
3
1
1
70
3
40
0
13
5
8
0
1
13
0
13
4
4
0
2
0
4
0
2
1
0
0
1
0
3
2
24
9
1
1
1
1
1
1
1
1
1
1
1
9
1
1
3
7
28
64
4
20
36
6
4
16
13
12
11
1
1
9
3
4
1
265
20
Release Rectangles:
year:
Quarter:
Rectangle
REC40F2
REC40F7
REC39F5
REC39F7
REC38Fl
REC38F2
REC38F7
REC37Fl
REC37F2
REC37F3
REC37F4
REC37F5
REC37F6
REC37F7
REC36FO
REC36F1
REC36F2
REC36F3
REC36F4
REC36F5
REC36F6
REC36F7
REC35F2
REC35F3
REC35F4
REC35F5
REC35F7
REC34F2
REC34F3
REC34F4
REC34F5
REC33F2
REC33F3
REC33F4
REC32Fl
REC32F2
REC32F3
REC31Fl
REC31F2
REC31F3
REC30E7
REC30FO
REC30Fl
REC29E6
REC29FO
REC29Fl
REC28E4
REC27E9
Total
No position
Not used:
APPENDIX I
0
3
REC35F5
REC34F5 (Western part of Dutch Waddensea, 3rd quarter)
1
4
1
2
2
3
4
1
3
2
3
4
1
2
3
4
4
4+
1
2
Total
1
1
1
1
1
1
1
1
1
1
3
1
1
1
1
1
1
25
13
1
1
2
2
3
3
2
1
3
1
5
7
1
33
14
1
3
3
2
1
3
25
2
1
3
1
1
1
1
1
2
3
15
3
1
2
6
29
7
47
1
4
7
10
68
1
1
4
1
1
3
6
97
91
2
10
19
1
4
1
1
4
43
12
7
25
7
13
9
4
5
1
1
2
1
4
2
1
1
4
5
1
2
3
1
1
1
25
4
1
15
9
1
6
2
3
1
1
1
1
1
1
2
2
1
1
1
1
1
8
1
1
1
7
1
3
1
1
1
1
1
1
1
2
1
1
2
2
1
2
2
1
1
1
2
1
1
1
2
1
1
1
1
1
1
1
1
1
1
1
1
1
63
0
1
81 107 299 141
3
2
4
5
50
6
18
4
99
4
32
3
12
1
8
3
7
0
7
1
4
0
1
2
3
0
8
1
22
10
1
1
1
1
2
1
1
1
1
3
1
1
1
1
1
1
4
4
6
48
5
4
5
14
91
25
2
5
32
215
263
5
48
81
2
3
27
3
14
1
2
4
2
1
1
2
1
1
940
39
•
Release Rectangles:
•
year:
Quarter:
Rectangle
REC45F2
REC44F9
REC41F5
REC40F2
REC40F6
REC40F7
REC39Fl
REC39F4
REC39F5
REC39F6
REC39F7
REC38F5
REC38F6
REC37FO
REC37Fl
REC37F2
REC37F3
REC37F4
REC37F5
REC37F6
REC37F7
REC37F8
REC36Fl
REC36F2
REC36F3
REC36F4
REC36F5
REC36F6
REC35Fl
REC35F2
REC35F3
REC35F4
REC35F5
REC35F6
REC35F7
REC34F1
REC34F2
REC34F3
REC34F4
REC34F5
REC33F2
REC33F3
REC33F4
REC32FO
REC32F2
REC32F3
REC31F1
REC31F2
REC31F3
REC31F4
REC30E8
REC30FO
REC30Fl
REC29E5
REC29E6
Total
No position
Not used:
APPENDIX I
0
4
REC35F5
1
1
2
3
REC34F5 (Western part of Dutch Waddensea, 4th quarter)
4
2
1
2
3
4
3
1
2
3
4
4
4+
2
Total
3
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
2
2
1
1
1
1
1
4
2
2
2
1
1
1
2
2
2
1
3
1
3
1
2
4
1
1
1
1
2
1
1
3
2
5
25
1
1
2
7
5
1
19
57
36
1
6
14
13
2
1
2
6
4
4
1
1
1
2
9 21
103 117
51
8
1
1
15 17
23 31
1
16
13
3
1
13
5
3
24
5
1
3
20
4
4
11
8
10
11
3
1
22
9
1
6
7
2
2
2
3
1
1
1
2
13
1
2
34
1
13
3
2
38
2
1
3
3
13 94
45 166
4
4
15
57
2
11
9
2
3
11
7
1
3
1
2
1
1
3
5
2
6
13
9
1
2
1
3
1
6
4
1
1
1
1
3
1
1
5
3
1
12
19
8
7
1
1
9
3
5
3
1
5
1
3
1
1
1
1
1
2
1
1
1
2
2
2
3
8
1
4
1
1
5
3
1
1
1
1
1
1
1
1
1
1
1
2
2
1
1
1
1
1
1
55 202 535 375 125 102 134
9 19
3 16
2
5
9
60
6
38
8
19
2
21
6
11
2
5
4
1
0
1
1
3
2
7
0
46
11
1
1
2
2
1
2
3
1
3
1
2
2
3
2
3
1
5
7
10
8
7
1
1
5
5
18
93
16
5
10
53
209
290
5
1
1
10
105
288
156
13
126
85
1
44
49
1
16
3
1
1
11
2
1
1
1694
94
REC34F5 (Dutch coast 1st quarter)
Release Rectangles:
year:
Quarter'
Rectangle
REC38Fl
REC37Fl
REC37F2
REC37F4
REC37F7
REC36FO
REC36F2
REC36F4
REC36F5
REC36F6
REC35Fl
REC35F2
REC35F3
REC35F4
REC35F5
REC34F2
REC34F3
REC34F4
REC34F5
REC33F2
REC33F3
REC33F4
REC32F2
REC32F3
REC31F2
REC31F4
Total
No position
Not used:
0
1
1
2
J
2
3
4
2
1
2
3
4
3
1
4+
2
3
4
Total
1
1
2
2
1
1
1
1
2
5
2
1
1
15
60
1
1
1
1
1
1
1
1
2
1
4
3
1
26
1
1
52
23
1
1
1
1
1
1
1
1
7
21
7
61
4
1
8
10
3
4
1
6
5
2
3
1
1
2
3
6
1
3
5
2
2
1
10 137 127
0
1
3
31
0
27
1
7
22
3
6
1
4
1
8
4
4
6
2
32
0
1
1
1
5
1
1
4
6
1
1
1
1
20
1
3
1
1
1
1
1
3
1
2
1
1
1
1
1
8
1
1
0
0
0
0
0
2
0
1
1
0
1
0
0
4
4
1
4+
3
2
20
138
30
4
36
47
4
16
3
1
401
11
0
0
2
year:
Quarter:
Rectangle
REC41E7
REC39Fl
REC39F2
REC38F5
REC38F7
REC37FO
REC37F2
REC37F4
REC36F3
REC36F4
REC36F5
REC35F2
REC35F3
REC35F4
REC34F3
REC34F4
REC34F5
REC33F2
REC33F3
REC33F4
REC32F2
REC32F3
REC31F2
REC30FO
REC29F1
REC2BE9
Total
No position
APPENDIX I
4
1
Release Rectangles:
Not used:
3
REC34F4
3
4
1
1
REC33F4
2
3
(Dutch coast 2nd quarter)
4
2
1
2
3
4
3
1
2
Total
1
1
1
1
1
1
1
1
4
1
14
1
4
1
1
2
1
11
2
14
1
1
5
1
4
7
1
1
10
1
1
1
1
1
1
1
3
1
16
3
2
3
2
3
3
3
1
5
4
1
1
1
1
3
1
1
2
2
4
2
1
2
3
1
1
1
1
1
1
17
30
1
2
1
1
2
1
1
1
24
1
43
0
20
2
2
1
36
2
1
1
25
2
3
11
11
0
0
0
0
0
4
1
1
1
1
1
1
1
1
1
2
4
3
7
9
26
13
61
1
1
1
4
1
2
5
0
1
2
1
1
1
192
0
0
0
0
0
0
7
37
12
•
Release Rectangles:
•
year:
quarterRectangle
REC40F7
REC39F7
REC38F1
REC38F2
REC38F7
REC37F2
REC37F4
REC37F5
REC37F7
REC36FO
REC36F2
REC36F3
REC36F4
REC36F5
REC36F6
REC36F7
REC35F2
REC35F3
REC35F4
REC35F5
REC35F7
REC34F2
REC34F3
REC34F4
REC34F5
REC33F2
REC33F3
REC33F4
REC32Fl
REC32F2
REC32F3
REC31F2
REC31F3
REC30E7
REC30FO
REC30Fl
REC29E6
REC29FO
REC29F1
REC28F1
REC27E9
Total
No position
Not used:
APPENDIX I
0
3
REC34F4
4
1
1
2
REC34F5
3
4
REC33F4
(Dutch coast 3rd quarter)
2
1
3
2
4
3
1
2
3
4
4
4+
1
Total
2
1
1
1
1
1
1
1
1
1
2
1
1
1
1
3
1
1
2
1
1
1
16
2
1
6
35
1
2
1
2
4
1
4
7
47
2
13
3
3
5
5
76
74
1
11
29
5
2
6
3
1
1
3
1
1
1
12
1
1
4
33
5
4
18
1
2
1
1
2
1
11
2
2
1
2
6
11
8
6
5
1
3
2
1
1
1
2
1
1
1
1
1
1
1
1
1
2
2
1
1
1
1
2
1
1
3
23
3
1
11
9
1
1
5
4
3
1
1
1
6
1
1
1
1
1
1
1
1
1
1
2
1
2
2
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
20
0
49
1
1
80 235
0
1
91
2
47
4
1
1
15
3
81
0
4
3
5
4
5
8
48
8
2
5
30
168
191
3
46
80
1
3
26
11
1
1
1
1
1
2
1
1
1
1
1
3
1
1
22
1
7
2
6
0
6
0
8
1
3
0
1
0
3
0
7
1
7
13
1
4
2
1
1
2
1
1
681
16
REC34F5 (Dutch coast 4th quarter)
Release Rectangles:
year:
QuarterRectangle
REC37FO
REC37F1
REC37F2
REC36F4
REC36F5
REC35Fl
REC35F2
REC35F3
REC35F4
REC35F5
REC34F1
REC34F2
REC34F3
REC34F4
REC34F5
REC33F2
REC33F3
REC33F4
REC32F2
REC32F3
REC31F2
REC31F4
REC29E6
Total
No position
Not used:
1
1
0
4
2
3
4
3
4
3
1
2
3
4
4
4+
1
2
Total
3
1
1
1
3
3
2
1
10
65
1
1
1
1
2
1
1
2
1
29
4
14
2
7
52
7
73
6
2
5
1
2
2
10
4
21
3
1
5
4
1
1
2
74 162 140
1
2
2
21
4
1
1
33
0
1
1
4
1
1
33
4
1
1
2
55
47
9
2
3
1
1
1
3
1
1
1
2
3
1
1
1
13
1
2
7
2
2
3
1
1
1
1
3
1
11
1
8
1
1
1
1
1
2
1
12
0
1
20
0
2
0
2
0
1
0
0
0
0
0
0
0
1
0
4
3
1
2
3
4
0
0
1
1
13
157
136
1
16
39
4
14
3
1
1
487
11
37
year:
quarter:
Rectangle
REC42F3
REC40F7
REC39Fl
REC38F4
REC37FO
REC37F2
REC37F3
REC36Fl
REC36F2
REC36F3
REC36F7
REC35F2
REC35F3
REC35F4
REC34F2
REC34F3
REC34F4
REC33F2
REC33F3
REC33F4
REC32F2
REC32F3
REC32F4
REC31Fl
REC31F2
REC31F3
REC30F1
REC29E5
0
1
REC32F4
(Scheldt Estuary, 1st quarter)
1
2
3
4
1
2
2
3
4
1
2
3
4+
Total
1
1
1
1
1
2
1
1
1
2
1
1
1
1
2
3
3
2
9
6
4
23
8
17
142
36
2
37
11
2
1
1
1
1
1
1
1
1
1
1
1
2
1
1
1
4
13
2
1
1
2
3
3
2
2
41
13
11
3
5
58
4
2
1
1
7
17
4
7
4
5
2
1
1
1
1
1
1
3
1
1
1
1
1
2
3
3
2
1
14
5
1
2
1
1
1
2
1
13
1
1
1
2
1
1
3
2
1
1
1
1
1
1
1
REC29E6
APPENDIX I
2
1
Release Rectangles:
REC29Fl
REC28FO
Total
No position
Not used:
2
1
1
2
1
1
1
1
19
0
84 102
3
5
20
2
13
3
36
1
26
1
10
0
2
0
9
0
1
0
1
1
0
0
2
0
0
0
0
0
1
0
2
14
2
1
326
16
•
Release Rectangles:
•
year:
quarter:
Rectangle
REC42F6
REC38Fl
REC37Fl
REC37F4
REC35F2
REC35F3
REC35F4
REC34F2
REC34F3
REC33F3
REC33F4
REC32F2
REC32F3
REC31F2
REC31F3
REC30E6
REC30F1
REC29E5
REC28E9
Total
No position
Not used:
0
2
REC32F4
3
2
3
4
2
1
2
3
3
1
4
2
3
4
4
1
4+
Total
1
1
1
1
1
1
1
2
1
1
2
2
3
1
37
3
1
4
1
1
1
2
2
1
3
1
8
1
4
1
1
1
1
1
1
2
2
1
1
1
1
1
0
0
45
0
10
0
1
9
1
12
0
11
2
4
2
5
0
3
0
2
0
2
0
0
0
1
1
1
0
0
0
0
0
2
3
4
4
1
2
1
2
1
1
1
3
1
3
1
1
7
12
1
6
56
6
2
1
1
1
1
106
8
4
0
3
year:
quarter:
Rectangle
REC44E8
REC40F2
REC40F6
REC38F2
REC37F2
REC35FO
REC35F2
REC35F3
REC35F4
REC34F2
REC34F3
REC34F4
REC33F2
REC33F3
REC33F4
REC32F1
REC32F2
REC32F3
REC32F4
REC31E7
REC31Fl
REC31F2
REC31F3
REC30FO
REC30Fl
REC29E5
REC29E6
REC29Fl
REC28E4
REC28E5
REC28E9
REC27E9
Total
No position
APPENDIX I
1
1
1
REC32F4 (Scheldt Estuary, 3rd quarter)
Release Rectangles:
Not used:
4
(Scheldt Estuary, 2nd quarter)
4
1
1
2
3
4
2
1
2
3
4
3
1
4+
Total
1
29
15
56
27
15
6
14
3
4
2
1
1
6
1
1
1
1
1
1
4
4
1
9
13
6
5
60
15
2
28
297
95
1
2
43
14
7
5
2
2
4
1
1
3
1
631
1
1
2
2
2
0
2
2
1
0
0
1
18
1
1
1
1
1
1
2
1
1
3
3
20
2
30
1
10
5
6
1
1
18
1
1
4
8
9 103 130
27 10
4
1
1
6 11 12
7
1
4
1
1
2
1
1
1
1
1
2
1
2
2
7
2
12
1
4
3
2
3
1
1
1
1
10
3
1
4
15
1
4
1
3
1
4
1
1
8
1
1
1
1
1
1
1
5
1
3
4
2
1
3
1
2
5
1
1
1
1
6
2
2
1
2
1
2
1
2
1
1
1
1
1
1
1
1
2
24
35
0
0
46 152 196
0
1
3
1
19
15
Release Rectangles:
year:
<Juarter:
Rectangle
REC44E7
REC40F2
REC39F2
REC39F6
REC38F4
REC38F6
REC37F1
REC37F2
REC37F7
REC36FO
REC36F3
REC36FS
REC3SF1
REC3SF2
REC35F3
REC35F4
REC34F2
REC34F3
REC34F4
REC33F2
REC33F3
REC33F4
REC32F1
REC32F2
REC32F3
REC32F4
REC31F1
REC31F2
REC31F3
REC30FO
REC30Fl
REC29E5
REC29E6
REC28E9
REC28FO
REC27E9
Total
No position
Not used:
0
4
REC32F3
1
1
4
2
1
(Scheldt Estuary, 4th quarter)
2
4
3
3
1
2
3
4
4
4+
1
2
3
Total
0
0
1
1
1
1
2
1
1
1
1
1
1
2
1
1
10
4
8
20
4
7
65
12
1
20
335
30
2
43
5
8
2
1
5
1
2
2
603
33
1
1
1
1
1
1
1
1
1
1
1
2
1
3
1
1
1
1
4
2
6
12
6
3
3
11
2
1
25
3
1
1
1
1
1
1
2
1
14
5
2
1
1
4
5
3
2
4
95 146
13
4
30
1
5
4
32
1
11
3
12
7
4
2
1
1
5
1
1
1
1
3
2
3
1
1
2
11
5
1
1
1
1
1
1
2
2
2
3
4
1
2
1
3
1
3
1
2
1
2
1
1
1
1
1
5
0
25 135 205
0 12
1
70
3
33
1
1
1
56
4
]
32
4
16
6
10
0
7
1
2
0
1
0
1
0
4
0
1
1
0
year:
<Juarter:
Rectangle
REC33F3
REC32F1
REC32F3
REC31Fl
REC31F2
REC30Fl
REC29Fl
Total
No position
APPENDIX I
3
REC31F3
1
Release Rectangles:
Not used:
2
REC32F4
0
2
REC31F1
3
4
1
1
REC31F2
2
3
(North Sea French/Be1gian border 2nd quarter)
4
2
1
2
3
4
3
1
2
3
4
4
1
4+
Total
1
0
8
3
2
1
1
1
1
1
1
6
0
0
0
0
0
8
3
2
2
0
0
1
0
0
0
0
0
0
0
0
1
1
1
1
14
3
1
22
0
0
0
0
0
0
0
0
0
0
0
0
0
2
6
16
•
Release Rectangles:
•
0
3
year:
quarter:
Rectangle
REC37F1
REC37F3
REC35F2
REC34F2
REC33F3
REC32F2
REC32F3
REC31Fl
REC31F2
REC30FO
REC30Fl
REC29E5
REC29E6
REC29F1
REC28E9
REC28FO
REC27E9
Total
No position
Not used:
REC31Fl
4
APPENDIX I
2
3
4
(North Sea French/Belgian border 3rd quarter)
2
1
2
3
4
3
1
2
3
4
4
4+
1
Total
2
1
1
1
1
1
1
1
2
15
41
1
75
4
5
8
1
28
4
1
7
1
4
2
1
2
1
1
2
1
3
1
1
1
1
1
1
2
1
1
1
1
61
0
82
0
9
0
35
0
1
1
9
0
11
0
5
0
6
0
7
0
0
0
0
0
0
0
1
0
1
0
1
0
1
0
2
0
1
1
1
1
2
1
2
19
171
1
18
1
1
6
2
2
1
231
0
30
Release Rectang1es:
year:
quarter"
Rectangle
REC45E7
REC45F2
REC44E8
REC44E9
REC44Fl
REC43E8
REC42E7
REC42E8
REC42E9
REC41E7
REC41E8
REC41E9
REC41FO
REC41Fl
REC41F2
REC40E7
REC40E8
REC40E9
REC40FO
REC40Fl
REC40F2
REC40F4
REC40F5
REC40F6
REC39E8
REC39E9
REC39FO
REC39Fl
REC39F2
REC39F3
REC39F5
REC38E8
REC38E9
REC38FO
1
1
REC31F2
0
2
REC33F1
3
4
1
1
REC32Fl
2
3
REC31FO
4
2
1
REC31F1
2
3
(Thames Estuary 2nd quarter)
4
3
1
4
2
3
4
4+
Total
1
1
1
1
1
1
1
1
3
1
1
2
2
1
1
2
1
1
1
1
1
1
1
1
1
1
1
1
1
1
3
1
1
1
1
1
2
1
1
1
1
1
1
1
2
2
1
1
2
1
1
1
1
1
1
1
2
2
1
1
1
1
1
2
4
1
1
1
1
3
1
1
1
1
1
1
17
1
2
1
1
1
5
1
4
2
2
4
2
1
1
1
2
1
7
1
2
5
2
1
1
5
2
6
3
5
2
1
1
14
4
Release Rectangles:
CONTINUED
year:
Q;uarter:
Rectangle
0
2
REC38Fl
REC38F2
REC38F4
REC37E9
REC37FO
REC37Fl
REC37F2
REC37F3
REC37F6
REC36FO
REC36F1
REC36F2
REC36F5
REC36F6
REC35FO
REC35F1
REC35F2
REC35F3
REC35F4
REC34F1
REC34F2
REC34F3
REC34F4
REC33FO
REC33F1
REC33F2
REC33F3
REC33F4
REC32FO
REC32Fl
REC32F2
REC32F3
REC31E5
REC31FO
REC31Fl
REC31F2
REC30E4
REC30E9
REC30FO
REC30Fl
REC29E5
REC29E6
REC29Fl
REC28E5
REC28E6
REC28E9
REC33Fl
3
2
4
1
1
1
1
1
REC32F1
REC31FO
2
1
2
3
4
1
2
1
1
1
1
1
3
1
1
3
4
2
1
3
1
2
1
2
4
3
16
8
3
2
1
1
REC31F1
2
3
(Thames Estuary 2nd quarter)
4
3
1
2
3
4
4
1
1
4+
Total
2
1
1
1
2
1
1
2
1
1
3
1
3
1
1
1
5
1
3
2
3
1
1
1
1
2
1
1
1
1
1
2
7
1
4
19
52 266
2
1
1
1
2
2
2
1
1
2
8
1
8
1
6
2
1
25
1
1
15
3
1
2
1
1
1
1
4
1
1
1
1
1
2
1
1
4
1
2
1
1
19
3
2
1
2
1
1
1
1
1
13
2
28
2
1
3
2
4
1
24
2
13
4
24
11
2
1
1
6
1
1
3
1
1
36
4
91
20
1
14
1
1
8
1
2
2
9
1
1
2
1
5
2
54
9
18
5
1
13
1
1
1
29
5
2
10
3
1
1
3
4
2
1
1
1
1
2
1
2
1
1
1
1
7
3
1
4
39
20
7
2
1
14
13
8
2
3
2
2
19
8
1
7
51
8
2
1
383
10
4
1
4
81
23
1
1
9
212
3
1
1
146
30
3
7
3
1
2
1
--------------------------------------------------------------------------------------Total
No position
Not used:
APPENDIX I
110 491
0 10
94
11
93 155
34 33
96
25
58
9
24
19
35
16
16
9
10
6
9
11
11
4
9
6
6
3
3
2
27
10
73
18
1247
208
•
REC35FO (Wash, 2nd quarter)
Release Rectangles:
0
2
year:
quarter:
Rectangle
REC44E8
REC42E7
REC42E8
REC42E9
REC4lE7
REC41E8
REC41E9
REC40E7
REC40E8
REC39E8
REC39F5
REC38E8
REC38E9
REC38FO
REC37E9
REC37FO
REC36FO
REC36F1
REC36F2
REC35FO
REC35Fl
REC33F2
REC30E8
Total
No position
Not used:
3
2
3
4
2
1
2
3
4
3
1
2
3
4
4
1
4+
Total
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
2
2
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
2
1
1
2
0
0
0
1
0
3
0
4
2
6
3
4
3
1
3
5
2
4
1
1
3
0
2
3
4
1
3
0
0
2
1
2
3
4
4
1
2
1
9
5
1
2
1
1
2
4
1
1
2
2
1
3
6
4
2
4
1
2
1
2
1
1
1
46
32
1
0
3
year:
QuarterRectangle
REC44E6
REC44E7
REC43E8
REC42E8
REC41E7
REC41E8
REC40E8
REC40E9
REC39E8
REC39FO
REC39F1
REC39F3
REC38E8
REC38E9
REC37E9
REC37FO
REC37Fl
REC37F2
REC36FO
REC36F1
REC36F2
REC35F2
REC33F2
REC32Fl
Total
No position
APPENDIX I
1
1
1
Release Rectangles:
Not used:
4
REC36FO
4
1
1
2
REC35FO (Wash, 3rd quarter)
3
4
2
1
2
3
4
3
1
4+
Total
1
1
1
1
2
1
2
2
1
1
1
1
1
2
1
2
1
1
1
1
1
3
1
2
1
2
1
1
1
2
1
1
2
2
1
1
1
1
1
2
2
1
2
1
1
2
1
1
1
1
1
1
1
1
1
1
1
2
1
1
1
1
1
1
1
1
1
1
1
1
2
0
1
0
3
0
4
0
1
12
2
8
0
10
1
9
0
7
3
3
4
2
4
3
1
4
3
1
2
4
1
1
1
14
15
1
19
1
1
1
1
9
3
3
2
11
1
2
1
6
10
16
5
2
1
6
2
1
1
1
1
88
37
Release Rectangles:
0
2
year:
quarterRectangle
REC41E9
REC40E9
REC38E9
REC38Fl
REC38F2
REC37FO
REC36F3
REC36F4
REC35FO
REC34F2
REC34F3
REC33Fl
REC33F2
REC33F3
REC32Fl
REC32F2
REC31Fl
REC31F2
REC30E9
REC30FO
REC30Fl
REC29E5
REC29E7
REC28E9
Total
No position
Not used:
REC30FO
1
3
2
3
4
2
1
3
2
3
4
]
2
3
4
4
1
4+
Total
1
1
1
1
1
1
1
1
1
1
1
1
1
1
1
2
1
3
75
2
2
97
2
3
3
1
1
34
1
1
2
1
21
1
1
3
2
46
1
2
22
4
3
1
3
2
5
1
3
2
1
6
0
9
2
1
0
3
1
3
1
1
1
0
1
1
0
3
4
1
2
3
4
4
1
2
1
1
1
2
0
1
0
5
1
1
]
79 106
0
0
44
0
30
8
54
2
25
0
5
4
5
2
8
5
1
1
1
1
3
2
1
1
1
1
1
1
1
1
2
5
15
1
9
322
5
2
1
1
380
27
14
REC30FO (Rye, 3rd quarter)
0
3
4
1
1
4
2
5
4
0
2
0
5
0
2
2
4
1
5
0
3
4
]
1
2
3
2
4+
Total
1
2
1
1
3
0
0
0
1
0
2
0
0
0
0
0
1
0
0
0
0
0
0
0
0
0
3
1
3
4
4
1
4+
2
22
2
1
26
0
0
Release Rectangles:
0
2
year:
quarter:
Rectangle
REC30E6
REC30E8
REC30E9
REC30FO
REC29E5
REC29E6
REC29E7
REC28E6
Total
No position
APPENDIX I
]
1
year:
quarter"
Rectangle
REC31F1
REC30FO
REC30Fl
REC29Fl
Total
No position
Not used:
4
1
Release Rectangles:
Not used:
REC30Fl (Rye, 2nd quarter)
22
REC30E8
REC30E9 (PooIe, 2nd quarter
1
3
4
]
48
3
4
6
10
2
3
4
2
1
2
23
3
4
7
1
7
2
3
3
4
Total
1
1
2
1
1
1
1
1
22
0
55
0
6
2
11
0
30
2
7
0
9
0
1
3
2
4
0
0
0
1
0
1
0
1
0
0
0
0
0
0
0
2
0
3
20
131
6
8
1
3
1
1
152
6
•
Release Rectangles:
year:
quarter:
Rectangle
REC28E3
REC28E7
REC28E8
REC28E9
REC27E8
REC27E9
Total
No position
Not used:
0
2
REC28E8
3
4
1
1
1
6
0
1
3
4
2
1
2
3
4
3
1
2
3
4
4
1
4+
Total
1
1
0
0
2
0
1
1
0
0
1
0
0
0
0
0
1
0
1
0
0
0
1
0
0
0
0
0
1
1
0
1
0
0
0
1
0
1
1
6
2
3
2
15
0
1
0
4
REC28E8
1
1
2
3
REC27E8
4
2
1
REC27E9
2
3
(Bay de Seine, 4th quarter)
4
3
1
2
3
4
4
4+
1
2
3
Total
1
1
2
2
1
2
2
1
5
0
1
1
2
1
3
2
6
9
16
8
0
28
0
3
8
1
17
0
2
1
1
1
1
1
1
3
1
2
2
2
1
4
0
3
0
6
0
3
0
2
0
1
0
1
0
1
1
1
0
1
0
0
0
0
0
0
0
4
4
1
2
3
0
1
1
9
5
1
16
45
2
83
0
4
Release Rectangles:
year:
quarter:
Rectangle
REC30FO
REC30Fl
REC29E5
REC29E6
REC29Fl
REC28E4
REC28E9
REC28FO
REC28Fl
REC27E9
BEC26E7
Total
No position
APPENDIX I
2
1
year:
quarter:
Rectangle
REC29E5
REC29E6
REC29E8
REC28E6
REC28E8
REC28E9
REC27E3
REC27E8
REC27E9
REC27FO
Total
No position
Not used:
1
1
(Bay de Seine, 2nd quarter)
1
Release Rectangles:
Not used:
4
REC27E8
0
3
REC29Fl
4
1
1
1
REC28Fl
2
3
1
1
4
(Bay de Somme, 3rd quarter)
2
1
2
1
1
3
4
3
1
2
3
4+
Total
1
1
6
7
3
7
10
1
2
1
1
2
2
1
1
1
1
0
1
0
1
7
0
7
0
3
0
9
0
11
0
1
0
2
0
4
0
0
0
2
0
0
0
0
0
2
0
0
0
3
0
1
21
2
3
1
1
37
1
2
2
1
2
1
53
0
REC29Fl (ßay de Somme, 4th quarter)
Release Rectangles:
0
4
year:
quarter:
Rectangle
REC31EO
REC30E9
REC30FO
REC30F1
REC29E5
REC29E6
REC29E7
REC29FO
REC29Fl
REC28E3
REC28E5
REC28E6
REC28E8
REC28E9
REC28FO
REC28F1
REC27E8
REC27E9
Total
No position
Not used:
1
1
1
1
2
1
2
3
4
3
1
2
3
4
4
4+
1
2
Total
3
4
1
1
2
1
2
2
2
1
9
7
1
21
6
1
1
1
1
4
1
1
1
1
2
1
1
1
1
1
1
3
2
1
1
1
1
5
32
0
1
12
0
3
1
3
2
1
1
10
0
2
9
0
4
0
6
0
15
0
1
8
0
1
1
1
9
2
5
2
4
55
2
1
7
0
1
0
3
0
1
0
1
0
1
1
0
0
0
2
0
3
0
1
1
1
5
7
4
1
13
115
0
3
REC26E7 (Bay du St. Michel, 2nd quarter)
0
2
4
1
1
2
3
4
2
1
2
3
4
3
1
2
3
4
4
1
4+
3
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
1
1
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
2
3
Total
2
2
0
0
Release Rectangles:
year:
quarter:
Rectangle
REC30E6
REC29E5
REC29E6
REC28E6
REC28FO
REC27E8
REC26E8
Total
No position
APPENDIX I
4
1
year:
quarter:
Rectangle
REC26E7
Total
No position
Not used:
3
1
Release Rectangles:
Not used:
2
0
4
REC28E8
1
1
2
3
REC26E8 (Bay du St. Michel, 4th quarter)
4
2
1
2
1
1
1
3
4
3
1
2
3
4
4
4+
1
Total
1
1
1
1
0
0
1
1
0
1
18
20
0
2
0
0
0
1
0
3
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
0
22
1
2
2
1
1
1
19
27
0
APPENDIX II
Summary of Tagging Data of plaice in the length range <25.0 cm
release area:
release perlod:
source:
e
No. of
Qqa rter returns
0 3
1
0 4
1
1 1
4
8
1 2
3
1 3
1
1 4
8
2 1
2 2
12
2 3
9
2 4
3
3 1
1
3 2
9
3 3
2
3 4
1
release area:
release period:
source:
No. of
Quarter returns
0 3
7
0 4
3
1 1
12
74
1 2
16
1 3
12
1 4
2 1
12
2 2
43
2 3
19
2 4
14
7
3 1
3 2
12
3 3
11
3 4
2
release area:
release period:
source:
No. of
Quarter returns
9
0 3
0 4
3
1 1
2
1 2
11
1 3
5
2 4
3
2 1
1
4
2 2
2 3
3
2 4
1
3 1
1
APPENDIX II
Hvide Sande
2nd quarter
Netherlands
D
11.9
5.1
84.0
66.8
64.6
5.6
98.2
71.1
48.7
71.8
98.2
95.1
159.3
199.5
'f
294.8
192.2
235.0
268.6
236.6
270.0
258.8
248.8
269.8
242.4
284.8
234.2
234.9
246.4
aA 2
0.0
0.0
1118.6
512.8
387.8
0.0
279.8
218.4
78.6
156.1
0.0
378.5
211. 7
0.0
V
11. 9
1.3
8.1
4.6
1.4
0.3
3.7
2.0
1.6
2.3
3.1
1.5
3.7
4.8
t
1.0
4.0
8.8
11.0
14.0
17.0
20.8
23.1
25.9
28.7
32.0
35.1
37.5
42.0
V*t
11. 9
5.1
70.6
51. 0
19.4
5.6
76.0
46.0
42.1
64.8
98.2
51.1
139.0
199.5
t*a A 2
0.0
0.0
9787.6
5641. 3
5429.4
0.0
5805.6
5041.2
2034.4
4474.5
0.0
13288.7
7937.9
0.0
aA 2
1425.4
239.7
297.7
327.9
293.4
236.0
679.1
304.8
117.7
189.3
360.1
239.3
113.2
141. 4
V
10.9
8.4
4.4
1.1
1.5
2.6
3.3
1.9
1.8
1.4
4.0
1.4
1.5
2.1
t
1.1
3.0
7.8
9.9
12.8
15.9
19.5
21.8
25.1
27.5
30.7
33.8
37.5
39.5
V*t
12.4
25.1
34.4
10.9
19.0
41. 6
64.7
42.3
45.1
39.5
121. 4
45.7
54.4
84.0
t*a A 2
1629.0
719.0
2332.1
3257.0
3759.5
3756.7
13243.1
6655.4
2954.1
5204.7
11061.3
8077.0
4249.7
5583.6
aA 2
81.1
333.3
135.6
339.8
363.3
239.6
0.0
219.9
33.1
0.0
0.0
V
7.6
10.6
4.1
2.3
2.4
11.8
6.7
3.3
6.1
5.4
3.5
t
2.1
4.7
8.0
11. 5
13.6
16.3
19.0
23.3
26.3
28.0
33.0
V*t
16.0
49.6
32.7
26.7
32.2
193.4
127.1
77.1
161.7
152.5
114.8
t*a A 2
171.1
1555.3
1084.9
3923.4
4941. 2
3913.4
0.0
5112.2
871. 9
0.0
0.0
Hvide Sande
3rd quarter
Netherlands
D
'J1
28.8
32.4
48.0
31. 6
44.5
62.3
101. 8
65.5
58.7
65.9
130.7
81. 6
64.8
102.8
30.1
338.3
244.8
332.8
264.4
241.7
234.9
274.9
253.6
226.8
221.5
284.4
282.0
241. 6
Sylt
2nd quarter
Netherlands
D
'J1
20.1
51.8
43.4
50.5
70.1
201. 9
127.1
79.9
162.6
152.5
114.8
163.2
223.4
261.2
288.2
268.6
248.1
260.9
223.4
236.2
45.4
275.0
1
release area:
release period:
source:
No. of
Quarter returns
0 3
2
1 1
1
7
1 2
1 3
5
1 4
4
2 1
2
2 2
10
2 3
2
2 4
2
3 1
5
3 2
7
7
3 3
4 1
3
4+
7
Sylt
3rd quarter
Netherlands
D
24.6
109.1
53.5
49.5
79.9
54.5
57.0
53.0
44.9
58.4
87.2
143.5
124.0
104.5
'P
324.4
8.2
235.9
256.0
271. 3
302.4
264.5
284.3
271. 4
242.0
292.3
248.8
236.7
275.0
release area:
release period:
source:
Sylt
4th quarter
Netherlands
No. of
Quarter returns
1 2
1
1 3
1
2 2
2
3 4
1
4 2
1
D
67.7
28.6
99.2
65.7
38.7
release area:
release period:
source:
Norderney
3rd quarter
Netherlands
No. of
Quarter returns
0 3
2
1 2
1
release area:
release period:
source:
No. of
Quarter returns
0 4
1
7
1 1
1 2
6
7
1 3
2
1 4
2 1
4
12
2 2
2 3
6
2 4
3
3 1
3
4
3 2
3 3
5
4 1
2
4 2
4
4 3
2
4+
3
APPENDIX I I
D
44.0
158.9
'P
8.2
168.3
324.1
315.7
342.8
'P
290.4
259.1
a"2
369.3
0.0
266.7
209.0
385.3
15.6
107.7
14.0
20.0
3.8
151. 6
260.5
223.9
122.1
V
23.3
15.6
4.7
3.7
2.9
2.8
2.0
2.2
1.6
1.9
2.4
3.3
2.9
1.5
t
1.0
7.0
8.7
11. 6
15.0
18.5
21.5
24.0
26.5
30.2
32.6
36.1
41.3
59.3
V*t
23.3
109.1
41.0
43.0
43.8
52.3
43.3
52.7
42.8
58.0
77 .1
120.7
118.4
87.7
t*a"2
369.3
0.0
2323.8
2424.1
5779.8
289.0
2316.0
336.1
531.1
114.9
4936.4
9416.4
9255.9
7241. 5
•
a"2
0.0
0.0
195.0
0.0
0.0
a"2
719.5
0.0
t
8.0
11.0
18.5
37.0
44.0
V*t
67.7
28.6
97.5
65.7
38.7
t*a"2
0.0
0.0
3606.7
0.0
0.0
t
V
21.6 2.0
15.9 10.0
V*t
43.1
158.9
t*a"2
1439.0
0.0
t
1.0
4.4
6.5
10.0
12.0
15.5
19.2
21.8
24.7
28.0
31. 3
34.4
39.5
43.0
45.5
57.0
V*t
9.6
180.7
114.3
64.5
102.1
155.8
64.2
63.6
95.6
111.7
105.0
58.9
184.2
67.8
88.7
150.5
t*a"2
0.0
9291.3
5588.2
7916.3
8193.0
5845.5
8229.3
6535.5
1367.7
2951.5
17484.3
3585.1
3271.4
9801.5
2285.3
19818.3
V
8.5
2.6
5.3
1.8
0.9
Norderney
4th quarter
Netherlands
D
9.6
184.5
126.4
96.1
104.8
159.0
78.0
70.9
96.5
118.6
131.2
62.4
189.6
86.6
95.6
174.3
'P
58.6
239.1
240.7
290.2
273.4
255.1
259.0
267.1
244.6
264.5
288.0
305.7
258.5
278.0
259.6
260.4
a"2
0.0
2098.0
859.7
791. 6
682.8
377 .1
429.4
299.3
55.4
105.4
559.5
104.2
82.8
227.9
50.2
347.7
V
9.6
40.8
17.6
6.4
8.5
10.1
3.4
2.9
3.9
4.0
3.4
1.7
4.7
1.6
1.9
2.6
2
release area:
release period:
source:
No. of
Quarter returns
0 3
222
48
0 4
1 1
36
1 2
224
181
1 3
1 4
45
72
2 1
117
2 2
94
2 3
34
2 4
18
3 1
24
3 2
3 3
16
3 4
6
4 1
8
4 2
9
4+
24
release area:
release period:
source:
No. of
Quarter returns
0 4
20
1 1
31
1 2
71
120
1 3
1 4
41
32
2 1
49
2 2
33
2 3
19
2 4
10
3 1
3 2
14
3 3
5
4 1
4
4 2
2
4 3
3
4+
8
release area:
release period:
source:
No. of
Quarter returns
164
0 1
0 2
491
0 3
399
113
0 4
1 1
131
164
1 2
76
1 3
1 4
29
2 1
14
18
2 2
2 3
5
2 4
2
3 1
3
3 2
6
3 3
3
4+
8
APPENDIX I r
eastern part of Dutch Waddensea
3rd quarter
Netherlands
D
4.9
14.8
69.7
49.1
60.3
84.5
137.8
81.2
78.8
129.9
130.4
107.3
131. 3
104.7
132.9
142.9
157.3
'P
324.2
305.1
257.6
273.6
292.2
268.4
257.6
275.3
304.7
286.5
280.1
303.2
294.9
255.7
273.0
270.7
288.6
a"2
45.5
149.1
997.0
566.6
235.5
286.1
475.4
223.1
145.2
293.4.
224.8
320.8
224.2
115.8
61.1
113.1
113.3
V
3.0
3.7
9.7
3.5
4.0
4.9
6.3
3.0
2.6
4.0
3.6
2.4
3.3
2.4
3.0
3.0
2.2
t
1.0
2.7
6.0
8.8
11. 9
14.7
18.1
21.2
23.7
26.6
30.1
33.2
36.1
39.3
41. 5
44.8
65.9
V*t
3.0
10.2
58.6
30.6
47.3
71. 6
114.1
63.0
62.4
106.1
106.9
78.2
118.6
94.3
124.4
134.0
145.3
t*a"2
45.5
406.8
6010.0
4963.2
2801.6
4203.2
8610.6
4719.0
3443.5
7801. 0
6756.2
10652.2
8100.5
4554.4
2537.3
5062.4
7464.8
eastern part of Dutch Waddensea
4th quarter
Netherlands
D
6.8
105.2
88.7
84.9
92.5
133.2
124.7
95.8
140.2
144.2
147.2
186.8
188.0
83.5
152.7
167.8
'P
283.8
250.4
250.3
274.4
272.5
255.2
260.3
294.5
287.5
261.0
280.1
299.1
291. 9
330.5
295.5
270.0
a"2
185.7
2277.5
774.3
550.1
336.2
248.6
482.9
283.3
382.8
180.0
373.5
282.1
715.8
64.1
53.4
154.2
V
4.0
21.7
11. 5
6.3
5.9
7.6
5.1
3.1
4.6
4.8
3.7
5.1
2.4
1.6
3.2
2.3
t
1.3
4.0
7.0
9.9
13.0
16.1
18.9
21.8
24.3
27.6
30.9
33.2
40.0
43.0
46.0
62.1
V*t
5.0
86.8
80.7
62.3
76.8
122.6
97.1
67.7
111.1
133.8
115.4
170.4
96.7
68.9
148.9
144.5
t*a"2
232.1
9110.1
5419.9
5418.4
4361.8
3992.9
9107.0
6181. 2
9308.0
4967.6
11523.7
9365.5
28633.6
2756.5
2458.2
9582.0
a"2
506.9
338.2
257.9
226.6
268.5
236.5
185.1
83.5
222.0
340.4
547.7
180.3
57.6
185.1
69.9
128.6
V
3.8
4.0
4.1
4.4
5.8
2.9
2.4
3.2
3.2
3.0
4.7
4.0
2.4
2.9
4.0
1.7
t
1.5
3.7
6.6
9.2
13.0
15.7
18.5
21.5
25.2
28.0
30.6
33.5
37.0
39.5
42.3
63.1
V*t
5.9
14.8
27.2
40.6
75.4
45.4
45.1
68.7
79.5
84.6
144.6
134.5
89.8
114.4
167.9
108.6
t*a"2
785.1
1258.3
1702.9
2083.6
3482.4
3719.1
3420.1
1793.1
5596.6
9530.6
16759.8
6039.6
2130.7
7310.7
2960.4
8120.1
Western Waddensea
1st quarter
Netherlands
D
15.8
24.5
40.1
54.5
83.0
63.0
66.1
74.3
93.6
97.1
163.7
151. 9
97.4
131.4
176.2
128.6
'Jl
280.0
257.1
262.1
255.7
240.9
244.5
261.1
241. 6
237.8
242.5
319.1
255.8
268.7
236.6
310.0
292.7
3
release area:
release perlod:
source:
No. of
Quarter returns
43
0 3
0 4
25
24
1 1
70
1 2
1 3
40
13
1 4
8
2 1
2 2
13
2 3
13
2 4
4
3 1
2
4
3 2
2
3 3
4 1
1
4+
3
release area:
release perlod:
source:
No. of
Quarter returns
63
0 3
81
0 4
107
1 1
1 2
299
1 3
141
50
1 4
2 1
18
2 2
99
2 3
32
2 4
12
3 1
8
3 2
7
3 3
7
3 4
4
4 1
1
4 2
3
4+
8
release area:
release period:
source:
No. of
Quarter returns
0 4
55
1 1
202
1 2
535
1 3
375
1 4
125
2 1
102
2 2
134
2 3
60
2 4
38
3 1
19
3 2
21
3 3
11
3 4
5
4 1
1
4 2
1
4 3
3
4+
7
APPENDIX Ir
Western Waddensea
2nd quarter
Netherlands
D
16.0
28.3
92.2
62.2
72 .0
89.4
124.2
84.7
100.4
78.6
81. 3
91.7
86.9
60.4
89.5
\}I
277 .1
238.8
233.0
227.7
238.7
238.1
230.8
229.2
246.8
277.8
232.9
240.1
250.0
252.7
283.2
a~2
227.5
168.6
677.7
295.9
208.3
164.2
173.4
170.3
183.7
181. 0
19.6
123.5
33.5
0.0
91. 5
V
4.5
5.1
10.0
5.3
4.6
5.2
5.9
2.7
3.4
2.2
2.6
2.2
2.2
1.3
1.2
t
1.8
5.0
8.5
10.8
13.9
16.5
20.5
23.2
26.2
28.3
31. 0
35.0
37.5
45.0
66.0
V*t
8.0
25.4
85.2
57.5
64.1
86.1
121. 3
63.6
90.2
61.2
79.6
77.8
81. 6
60.4
82.0
402.0
842.9
5788.5
3195.3
2895.2
2715.8
3554.8
3942.3
4804.5
5112.6
607.2
4324.0
1255.9
0.0
6042.0
V
7.2
0.8
1.8
2.1
2.5
2.9
3.9
2.4
2.2
2.0
3.4
1.5
0.9
2.0
4.3
1.1
1.7
t
1.3
2.8
5.8
8.7
11.4
14.5
17.9
20.4
23.2
26.0
30.0
32.3
35.9
38.5
42.0
44.0
70.0
V*t
9.0
2.2
10.4
18.1
28.8
42.3
69.5
48.4
50.7
51.2
103.4
46.9
33.7
77.8
181.7
46.4
115.9
562.4
821. 3
1879.6
1970.6
1655.8
2975.4
3813.7
3602.8
5638.4
8467.9
6422.7
1080.1
5228.9
3152.4
0.0
1236.8
15459.5
V
·3.6
8.0
2.9
3.6
4.2
5.4
3.2
3.2
2.2
3.6
2.4
1.5
3.5
3.1
0.9
1.1
1.4
t
1.2
3.2
6.4
9.5
11. 9
15.5
18.1
21.0
24.0
27.4
30.1
33.7
36.0
38.0
41.0
45.7
61. 6
V*t
4.1
25.1
18.8
34.3
49.9
84.4
57.8
67.0
51. 9
99.6
71.2
50.2
126.5
117.2
35.9
48.5
88.2
520.5
2186.3
1373.2
1771.2
3093.8
4318.5
5035.9
4033.6
4040.7
10609.4
5806.6
5495.5
10471. 7
0.0
0.0
11059.7
22663.9
t*a~2
Western Waddensea
3rd quarter
Netherlands
D
19.1
21.0
30.7
31.7
39.2
52.8
80.7
60.6
73.3
78.3
104.6
49.8
68.7
92.9
181.7
47.2
131. 9
'f
66.2
95.9
204.7
234.9
243.9
233.4
236.1
236.1
248.1
267.6
230.4
228.9
286.3
238.6
308.9
223.0
254.0
a~2
448.5
298.3
321. 8
226.6
144.7
204.9
213.2
176.5
242.8
325.7
214.1
33.5
145.8
81. 9
0.0
28.1
220.8
t*a~2
Western Waddensea
4th quarter
Netherlands
D
15.9
35.0
27.9
44.1
63.0
94.4
74.9
81.2
69.7
131. 5
90.7
83.8
153.8
117.2
35.9
97.4
143.5
'f
108.4
235.6
244.9
247.4
239.9
240.0
241.1
251.6
268.1
238.9
241.2
270.5
246.1
238.0
270.0
311.4
282.0
a~2
447.3
693.3
213.6
187.2
260.1
278.6
277.6
192.5
168.2
386.9
192.6
162.9
290.9
0.0
0.0
242.2
368.1
t*a~2
4
release area:
release period:
source:
No. of
Quarter returns
0 1
10
137
0 2
127
0 3
31
0 4
27
1 1
32
1 2
20
1 3
8
1 4
3
2 1
2
2 2
1
2 3
3 2
2
1
3 3
•
release area:
release period:
source:
No. of
Quarter returns
24
0 2
43
0 3
20
0 4
2
1 1
36
1 2
25
1 3
2
1 4
3
2 1
11
2 2
11
2 3
4
2 4
3 1
1
3 2
2
3 3
5
4 1
2
4+
1
release area:
release period:
source:
No. of
Quarter returns
0 3
20
0 4
49
80
1 1
1 2
235
1 3
91
47
1 4
2 1
15
2 2
81
22
2 3
7
2 4
3 1
6
3 2
6
3 3
8
3
3 4
4 1
1
3
4 2
4+
7
APPENDIX II
Dutch coast
1st quarter
Netherlands
D
11.8
27.0
35.9
45.5
66.4
60.0
66.4
77.6
59.1
53.4
159.9
95.7
180.2
'l'
282.2
244.3
246.2
250.0
246.6
235.0
245.1
236.1
232.9
270.0
306.9
287.0
287.4
aA 2
75.7
324.3
110.0
137.9
245.8
170.7
105.0
104.6
34.3
95.5
0.0
133.9
0.0
V
5.7
6.3
5.1
4.1
4.2
3.3
3.2
3.2
2.3
0.7
5.5
2.1
4.4
t
1.5
3.4
6.3
8.5
12.4
14.7
17.9
21.3
25.0
27.0
29.0
39.0
41.0
V*t
8.5
21.1
31. 9
35.2
52.0
49.0
57.7
67.2
56.4
18.3
159.9
82.4
180.2
t*a A 2
113.6
1086.6
694.5
1178.6
3041. 2
2512.9
1874.1
2223.6
857.8
2579.5
0.0
5221.1
0.0
aA 2
273.1
478.9
253.2
5.2
396.3
85.1
61. 4
236.4
443.4
108.1
328.4
0.0
5.7
125.1
231.1
0.0
t
V
9.0 1.2
3.1 2.7
3.4 4.8
3.2 9.0
1.2 11. 3
1.9 13.6
2.5 16.5
3.4 20.7
1.8 22.8
1.5 26.1
1.4 29.5
5.4 32.0
0.1 35.5
1.5 37.2
2.9 43.5
0.3135.0
V*t
10.9
8.6
16.4
28.7
14.1
25.5
41.7
70.4
42.1
39.9
41.0
172 .2
3.0
56.9
126.9
36.1
t*a A 2
330.0
1314.2
1228.2
46.4
4491. 9
1154.3
1012.3
4886.3
10117.0
2819.5
9689.1
0.0
201. 2
4652.3
10051.0
0.0
t
1.1
2.6
5.8
8.6
11.2
14.1
18.0
20.2
23.1
25.7
30.5
32.3
35.9
38.3
42.0
44.0
70.3
V*t
22.2
6.7
4.9
9.7
21.3
32.2
67.7
41. 4
49.3
20.2
80.3
40.2
45.3
66.0
181.7
46.4
82.1
t*a A 2
582.1
502.4
1433.0
2108.1
1779.0
3009.2
4466.3
3217.4
5799.1
2897.8
9660.4
928.5
6856.7
3253.1
0.0
1236.8
4367.9
Dutch coast
2nd quarter
Netherlands
D
17.2
26.9
33.6
28.7
45.3
35.1
49.4
75.0
65.7
47.0
99.3
172 .2
14.4
64.5
127.0
36.1
'l'
150.5
303.5
210.3
271. 0
293.4
322.3
315.9
239.2
315.4
329.4
291.1
231.2
180.5
329.9
222.9
19.7
Dutch coast
3rd quarter
Netherlands
D
30.6
17.6
29.8
29.4
36.2
46.9
80.8
55.0
68.2
47.1
85.4
43.5
80.6
84.2
181.7
47.2
95.5
\}l
93.9
136.3
144.6
234.5
251.9
246.2
239.1
236.3
237.2
312.7
231.2
227.8
265.9
246.8
308.9
223.0
266.4
aA2
529.2
193.8
246.0
245.9
159.3
213.0
248.1
159.0
250.7
112.7
316.7
28.7
191.1
84.9
0.0
28.1
62.1
V
20.2
2.6
0.8
1.1
1.9
2.3
3.8
2.0
2.1
0.8
2.6
1.2
1.3
1.7
4.3
1.1
1.2
5
release area:
release period:
source:
No. of
Quarter returns
33
0 4
74
1 1
1 2
162
1 3
140
21
1 4
2 1
12
2 2
20
2 3
11
8
2 4
2
3 1
3 2
2
3 3
1
1
4 3
release area:
release period:
source:
No. of
Quarter returns
19
0 1
84
0 2
0 3
102
0 4
20
13
1 1
36
1 2
26
1 3
10
1 4
2 1
2
9
2 2
2 3
1
2 4
1
3 2
2
4+
1
release area:
release period:
source:
No. of
Quarter returns
0 3
45
0 4
10
1 1
9
1 2
12
1 3
11
1 4
4
2 1
5
2 2
3
2 3
2
2 4
2
3 2
1
3 3
1
4+
1
APPENDIX II
Dutch coast
4th quarter
Netherlands
D
14.6
10.6
21.7
32.7
44.5
57.5
57.7
81.3
48.7
124.5
43.3
48.1
18.1
'P
87.6
241.1
246.7
240.1
252.6
257.4
245.9
263.4
234.4
227.6
261. 6
355.7
270.0
a"2
279.8
192.3
125.1
93.2
88.0
204.0
278.6
164.7
25.3
0.2
11. 9
0.0
0.0
V
12.1
2.6
2.7
2.8
3.1
3.0
2.7
3.2
1.9
4.5
1.3
1.4
0.4
t
1.1
2.4
6.4
9.6
12.0
16.1
18.1
20.9
24.3
27.5
31. 0
35.0
46.0
V*t
13.2
6.2
17.2
27.1
37.3
48.3
49.2
66.8
46.4
124.5
40.9
48.1
18.1
t*a"2
305.2
459.9
802.5
898.4
1059.7
3281.1
5056.9
3443.2
613.6
5.1
369.1
0.0
0.0
a"2
371.3
205.0
181. 6
330.9
288.3
316.1
451.7
694.4
156.6
123.4
0.0
0.0
2.5
0.0
V
8.1
7.8
5.6
5.2
5.0
2.8
3.1
3.2
5.6
2.1
7.7
1.7
1.3
2.2
t
1.7
4.3
6.7
9.6
12.7
15.9
18.4
21.2
25.5
27.7
30.0
33.0
39.5
81.0
y*t
13.6
33.1
37.4
50.1
62.8
44.6
57.6
68.7
142.8
56.9
232.3
56.0
52.3
174.3
t*a"2
625.3
873.5
1212.6
3176.8
3659.0
5031.8
8321.2
14721. 7
3993.6
3413.9
0.0
0.0
100.4
0.0
a"2
452.3
133.4
545.5
50.4
647.2
64.4
109.5
459.1
9.8
104.1
0.0
0.0
0.0
y
16.5
8.9
7.8
3.4
4.4
3.8
3.0
4.5
3.7
5.7
4.8
5.3
5.1
t
2.0
4.6
8.2
11.0
13.7
16.8
20.8
22.0
25.5
29.0
34.0
38.0
60.0
y*t
33.7
40.9
64.0
37.7
60.2
64.4
61. 7
98.2
94.1
163.9
162.0
203.0
304.9
t*a"2
924.7
613.7
4485.5
554.2
8883.8
1078.9
2277.0
10100.4
251. 0
3020.3
0.0
0.0
0.0
Scheldt Estuary
1st quarter
Netherlands
D
13.9
37.5
45.6
64.9
76.6
57.1
80.6
111. 5
149.1
71.7
232.3
56.0
53.0
174.3
'P
255.3
267.7
282.4
294.8
300.0
291.0
304.9
293.1
345.1
280.8
331.4
344.7
273.2
231.7
•
Scheldt Estuary
2nd quarter
Netherlands
D
37.2
43.0
80.6
42.1
83.5
69.6
74.6
100.0
94.3
172.6
162.0
203.0
304.9
'P
278.9
317 .2
269.4
279.2
290.7
327.0
274.1
259.4
319.9
342.1
1.5
329.0
15.4
6
release area:
release period:
source:
No. of
Quarter returns
0 3
24
0 4
35
1 1
46
152
1 2
196
1 3
29
1 4
15
2 1
2 2
56
27
2 3
2 4
15
6
3 1
14
3 2
3
3 3
4
3 4
4 1
2
1
4 2
4+
6
•
release area:
release period:
source:
No. of
Quarter returns
5
0 4
25
1 1
135
1 2
205
1 3
70
1 4
33
2 1
56
2 2
2 3
32
16
2 4
3 1
10
3 2
7
3 3
2
3 4
1
4 1
1
4 2
4
4+
1
release area:
release period:
source:
No. of
Quarter returns
8
0 3
0 4
3
1 1
2
6
1 2
1 3
2
2 2
1
APPENDIX I r
Scheldt Estuary
3rd quarter
Netherlands
D
'l'
6.4
8.7
22.2
36.9
42.2
45.3
82.7
71.2
62.1
51.8
94.5
80.0
50.7
144.1
197.2
43.5
168.0
293.8
65.0
263.6
271.7
283.2
293.3
257.0
276.5
303.3
299.4
258.0
307.5
297.6
288.7
244.5
290.1
284.2
a~2
125.8
43.2
176.6
95.3
112.9
76.5
244.1
279.7
114.4
29.4
142.5
310.2
13.3
544.1
71. 9
0.0
378.0
V
2.4
1.2
3.3
3.9
3.3
2.7
4.4
2.8
2.1
1.9
3.0
2.0
1.3
2.9
4.6
0.9
1.8
t
1.5
3.0
5.7
8.7
10.8
14.0
17.4
20.1
23.4
25.3
28.7
32.1
35.3
37.3
42.0
46.0
63.0
V*t
3.7
3.7
18.7
33.7
35.8
38.3
76.5
56.1
50.2
47.2
86.4
62.6
47.5
107.9
194.9
43.5
112.9
V
2.1
7.4
4.8
3.9
3.2
3.8
3.0
2.7
2.4
3.7
3.0
1.7
5.1
2.7
1.9
3.7
t
1.4
3.9
6.4
9.1
12.0
15.7
18.6
21.8
24.4
28.1
30.3
34.5
36.0
39.0
42.0
61. 0
V*t
3.0
29.0
30.9
35.7
38.1
59.7
56.0
59.4
58.5
103.7
92.3
57.2
185.1
105.2
78.2
223.3
22.2
2541.2
585.7
867.4
1091.5
5774.6
5009.5
8167.8
3482.6
8354.5
18437.6
20.0
0.0
0.0
7881.9
0.0
V*t
1.7
10.2
5.4
3.0
27.4
69.2
t*a"2
23.2
46.6
55.7
1345.8
98.7
0.0
t*a~2
194.0
128.3
1009.8
831.1
1220.6
1068.6
4247.2
5633.1
2673.2
742.3
4085.5
9949.3
468.9
20267.2
3019.0
0.0
23815.4
Scheldt Estuary
4th quarter
Netherlands
D
3.0
31.3
34.5
41.5
47.2
83.4
66.7
85.0
71. 7
118.7
108.5
57.3
185.1
105.2
91.7
223.3
'l'
270.0
257.6
273.9
294.0
287.9
287.9
271. 8
326.1
307.1
259.3
309.1
294.8
318.2
345.9
266.7
251.2
a~2
15.9
648.3
91. 5
94.8
91. 3
368.6
269.0
374.5
142.9
297.3
608.8
0.6
0.0
0.0
187.7
0.0
t*a~2
North Sea, French/Belgian Border
2nd quarter
France
D
3.5
10.2
7.6
32.4
27.7
69.2
'l'
2.3
64.8
68.4
282.6
260.6
31.5
a"2
10.9
11. 6
8.0
122.3
6.8
0.0
V
t
0.8 2.1
2.5 4.0
0.8 7.0
0.3 11.0
1.9 14.5
3.0 23.0
7
release area:
release period:
source:
No. of
Quarter returns
61
0 3
0 4
82
1 1
9
1 2
35
1 3
9
1 4
11
2 1
5
2 2
6
7
2 3
3 3
1
3 4
1
4 1
1
4 2
1
4+
2
release area:
release period:
source:
No. of
Quarter returns
0 2
2
0 4
1
1 1
3
1 2
4
1 3
6
1 4
4
2 1
1
2 2
5
2 3
4
2 4
1
3 2
3
3 3
1
4 1
2
4+
9
release area:
release period:
source:
No. of
Quarter returns
2
0 3
0 4
1
1 1
3
1 2
4
1 3
12
8
1 4
2 1
10
9
2 2
7
2 3
2 4
3
3 1
2
3
3 2
3 3
4
3 4
1
4 1
4
4 2
1
4+
14
APPENDIX Ir
North Sea, French/Belgian Border
3rd quarter
France
D
'P
6.3
3.7
18.8
17.2
35.6
28.4
28.7
46.4
76.1
118.6
7.7
100.2
109.4
133.6
271. 3
58.4
71. 3
57.8
210.1
6.9
271.0
43.9
287.4
16.0
24.2
222.4
223.0
334.8
aA2
87.6
23.4
58.4
62.5
121. 4
151. 5
61. 2
104.4
517.8
0.0
0.0
0.0
0.0
307.7
V
1.2
0.4
1.7
0.2
2.3
1.3
0.6
1.3
1.6
3.1
0.2
2.3
2.5
1.0
t
1.1
2.3
6.4
9.3
12.4
14.8
18.6
22.0
23.6
38.0
39.0
43.0
44.0
61. 5
V*t
1.3
0.8
11.1
1.5
28.5
20.0
11. 9
27.7
36.8
118.6
7.7
100.2
109.4
64.1
t*a A 2
97.7
54.9
376.6
580.1
1510.8
2244.4
1138.4
2296.2
12205.1
0.0
0.0
0.0
0.0
18926.5
aA2
0.0
0.0
144.7
419.8
229.5
325.9
0.0
124.4
228.9
0.0
461.1
0.0
4.8
276.7
V
0.6
14.4
12.6
8.3
5.7
5.9
5.5
5.1
3.5
1.3
3.0
5.4
1.6
2.0
t
1.0
6.0
8.3
10.8
14.5
16.5
21.0
23.2
26.5
30.0
35.3
38.0
43.5
62.3
V*t
0.6
86.2
104.6
89.7
82.3
96.6
115.0
118.5
93.6
39.8
104.7
203.5
71.0
125.4
t*a A 2
0.0
0.0
1206.2
4512.8
3328.3
5377.9
0.0
2887.2
6064.9
0.0
16293.6
0.0
208.7
17245.1
a"'2
282.1
0.0
1053.7
519.9
601. 8
80.3
184.0
88.4
134.2
21. 2
64.4
32.8
96.2
0.0
38.5
0.0
71. 4
V
22.5
12.9
18.7
10.7
5.4
5.7
4.4
4.8
3.5
1.7
2.0
4.5
1.9
1.1
2.3
1.8
1.1
t
1.5
4.0
7.7
9.8
12.8
15.5
19.4
22.0
25.3
28.3
31.0
34.0
37.0
39.0
43.3
45.0
79.5
y*t
33.7
51. 7
143.6
104.4
69.4
88.0
85.8
105.7
88.4
48.5
63.1
153.5
69.8
41.3
98.9
80.0
91.2
t*a A 2
423.2
0.0
8078.5
5068.6
7722.9
1244.5
3568.7
1943.9
3393.8
601. 7
1996.8
1114.7
3558.7
0.0
1664.0
0.0
5674.3
Wash
2nd quarter
UK, England
D
'P
0.6
86.2
104.9
100.7
93.2
103.9
115.0
120.7
101. 6
39.8
158.3
203.5
71.1
162.3
90.0
4.3
6.4
16.0
0.8
43.8
1.1
1.9
4.9
28.5
6.6
4.0
359.0
3.2
•
Wash
3rd quarter
UK, England
D
'P
33.8
51.7
146.5
104.8
88.9
88.6
92.0
106.4
94.4
51.1
77.2
156.6
81.3
41.3
99.3
80.0
105.0
353.9
48.9
353.3
5.2
9.0
2.5
3.5
358.8
13.5
12.5
36.1
349.6
13.8
7.4
8.6
358.9
15.9
8
release area:
release period:
source:
No. of
Quarter returns
0 2
110
0 3
491
94
0 4
1 1
93
155
1 2
96
1 3
58
1 4
2 1
24
35
2 2
16
2 3
10
2 4
3 1
9
11
3 2
3 3
9
3 4
6
4 1
3
4+
27
release area:
release period:
source:
No. of
Quarter returns
0 2
79
0 3
106
0 4
44
30
1 1
1 2
54
1 3
25
1 4
5
2 1
5
2 2
6
9
2 3
2 4
1
3 1
3
3 2
3
3 3
1
4 1
1
4+
8
release area:
release period:
source:
No. of
Quarter returns
0 3
4
0 4
2
5
1 1
1 2
5
1 3
3
2 1
1
2 2
2
2 4
2
3 1
1
3 3
1
APPENDIX TI
Thames
2nd quarter
UK, England
D
7.9
12.2
31. 6
84.0
56.5
61.0
83.9
90.2
113.2
155.9
139.6
131.1
194.6
134.3
167.4
249.8
170.1
'l'
87.9
95.8
45.5
354.4
3.1
8.8
15.0
13.4
352.7
342.6
3.4
345.8
344.5
353.2
341.8
22.5
353.6
aA2
199.8
146.8
484.9
826.2
638.8
480.8
473.2
274.2
656.4
688.1
551.1
228.5
671. 7
187.5
311. 0
354.7
247.0
V
2.8
1.1
2.3
6.6
2.4
2.7
3.5
3.3
3.2
4.2
3.4
3.3
4.1
3.4
3.2
5.3
2.3
t
1.0
2.4
5.3
8.8
11. 8
14.5
17.3
20.5
23.6
26.4
29.4
32.4
35.6
38.1
41. 8
45.3
64.3
V*t
2.9
2.7
12.0
58.4
28.1
39.5
60.5
67.5
74.3
112.1
101. 4
106.8
147.6
128.0
134.7
238.9
146.8
t*a A 2
201.6
356.1
2574.3
7257.7
7562.7
6976.2
8166.7
5633.3
15473.0
18190.8
16202.9
7415.1
23937.8
7146.9
13011.8
16080.6
15891. 7
aA2
106.8
47.6
144.2
545.5
100.6
307.0
414.6
57.4
647.6
296.3
0.0
1.1
142.6
0.0
0.0
248.2
V
0.3
0.4
1.4
1.6
0.4
1.1
1.9
1.1
3.3
1.3
0.3
0.2
1.3
0.1
5.0
0.7
t
1.0
3.1
5.9
9.3
11. 9
14.6
18.2
21. 6
24.3
26.8
30.0
33.3
35.7
38.0
46.0
57.8
V*t
0.3
1.2
8.3
14.8
4.6
15.4
34.4
24.2
80.4
35.0
8.0
8.2
46.3
3.3
231.2
40.2
t*a A 2
106.8
146.8
851. 9
5091.3
1192.9
4495.0
7545.8
1240.5
15758.2
7933.7
0.0
35.4
5086.7
0.0
0.0
14331. 6
aA2
5.5
6.5
3.5
2.0
25.7
0.0
0.7
9.2
0.0
0.0
y
1.9
1.6
0.6
0.1
0.9
1.4
0.1
0.5
0.3
0.4
t
1.8
4.0
7.2
10.2
12.7
19.0
22.0
27.5
31. 0
37.0
y*t
3.3
6.5
4.7
0.8
11.0
26.5
3.2
14.6
8.6
15.4
t*a A 2
9.6
26.1
25.2
20.8
325.9
0.0
14.3
253.1
0.0
0.0
Rye
2nd quarter
UK, England
D
'Y
4.5
5.5
13.7
33.6
13.7
27.2
51.4
28.1
95.8
49.9
8.0
8.5
60.4
3.3
231.2
51.0
197.8
13.9
48.0
17 .4
19.6
340.2
26.4
203.2
358.0
18.1
240.0
232.5
359.6
252.4
357.4
356.1
Rye
3rd quarter
UK, England
D
4.0
6.8
5.7
3.6
15.1
26.5
4.1
18.0
8.6
15.4
'l'
243.5
256.5
226.7
89.4
54.9
107.6
231. 5
173.7
234.3
251.0
9
release area:
release period:
source:
No. of
Quarter returns
0 2
22
0 3
55
0 4
6
1 1
11
1 2
30
7
1 3
1 4
9
2 1
3
2 2
4
2 4
1
3 1
1
3 2
1
4+
2
release area:
release period:
source:
No. of
Quarter returns
0 3
6
0 4
2
1 2
1
2 1
1
2 2
1
2 4
1
3 3
1
3 4
1
4+
1
release area:
release period:
source:
No. of
Quarter returns
0 4
5
1 1
8
1 2
28
1 3
17
1 4
4
2 1
3
2 2
6
2 3
3
2 4
2
3 1
1
3 2
1
3 3
1
3 4
1
4+
3
APPENDIX I I
Poole
2nd quarter
UK, Enlgand
D
2.2
2.6
3.1
14.6
2.5
3.8
22.1
32.9
27.2
103.7
40.6
1.0
2.3
\f
191. 6
212.4
112.2
249.5
160.0
201. 3
260.7
235.2
258.0
254.3
249.8
179.0
205.4
a"2
3.3
1.8
0.5
159.3
0.5
1.6
64.7
94.7
86.4
0.0
0.0
0.0
0.0
V
1.7
0.7
0.5
1.4
0.2
0.1
1.2
1.4
1.1
3.5
1.3
0.0
0.0
t
1.0
2.7
6.0
9.4
12.1
15.0
17.9
21.3
23.0
30.0
32.0
37.0
67.5
V*t
1.7
1.9
3.1
12.7
1.9
2.0
21. 9
30.3
26.4
103.7
40.6
1.0
2.2
t*a"2
3.3
4.8
3.0
1492.0
5.6
24.0
1157.3
2020.7
1987.3
0.0
0.0
0.0
0.3
a"2
37.7
4.3
0.0
0.0
0.0
0.0
0.0
0.0
0.0
V
1.3
0.6
0.4
0.1
5.5
6.6
0.8
0.5
0.1
t
2.8
6.0
12.0
22.0
23.0
29.0
38.0
42.0
47.0
y*t
3.6
3.5
4.4
1.2
126.5
191. 4
31. 6
19.1
4.6
t*a"2
106.9
26.0
0.0
0.0
0.0
0.0
0.0
0.0
0.0
a"2
17.2
10.8
35.8
39.3
24.4
0.4
64.4
391.5
0.5
0.0
0.0
0.0
0.0
180.4
V
0.4
0.6
0.4
0.3
0.7
0.3
1.8
3.5
0.1
0.3
0.8
0.2
0.2
0.8
t
1.6
4.5
7.2
9.6
12.5
16.3
19.3
21.3
24.0
28.0
31. 0
35.0
38.0
73.0
y*t
0.7
2.5
3.0
2.7
8.3
5.0
34.2
75.6
1.6
8.9
26.1
5.9
7.2
61. 3
t*a"2
27.6
48.4
258.1
376.6
304.8
6.8
1245.6
8351.4
12.8
0.0
0.0
0.0
0.0
13165.9
Bay de Seine
2nd quarter
France
D
7.6
5.8
4.4
1.2
126.5
191.4
31.6
19.1
4.6
\f
123.3
235.8
242.8
213.0
272.7
269.1
104.7
54.8
282.4
•
Baye de Seine
4th quarter
France
D
4.1
6.8
11.0
14.8
17 .0
5.5
35.1
75.9
3.6
8.9
26.1
5.9
7.2
86.6
\f
269.1
342.8
57.6
51. 5
72.6
2.4
310.4
284.8
202.8
154.0
40.1
90.0
270.0
289.0
10
release area:
release period:
source:
No. of
Quarter ret\lrnS
7
0 3
7
0 4
3
1 1
9
1 2
1 3
11
1 4
1
2 1
2
2 2
4
2 4
2
3 1
1
1
3 2
2
4 1
4+
3
•
release area:
release period:
source:
No. of
Quarter returns
0 4
10
1 1
9
32
1 2
1 3
12
4
1 4
2 1
6
2 2
15
2 3
8
7
2 4
3 1
1
3 2
3
3 3
1
3 4
1
4 1
1
4 3
2
4+
3
release area:
release period:
source:
No. of
Quarter returns
0 2
1
2 3
1
release area:
release period:
source:
No. of
Quarter returns
1 1
1
1 2
20
1 3
2
2 1
1
2 2
3
APPENDIX Ir
Bay de Somme
3rd quarter
France
D
8.1
4.8
3.3
7.8
10.0
3.2
79.5
96.4
11.9
74.9
52.5
52.7
44.9
'Y
351.6
40.2
260.3
336.5
322.1
232.0
244.2
269.1
231.0
227.1
203.9
231.7
260.0
a"2
88.6
11.1
1.1
12.5
11.0
0.0
227.9
377.2
3.2
0.0.
0.0
5.4
32.9
V
7.1
0.4
0.3
0.3
0.3
0.2
3.2
4.4
0.5
2.6
1.6
1.2
0.7
t
1.0
2.3
6.7
8.7
12.0
14.0
17.5
21.3
26.0
29.0
32.0
41.0
60.3
V*t
7.1
0.9
2.0
2.5
3.1
3.2
55.2
93.3
11. 9
74.9
52.5
50.8
42.4
t*a"2
88.6
25.4
7.1
108.6
132.3
0.0
3987.7
8015.3
84.0
0.0
0.0
221. 0
1986.4
a"2
171.1
203.1
106.1
71. 9
133.9
1335.0
242.0
208.1
12.1
0.0
16.0
0.0
0.0
0.0
8.6
10.9
V
4.6
2.7
2.0
1.0
2.8
6.4
2.7
5.4
0.9
0.4
1.3
4.8
0.4
1.6
0.6
0.8
t
1.4
4.7
7.5
9.9
13.3
16.5
18.9
21. 6
24.7
29.0
31.7
35.0
37.0
41.0
47.0
62.0
V*t
6.5
12.5
14.9
9.6
37.1
105.7
50.9
116.8
21.2
12.8
41.2
167.2
14.6
64.1
29.0
47.7
t*a"2
239.5
947.6
792.5
713.2
1773.9
22027.4
4582.1
4500.4
299.2
0.0
505.9
0.0
0.0
0.0
402.0
673.9
a"2
0.0
0.0
V
t
0.0 1.0
0.4 28.0
V*t
0.0
11.1
t*a"2
0.0
0.0
a"2
0.0
291.7
117.8
0.0
104.5
V
t
1.8 4.0
0.8 6.3
13.8 10.0
8.2 16.0
6.1 18.0
V*t
7.2
5.0
138.0
130.9
110.5
t*a"2
0.0
1852.1
1178.1
0.0
1881. 5
Bay de Somme
4th quarter
France
D
13.6
20.7
19.8
23.3
44.4
110.2
62.6
118.5
21.8
12.8
41.4
167.2
14.6
64.1
29.0
51.0
'Y
9.5
233.7
242.4
306.8
280.1
266.4
261. 9
264.1
241. 8
270.0
233.3
263.5
236.7
224.1
263.1
282.5
Bay du St. Michel
2nd quarter
France
D
0.0
11.1
'Y
0.0
186.8
Bay du St. Michel
4th quarter
France
D
7.2
16.9
138.4
130.9
112.8
'f
326.6
351.7
303.9
315.3
316.3
11
APPENDIX III
Maps of quarterly recaptures of plaice, by nursery release area and tagged with a length
< 25 cm, which have been in sea for more than 12 months.
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Appendix IV.
REC31FO
REC31Fl
REC31F2
REC31F3
REC32FO
REC32Fl
REC32F2
REC32F3
REC33Fl
REC33F2
REC33F3
REC33F4
REC34Fl
REC34F2
REC34F3
REC34F4
REC35FO
REC35F1
REC35F2
REC35F3
REC35F4
REC35F5
REC36E9
REC36FO
REC36F1
REC36F2
REC36F3
REC36F4
REC36F5
REC36F6
REC36F7
REC37E9
REC37FO
REC37Fl
REC37F2
REC37F3
REC37F4
REC37F5
REC37F6
REC37F7
REC38E9
REC38FO
REC38Fl
REC38F2
REC38F3
REC38F4
REC38F5
REC38F6
REC38F7
REC39E8
REC39E9
REC39FO
REC39Fl
REC39F2
REC39F3
REC39F4
REC39F5
REC39F6
REC39F7
REC40E7
REC40E8
REC40E9
REC40FO
REC40Fl
REC4CF2
REC40F3
REC4CF4
RI:;C4CF5
REC40F6
REC40F7
Relative fishinq effort in Dutch beam trawl hours by ICES rectanqle
Ql
Q2
Q3
Q4
19
809
385
4313
0
166
4602
4920
109
5325
12988
356
58
2839
15887
1886
335
846
2350
4986
3519
333
2
1625
9276
7626
10301
2895
4982
1283
17
4233
7623
3809
12694
17107
15557
3897
3425
691
3450
1189
1332
553
3289
21711
5777
10482
686
282
559
429
408
431
482
3549
5586
6060
424
127
452
57
48
20
788
844
5232
3076
858
926
767
3795
835
5743
85
660
50
5245
1325
1746
19052
3673
9
1053
21864
10958
253
762
1111
10305
15738
1058
0
405
4477
6394
12497
7997
16993
4789
2928
7420
7675
7572
1926
2006
4863
2905
6684
9165
7008
1754
2558
933
136
795
1434
3939
6710
1181
1427
745
4455
3227
420
315
645
4984
13114
94
977
246
37
112
6330
1198
3996
550
905
3841
204
4163
944
3900
0
713
1369
3172
3693
473
15945
2558
15
1431
12751
10463
183
1018
3874
10666
11575
563
25
746
2728
5801
15048
4366
15291
6642
721
8325
5023
4572
2720
4535
3742
5139
14483
13148
5127
4559
1914
631
459
1908
2408
7309
4312
656
2182
1447
5079
2141
831
489
762
4189
7689
46
128
90
34
212
6119
1092
2933
1285
1544
4676
0
2075
102
3414
14
233
960
3859
1435
834
13141
672
0
1108
8605
3045
252
471
3223
6817
4974
284
6
1063
4993
5437
10126
4425
6682
2271
149
6380
5753
7943
9102
7712
9964
4423
7337
4253
3457
1063
1665
700
1469
9663
6296
5067
3052
463
909
555
3468
662
675
770
1943
4461
1839
97
167
94
16
31
1477
1355
1868
1962
1440
1862
Ql
REC41E7
REC41E8
REC41E9
REC41FO
REC41Fl
REC41F2
REC41F3
REC41F4
REC41F5
REC41F6
REC41F7
REC42E7
REC42E8
REC42E9
REC42FO
REC42Fl
REC42F2
REC42F3
REC42F4
REC42F5
REC42F6
REC42F7
REC43E7
REC43E8
REC43E9
REC43FO
REC43F1
REC43F2
REC43F3
REC43F4
REC43F5
REC43F6
REC43F7
REC44E6
REC44E7
REC44E8
REC44E9
REC44FO
REC44F1
REC44F2
REC44F3
REC44F4
REC44F5
REC44F6
REC45E6
REC45E7
REC45E8
REC45E9
REC45FO
REC45Fl
REC45F2
REC45F3
REC45F4
REC45F5
REC45F6
95
42
13
17
153
6
154
90
3731
578
70
109
42
18
68
14
4
1765
913
378
2155
63
10
61
86
166
22
6
1
45
0
287
492
112
635
342
116
68
20
5
13
77
4
1
48
115
168
134
87
16
27
15
5
1
0
Q2
102
91
43
39
141
145
110
66
1872
1245
687
128
202
237
238
65
107
5433
376
103
2305
245
22
414
575
148
59
76
62
1541
88
2158
920
24
198
349
196
116
59
2147
2514
10052
22
5
52
151
164
142
72
30
1560
104
11
0
0
Q3
67
85
47
62
149
54
2
68
2034
2083
5267
93
138
78
257
82
72
2664
528
208
5678
538
24
213
260
498
120
91
33
2567
124
3639
1550
24
142
395
115
120
172
1446
1538
7390
8
1
58
144
90
58
56
81
657
64
17
0
0
Q4
64
47
21
31
252
217
484
94
2324
386
8479
89
112
27
426
36
11
1912
585
124
3220
422
9
118
53
375
58
18
4
882
11
990
797
92
478
414
106
81
65
325
639
2025
8
4
45
169
126
80
62
48
116
36
8
2
0