December 1982 arJ ECOLOGICAL SURVEY OF THE PROPOSED CEDAR B A S I N RESEARCH NATURaL AREA, SHASTA-TRINITY NATIONAL F O R E S T , CALIFORNIA ( P u r c h a s e order 40-9AD6-2-606) Todd Keeler-Wolf INTRODUCTION L o c a t i o n and P r i n c i p a l D i s t i n g u i s h i n g F e a t u r e s History of S c i e n t i f i c Interest JUSTIFICATIONS T h e P o r t O r f o r d Cedarof C C e d a r B a s i n T h e s i g n i f i c a n c e of the i n l a n d POC p o p u l a t i o n E n d e m i c and U n u s u a l K l a m a t h and T r i n i t y M o u n t a i n s Flora U n u s u a l B o g Species The High D i v e r s i t y of E r i c a c e a e Z o o l o g i c a l Justification PHYSICAL FEATURES G e o r n o r p h o l o g y and T o p o g r a p h y Geology Climate VEGETATION Shallow Lakes and Ponds B o g and M e a d o w P o r t O r f o r d Cedar Forest M i x e d C o n i f e r Forest Red F i r - M o u n t a i n H e m l o c k Forest Mountain Chaparral Rock O u t c r o p BOUNDARIES 10 IMPACTS RECOMMENDATIONS LITERATURE C I T E D APPENDIX V a s c u l a r Plant L i s t V e r t e b k a t e s Known o r Suspected from Cedar B a s i n Vegetation P l o t ~escriptions Description o f Arctostaphylos klamathensis Edwards INTRODUCTION Location and P r i n c i p a l D i s t i n u u i s h i n a F e a t u r e s : The Cedar Basin c a n d i d a t e RNA c o v e r s approximately 874 a c r e s of r.ed fir-Mountain hemlock, mixed c o n i f e r , and P o r t Orford c e d a r f o r e s t w i t h a d d i t i o n a l mountain c h a p a r r a l , rock o u t c r o p , l a k e , bog, and meadow communities i n t h e n o r t h e r n T r i n i t y Mountains of n o ~ t h w e s t e r n California. The proposed RNA i s on t h e S h a s t a - T r i n i t y N a t i o n a l F o r e s t i n extreme s o u t h - c e n t r a l S i s k i y o u County. I t l i e s w i t h i n p o r t i o n s of s e c t i o n s 25 and 36 o f Township 39 ,Na.rhh, Range 6 West, and s e c t i o n s 30 and 31 of T 39 N, R 5 W. E l e v a t i o n s range from c a . 5420 t o 7149 f e e t . ( s e e map 1). The a r e a i s about 1 2 road m i l e s west of M t . S h a s t a C i t y and may be. approached t o w i t h i n a q u a r t e r m i l e by a good, o i l e d road ( 4 0 N 2 6 ) . A p o o ~ l ymaintained j e e p road(39N05Y) l e a d s s o u t h o f f t h i s road c a . one m i l e t o Cedar Lake, i n s i d e t h e proposed boundaries. Cedar Basin c o n t a i n s t h e h i g h e s t e l e v a t i o n s t a n d s of Chamaecyparis l a w s o n i a n a . ( P o r t Oxford c e d a r ) . e r n l i m i t of i t s range h e r e . This s p e c i e s i s a l s o n e a r t h e e a s t - The t r e e s form a dense f o r e s t around l a k e s , s e e p s , and s t r e a m s up t o almost 6400ft. FOE s e v e r a l e c o l o g i c a l , phytogeographical, and g e n e t i c r e a s o n s t h e s e s t a n d s a r e unique and promise t o be t h e s u b j e c t of much 6 u t u r e r e s e a r c h . Also c o n t a i n e d w i t h i n thBs ,smar3.1 b a s i n a r e two h i g h l y r e s t r i c t e d s p e c i e s of p l a n t s , a manzanita, A r c t o s t a p h y l o s k l a m a t h e n s i s and a penstemon, Penstemon s p . nov. (as y e t n o t f o r m a l l y d e s c r i b e d ) , which so f a r appear t o be t o t a l l y r e s t r i c t e d t o t h e b a s i n . Several o t h e r s p e c i e s endemic t o the Klamath Province a l s o o c c u r h e r e . Another b o t a n i c a l v a l u e of t h e a r e a ' i s t h e bog v e g e t a t i o n around MAP 1 scale : 2 i n . = l r n i . Cedar and Lower C l i f f l a k e s . These bogs c o n t a i n s e v e r a l r a r e and unusual s p e c i e s i n c l u d i n g Drosera r o t u n d i f o l i a , D a r l i n g t o n i a c a l i f o r n i c a , and Menyanthes t r i f o l i a t a . Within t h e bogs and i n a d j a c e n t f o r e s t s and mountain c h a p a r r a l no l e s s t h a n n i n e genera and 13 s p e c i e s of E r i c a c e a e ( t h e Heath f a m i l y ) o c c u r , an e x t r e m e l y h i g h c o n c e n t r a t i o n f o r such a small a r e a . H i s t o r y of S c i e n t i f i c I n t e r e s t : Cedar Basin h a s been t h e s u b j e c t of b o t a n i c a l i n t e r e s t f o r several years. The C a l i f o r n i a Native P l a n t S o c i e t y h a s made two t r i p s t o t h e a r e a (July 1981, Oct. 1 9 8 2 ) t o i n v e s t i g a t e t h e bog and lower b a s i n f l o r a , and more r e c e n t l y t h e new s p e c i e s of manzanita. Mary Taylor, a s s i s t a n t S h a s t a - T r i n i t y N.F. botanist, also c o l l e c t e d twice i n t h e lower p o r t i o n of t h e b a s i n i n 1980. John Sawyer o f Humboldt S t a t e U n i v e r s i t y v i s i t e d t h e a r e a i n 1980, and Glen Keator of t h e S t r y b i n g Arboretum h a s c o l l e c t e d i n t h e b a s i n . I n June 1 9 7 6 t h e U n i v e r s i t y o f C a l i f o r n i a Santa Cruz C a l i f o r n i a N a t u r a l H i s t o r y C l a s s s p e n t t h r e e days s t u d y i n g v a r i o u s a s p e c t s of t h e ecology of t h e b a s i n . Apart from t h e v i s i t s a c t u a l l y made t o t h e a r e a , numerous o t h e r r e s e a r c h e r s have e x p r e s s e d i n t e r e s t i n preserving t h e hasin,primar- i l y because of i t s h i g h e l e v a t i o n , i n l a n d s t a n d s of P o r t Orford c e d a r (POC) and because of t h e p o t e n t i a l g e n e t i c d i f f e r e n c e s between t h e s e and t h e c o a s t a l s t a n d s of t h e s p e c i e s . Drs. These r e s e a r c h e r s i n c l u d e Adams, Hawk, Roth, and Zobel who have s t u d i e d t h e ecology of POC and c o n s i d e r t h e a r e a a s a v a l u a b l e g e n e t i c r e s o u r a e a s w e l l a s a p r o t e c t e d b a s t i o n a g a i n s t t h e r a p i d l y s p r e a d i n g and l e t h a l cedar r o o t r o t , p r e s e n t l y i n f l i c t i n g a g r e a t d e a l of damage on t h e coastal. POC p o p u l a t i o n s . JUSTIFICATIONS The P o r t Orford Cedar of Cedar Basin: Of primary importance t o f o r e s t e r s a r e t h e b a s i n ' s s t a n d s of POC, f o r which t h e a r e a r e c e i v e s i t s name. These s t a n d s a r e not o n l y t h e h i g h e s t e l e v a t i o n o f any POC groves i n t h e s p e c i e s ' n a t u r a l range (Steven Edwards, p e r s . cornm. ) , b u t a l s o a r e near t h e species ' e a s t e r n m o s t range l i m i t ( a b o u t 10 m i l e s e a s t a l o n g t h e upper Sacramento River) . POC has a l i m i t e d n a t u r a l range c e n t e r e d i n t h e Klamath Province of Southwestern Oregon and Northwestern C a l i f o r n i a . Most of t h e range i s c o a s t a l . However,a d i s t i n c t i n l a n d p o p u l a t i o n s e p a r a t e d from t h e c o a s t a l s t a n d s by a b o u t 50-60 m i l e s o c c u r s a l o n g t h e upper r e a c h e s o f t h r T r i n i t y , E a s t Fork of t h e T r i n i t y , and Sackamento R i v e r . d~ainages riffin in and C r i t c h f i e l d , 1 9 7 2 ) . A t p r e s e n t no e s t a b l i s h e d RNA's w i t h i n t h e R 5 r e g i o n i n c l u d e e x t e n s i v e s t a n d s o f POC, and o n l y one o t h e r c a n d i d a t e RNA :(Adorni, w i t h i n t h e main c o a s t a l populakion) is .being . considered. Cedar Basin i s t h e f i r s t c a n d i d a t e RNA withim t h e s p e c i e s ' i n l a n d d i s t r i b u $ i o n . The s p e c i e s , because of i t s l i m i t e d r a n g e , high economic v a l u e , and extremely t h r e a t e n e d n a t u r e due t o t h e r a p i d l y s p r e a d i n g r o o t r o t fungus, h a s been c o n s i d e r e d by D r . P e t e r Theisen, R6 r e g i o n a l g e n e t i c i s t , a s t h e s i n g l e most i m p o r t a n t t r e e s p e c i e s w i t h i n t h e region t o s t u d y and p r e s e r v e ( f i d e Ron K e l l y . ) .- The s i g n i f i c a n c e o f the i n l a n d POC p o p u l a t i o n : Raven and Axelrod (1978) c o n s i d e r POC t o be a s p e c i e s t h a t i s somewhat r e s t r i c t e d t o s e r p e n t i n e and o t h e r u l t r a m a f i c s o i l s . Griffin and C r i t c h f i e l d ( 1 9 7 2 ) note t h a t p a r t i c u l a r l y f a r t h e r i n l a n d POC a p p e a r s more r e s t r i c t e d t o s e r p e n t i n e s o i l s . I n the most r e c e n t :, review cif t h e s p e c i e s ' ecology (Zobel and Hawk, 1980) POC i s not considered t o be s t r o n g l y r e s t r i c t e d t o uktrarnafics on any p a r t of i t s range and Zobel and Hawk s t a t e a h i g h water t a b l e (permknent m o i s t u r e ) a s a more i m p o r t a n t l i m i t t o i t s d i s t r i b u t i o n . Despite t h i s , when a map of t h e s p e c i e s ' range (GrifiEin and G r i t c h f i e l d , 1972) i s compared t o a map of t h e u l t r a m a f i c o u t n r o p s i n t h e Klamath Province ( I r w i n , 1966) t h e main c o a s t a l and i n l a n d p o p u l a t i o n s c o i n c i d e almost e x a c t l y w i t h t h e two l a r g e s t u l t r a m a f i c e x p o s u r e s i n t h e r e g i o n . The i h l a n d d i s t r i b u t i o n l i e s almost e n t i r e l y on t h e T r i n i t y P l u t o n , a huge s h e e t - l i k e exposure of primariLy s e r p e n t i n i t e and gabbroic r o c k s w i t h a continuous w e s t e r n boundary 100 m i . Irwin 1 9 6 6 ) . l o n g . (Davis 1966, There i s no a c c u r a t e d a t e t o my knowledge f o r t h e time of exposure of t h i s s h e e t . However, i t i s c l e a r from f o s s i l evidence (Raven and Axelrod, 1978) t h a t POC had a much wider d i s t r i b u t i o n o f f of uLtramafics i n t h e mid T e r t i a r y ( t o a t l e a s t E. Oregon, W. Neva d a , and S . Idaho) . The p r e s e n t i n l a n d r k s t r i b t i 6 n : t o t h e s e r p e n t i n e b e l t probably o c c u r r e d i n t h e l a t e P l i o c e n e o r e a r l y P l e i s t o c e n e coinciding with i n c r e a s e d e r o s i o n a l p r o c e s s e s exposing s u b j a c e n t r o c k s (such a s t h e u l t r a r n a f i c s ) i n t h e Klamaths ( D i l l e r 1902, I r w i n 1966) and t h e c o n t i n u a l d r y i n g of t h e western U . S . u p l i f t of t h e c o r d i l l e r a n climate during t h e mountains, f o r c i n g t h e s p e c i e s t o move westward (Raven and Axelrod, 1978).Thus, t h e i n l a n d s t a n d s have proba b l y been i s o l a t e d f o r a t l e a s t one m i l l i o n y e a r s . Zobel and Hawk (1980) have shown t h a t a sample s t a n d of POC with- i n t h e i n l a n d d i s t r i b u t i o n ( C a s t l e Lake ca. 6 m i . NE o f t h e b a s i n . and s e v e r a l hundred f e e t lower i n e l e v a t i o n ) 1) had t h e s h o r t e s t grow- ing season, 2 ) t h e l o w e s t mean annual s o i l t e m p e r a t u r e , 3 ) t h e l o w e s t mean annual a i r t e m p e r a t u r e , 4 ) t h e c o l d e s t c o l d month a n n u a l l y , 5 ) b o t h t h e l o w e s t and t h e h i g h e s t annual Temperature Growth Index ;(lowe s t i n w i n t e r , h i g h e s t i n summer), and 6 ) t h e l o w e s t temperature recorded a t any of t h e i r s i t e s d u r i n g t h e s t u d y O OF) . Because of t h e long-term i s o l a t i o n i n a c o l d e r , more extreme c l i m a t e t h e l i k e l j h o o d of g e n e t i c d i f f e r e n t i a t i o n ! - i n t h e i n l a n d populations is r e l a t i v e l y high. T h i s d i f f e r e n t i a t i o n could be s i g - n i f i c a n t i n t h e maintenance = o f v i a b l e POC r e s e r v e s i n a number o f ways. I t h a s been suggested (Donald Zobel, Lewis Roth, i n l i t . ) t h a t t h e i n l a n d high e l e v a t i o n s t a n d s may even prove t o be r e s i s t a n t t o t h e I d e v a s t a t i n g Phytophthora r o o t r o t . Evidence from o t h e r commercially i m p o r t a n t s p e c i e s such a s t h e avocado (from r e s e a r c h conducted by Dr. Zentmire of U.C. R i v e r s i d e f i d e Ron K e l l y ) suggest t h a t i s o l a t e d montane stock ( i n t h i s c a s e from C e n t r a l Mexico) '.is t h e most r o o t r o t r e s i s t a n t form y e t d i s c o v e r e d . Regardless o f p o s s i b l e g e n e t i c d i f f e r e n c e s from t h e main c o a s t a l p o p u l a t i o n , s t a n d s such a s a t Cedar Basin r e p r e s e n t e a s i l y p r e s e r v e d and c o n t r o l l e d s i t e s which could be p r o t e c t e d from r o o t r o t i n f e s t a t i o n i f t h e proper p r e c a u t i o n s a r e t a k e n . For a review of t h e n e c e s s a r y p r e c a u t i o n s see K l i e junas and Adams (1980) . According t o r e c e n t s t u d i e s ( K l i e j u n a s and Adams 1980, Zoebel and Hawk 1980) and testamony from o t h e r worke?s ( i n l i t . ) , t h e non-native Phytophthosa - lateralis h a s f a n d w i l l probably c o n t i n u e t o s p r e a d r a p i d l y from Oregon ( v i a r o o t s t o c k imported t o B r i t i s h Col-umbia from F r a n c e , i n 1923) througho u t t h e range of POC. I t i n e v i t a b l y k i l l s b o t h l a r g e o l d growth trees and s a p l i n g s b e f o r e t h e y r e a c h marketable s i z e . It is easily s p r e a d from d r a i n a g e t o daainage i n mud t r a n s p o r t e d on t i r e s and f e n d e r s o f cars and t r u c k s . F o r t u n a t e l y a t t h i s time t h e r o o t rot h a s appar- e n t l y n o t spread t o any p a r t of t h e i n l a n d p o p u l a t i o n . Clearly, a w e l l c o n t r o l l e d and g e o g r a p h i c a l l y kell z a n t a i n e d a r e a i s necessary f o r t h e p r e s e r v a t i o n of u n i n f e c t e d t r e e s . An a r e a such a s Cedar Basin a t t h e head of a r i v e r d r a i n a g e i n a n u n i n f e c t e d r e g i o n , w i t h o n l y one l i t t l e - t r a v e l e d r o a d l e a d i n g i n t o t h e a r e a would be e a s i l y managed f o r r o o t r o t e x c l u s i o n . With t h e c u r r e n t v a l u e of POC over are $3000/MBF t h e r e d e f i n i t e economic a s w e l l a s s c i e n t i f i c and e s t h e t i c j u s t i f i a a t i o n s f o r m a i n t a i n i n g h e a l t h y p o p u l a t i o n s of t h i s s p e c i e s . There may be h o r t i c u l t u r a l a s w e l l a s s i l v i c u l t u r a l r e b e a r c h v a l u e s f o r t h e h i g h e l e v a t i o n i n l a n d p o p u l a t i o n of POC. Chamaecyparis lawsoniana i s one of t h e most widely p l a n t e d ornamental c o n i f e r s i n t h e North Temperate Zone. B a i l e y (1978) s t a t e s t h a t o v e r 8 0 h o r t - i c u l t u r a l " v a r i e t i e s " have been developed i n Europe and North America. Many o r a l l of t h e s e have o r i g i n a t e d from t h e main c o a s t a l p o p u l a t i o n and l i t t l e o r no b r e e d i n g r e s e a r c h on t h e i n l a n d p o p u l a t i o n h a s been conducted. Again, w i t h t h e d i s t i n c t c l i m a t e and l o n g i s o l a t i o n of t h e i n l a n d s t a n d s , i t i s l i k e l y t h a t d i f f e r e n t g e n e t i c t o l e r a n c e s t o environment a s w e l l a s p o s s i b l e morphological d i f f e r e n c e s i n f o l i a g e and s t a t u r e , may s e r v e t o i n c r e a s e t h e s p e c i e s ' o r n i m e n t a l value. From a more e c o l o g i c a l s t a n d p o i n t , t h e Cedar Basin POC i s unique with r e g a r d t o i t s a s s o c i a t e d f l o r a . Only i n t h i s and perhaps i n one o r two o t h e r high e l e v a t i o n b a s i n s i n t h e T r i n i t y Mountains, does POC a s s o c i a t e s o c l o s e l y w i t h t y p i c a l high montane t r e e s such a s Abies -magnif i c a , Pinus c o n t o r t a , P. m o n t i c o l a , and Tsuga mertensiana - . In some a r e a s a s i n t h e T e r r a c e Lake c i r q u e , a l l f o u r of t h e s e s p e c i e s may co-occur with C . lawsoniana ( s e e v e g e t a t i o n s e c t i o n ) . Hawk (1980) s t a t e POC i s remarkable Zobel and f o r i t s r e s t r i c t e d geographic range y e t i t s extreme t o l e r a n c e of a wide range of h a b i t a t s , spanning f o u r v e g e t a t i o n zones as d e f i n e d by F r a n k l i n and Dyrness ( 1 9 7 3 ) . A c t u a l l y , t h e Cedar Basin s t a n d s c l e a r l y e x t e n d i n t o a f i f t h vegetat i o n zone ( t h e Abies magnifica s h a s t e n s i s zone) and m a r g i n a l l y i n t o a s i x t h ( t h e Tsuga m e r t e n s i a n a zone) . A t t h e s e h i g h e l e v a t i o n s i n t h e c o l d n o ~ t h - f a u i n g Terrace Lake cirque, POC may t r a d e dominance p a r t i c u l a r l y with Tsuga mertensiana. T. - m e r t e n s i a n a , l i k e POC, p r e f e r s m o i s t , c o o l ground, and abave Terr a c e Lake t h i s i s provided by a v a l l e y bottom which channels runoff from snowpacks l i n g e r i n g i n t o l a t e J u l y and .not by permanent s t r e a m s and s e e p s which i s so o f t e n t h e h a b i t a t 6.5.;: POC i n o t h e r p a r t s of t h e basin. Hence, POC can be c o n s i d e r e d t o run t h e e n t i r e Eange of major c o n i f e r o u s f o r e s t t y p e s i n t h e Klamath Province, from the lowland c o a s t a l w e s t e r n hemlock and S i t k a spruce-dominated f o r e s t s , through Douglas f i r and mixed c o n i f e r f o r e s t , up i n t o t h e white f i r , red f i r , and even mountain hemlock-dominated f o r e s t s . There i s no o t h e r c o n i f e r o u s tree i n t h i s r e g i o n t h a t a s s o c i a t e s with more major f o r e s t types. However, o n l y i n t h e v i c i n i t y of t h e Cedar Basin c a n d i d a t e RNA i s POC known t o co-occur w i t h t h e most t y p i c a l dominant t r e e of t h e s u b a l p i n e zone i n t h e P a c i f i c Northwest (T. - mertensiana) and t h e most characterksjikc (A. - t r e e of t h e C a l i f o r n i a Canadian Zone magnifica). Endemic and Unusual Khamath and T r i n i t y Mountains F l o r a : The f l o r a of Cedar Basin s h a r e s t h e m a j o r i t y of i t s p l a n t s o t h e r mountain r e g i o n s of C a l i f o r n i a . However, with a b o u t 1 0 % of it@ f l o r a i s e i t h e r endemic t o t h e Klamath Province o r h a s a wider d i s t r i b u t i o n ( u s u a l l y n o r t h w e s t e r n North American), b u t i s known i n C a l i f o r n i a o n l y from t h e Klamath r e g i o n . The most s p e c i f i c group of endemic p l a n k s i n c l u d e two t a x a which s o far a s i s known, o n l y occur i n Cedar Basin. Both of t h e s e taxa w e r e d i s c b v e r e d d u r i n g t h e f i e l d work f o r t h h s r e p o r t . A r c t o s t a p h y l o s r k l a m a t h e n s i s ( ~ d w a r d s ,Keeler-Wolf , and Knight, 1983, attached ) i s a very d i s t i n c t i v e s p e c i e s o f manzanita known o n l y from t h e Terrace Lake b a s i n . The s p e c i e s i s a gray-green, low p r o s t r a t e p l a n t which l o c a l l y dominates t h e s h r u b l a y e r of t h e open r e d f i r f o r e s t and mountain c h a p a r r a l ( s e e F i g . 1 and Appendix 3 ) . g e n e r a l form it resembles i t a , A. nevadensis. t h e common and widespread pine-mat manzan- However, i n f l o r e s c e n c e form, l e a f c o l o r and g l a n d u l a r pubescence i n d i c a t e i t i s probably more r e l a t e d t o a n o t h e r low e l e v a t i o n Klamath endemic, A. k n i g h t i i , and a widespread chaparA r a l s p e c i e s , A. - viscida. I t s r e s t r i c t e d o c c u r r e n c e h e r e may have been t h e r e s u l t of p a s t h y b r i d i z a t i o n d u r i n g a d i f f e r e n t c l i m a t e i n t h e P l e i s t o c e n e when i t s p a r e n t forms occupied t h e same r e g i o n . The s p e c i e s i s d i s t i n c t i v e enough s o it i s u n l i k e l y t o occur more widel y i n o t h e r mountains of t h e Klamath Province (where b o t a n i c a l c o l l e c t i n g h a s been r e l a t i v e l y t h o ~ o u g h ) . I t should be c o n s i d e r e d very r a r e and endangered ( s e n s u Powell, 1974) by v i r t u e of i t s extremely l i m i t e d d i s t r i b u t i o n and s m a l l p o p u l a t i o n s i z e . The o t h e r taxon a p p a r e n t l y r e s t r i c t e d t o Cedar Basin i s a showy s p e c i e s of Penstemon, which a s y e t h a s been undescribed. Again, t h i s p l a n t i s l o c a l l y q u i t e common on t h e rock o u t c r o p s and s u n n i e r and d r i e r c l i f f s above T e r r a c e and Upper C l i f f Lakes. The s p e c i e s s h a r e s t h e pubescent a n t h e r s and rock-hugging h a b i t of two montane s p e c i e s known from t h e Klarnath r e g i o n , P. r u p i c o l a and P. d a v i d s o n i i ( n e i t h e r of which o c c u r s i n t h e b a s i n ) . The g l a c o u s , s e r r a t e l e a v e s resemble t h e f o l i a g e of P, r u p i c o l a , b u t t h e b l u e (and n o t magenta) f l o w e r s more c l o s e l y resemble P. d a v i d s o n i i (Fig. 2 ) . The u n i f o r m i t y of c h a r a c t e r i s t i c s t h r o u g h o u t t h e p o p u l a t i o n i n Cedar Basin and t h e absence of t h e s p e c i e s it most c l o s e l y resembles s u g g e s t t h a t it i s indeed a s e p a r a t e g e n e t i c e n t i t y , and n o t a s i m p l e r e c e n t h y b r i d . S p e c i f i c d e t e r m i n a t i o n w i l l have t o w a i t u n t i l t h e summer of 1983 when more m a t e r i a l c a n be c o l l e c t e d . A s w i t h t h e manzanita, t h i s s p e c i e s - i s probably r a r e enough by v i r t u e of i t s l i m i t e d range, t o :; FIGURE 1: The endemic Arctostaphy l o s klamathensis, dominant shrub of the open red f i r forest S W of Terrace L a k e F I G U R E 2: Penstemon sp. nov. a common p l a n t of gabbro outcrops S. of Upper C l i f f Lake. be c o n s i d e r e d v e r y r a r e and endangered. In a d d i t i o n t o t h e above two newly d i s c o v e r e d s p e c i e s which most c e r t a i n l y d e s e r v e a d d i t i o n a l s t u d y and p r o t e c t i o n , two o t h e r s p e c i e s known o n l y from t h e Northern T r i n i t y Mountains and a d j a c e n t S c o t t Mountains oco.ur i n t h e b a s i n . These a r e upi in us c r o c e u s (on t h e CNPS r a r e l i s t Powell, 1974) a yellow flowered herbaceoud. p e r e n n i a l l u p i n e r e s t r i c t e d t o r a t h e r d r y u l t r a m a f i c s o i l , and Mimulus primuloide s - - s s p . l i n e a r i f o l i u s , a yellow flowered bog and meadow i n h a b i t i n g monkeyflower. S e v e r a l more widespread Klamath Province endemics a l s o occur l o c a l l y and a r e on t h e CNPS l i s t s . These i n c l u d e L i l i u m washingtonianum v a r . purpurascens, C a s t i l l e j a arachnoidea, Lewisia l e a n a , and Charnaecyparis lawsoniana, i t s e l f . Other Rlamath endemics Bhd w i d e s ~ r e a ds p e c i e s known i n C a l i f o r n i a only. from t h e .Klamaths a r e - noted i n Appendix -1. Unusual Bog S p e c i e s : S e v e r a l widespread s p e c i e s on t h e CNPS r a r e l i s t a l s o occur i n the basin. and bogs. A l l of t h e s p e c i e s i n h a b i t m o i s t a r e a s such a s rocky s e e p s Two a r e a t t r a c t i v e and conspicuous p l a n t s of t h e bog and l a k e community a t Cedar and Lower C l i f f Lake: Drosera r o t u n d i f o l i a . . and Menyanthes t r i f o l i a t a ( F i g . 3 ) . Darlingtonia c a l i f o r n i c a ( C a l i f . p i t c h e r p l a n t ) an unusual and r a t h e r s c a r c e endemic t o t h e s e r p e n t i n e bogs of Northern C a l i f o r n i a and C o a s t a l Oregon i s a l s o common, p a r t i c u l a r l y a t Cedar Lake. Carex g i g a s , a n inconspicuous sedge of t h e m o i s t c i r q u e w a l l s e e p s above Terrace Lake i s c o n s i d e r e d t o be a r a r e s p e c i e s b o t h by Munz (1959) and Powell ( 4 9 7 4 ) . S e v e r a l o t h e r meadow and bog s p e c i e s a t Cedar Lake a r e unusual; Narthecium c a l i f o r n i c u m , T o f i e l d i a g l u t i n o s a , and S c h o e n o l i r i o n album a r e a l l p r i m i t i v e members of t h e L i l g f a m i l y endemic t o t h e C a l i f o r n i a region. FIGURE 3 : Drosera r o t u n d i f o l i a , Carex s p . , and K a L m i a c o l o n i z i n g submerged log i n Cedar Lake. Menyan--t h e ~i n shallow water ,beyond. F I G U R E 4 : Meadow f r i n g e a t Cedar Lake w i t h S i s y r i n c h ium idahoense and Pedicularis a t t o l e n s . .Two o t h e r n o t a b l e monocots a r e t h e a t t r a c t i v e and uncommon S i s y r i n chiurns, S. Elmeri (mountain golden-eyed g r a s s , f i g . 4 ) and S. idahoense, a glaucous-leaved p a l e b l u e flowered s p e c i e s . The uncommon and b e a u t i f u l g e n t i a n s Gentiana newberryi and G-amarella a l s o i n h a b i t t h e meadowy r i m around Cedar Lake. - - High D i v e r s i t y o f E r i c a c e a e : The The d e n s i t y and d i v e r s i t y of t h i s f a m i l y i n t h e b a s i n has been commented upon by s e v e r a l v i s i t o r s ( G . Keator, W. Rodrick, i n lit. ) . P e r s o n a l l y , I have never s e e n such a h i g h d i v e r s i t y of genera i n t h e E r i c a c e a e i n such a s m a l l a r e a . Nine genera and 1 3 s p e c i e s of e r i c a d s a r e known from t h e a r e a (see Appendix 1 ) . Many of t h e s e seem t o be f a r i n g w e l l i n t h e b a s i n because of t h e r e q u i r e d e d a p h i c c o n d i t i o n s . Leucothoe, G a u l t h e r i a , Kalmia, Ledwn, Rhododendron and some Vaccinium s p e c i e s a r e r e s t r i c t e d t o m o i s t more o r l e s s boggy a r e a s and s e v e r a l o t h e r genera (Cassiope, Phyllodoce and c e r t a i n A r c t o s t a p h y ~ o s , e . g . A . . k l a m a t h e n s i s ) may be l o c a l l y s u c c e s s f u l i n p a r t because of t h e i r t o l e r a n c e of t h e h a r s h u l t r a m a f i c s o i l . This may be p a r t i c u l a r l y '. t r u e of Cassiope and Phyllodoce, both of which axe u s u a l l y found a t s u b s t a n t i a l l y higher elevations. Zoological ~ u s t iifc a t i o n : A s c a l e d photo survey of animal t r a c k s was made by Ron K e l l y i n March and A p r i l 1980.. Subsequent i d e n t i f i c a t i o n by D r . W. Z. L i d i c k e r of t h e Museum of V e r t i b r a t e Zoology, U. C. Berkeley and Dr. Marshall White Dept. o f F o r e s t r y , U.C. Berkeley proved l a r g e d i v e r s i t y of c a r n i v o r e s i n h a b i t t h e b a s i n . that a Martens, b o b c a t s , mountain l i o n s , and b l a c k b e a r s l e f t f o o t p r i n t s i n t h e snow, a s d i d most l i k e l y f i s h e r s and wolverines. ticularly r a r e i n California. The l a s t two s p e c i e s a r e p a r - The wolverine i s e s p e c i a l l y r a r e w i t h 16 t h e o&kp r e c e n t r e p o r t s coming from t h e h i g h Northern S i e r r a and t h e Rlamath Mountains ( I n g l e s 1965, and M. White i n l i t . ). I n a d d i t i o n , two uncommon and s e n s i t i v e s p e c i e s of b i r d s , t h e s p o t t e d owl and p i l e a t e d woodpecker r e g u l a r l y u s e t h e f o r e s t e d a r e a s of t h e b a s i n . Both s p e c i e s p r e f e r o l d growth and overmature f o r e s t s and c e r t a i n l y f i n d s u i t a b l e h a b i t a t i n t h e c e d a r groves and t h e mixed c o n i f e r f o r e s t s o f t h e lower b a s i n . Maintanence of t h e b a s i n i n i t s l e a s t d i s t u r b e d s t a t e i s t h e most i m p o r t a n t f a c t o r i n i n s u r i n g t h e continued r e s i d e n c e of both t h e c a r n i v o r e s and b i r d s . PHYSICAL FEATURES -Geomorphology and Topography: Cedar Basin l i e s a t t h e head o f t h e South Fork of t h e Sacramento River. The b a s i n i s o r i e n t e d t o t h e n o r t h e a s t and h a s had a l l of i t s major f e a t u r e s shaped by g l a c i a t i o n i n t h e P l e i s t o c e n e . The b a s i n was dominated by two g l a c i e r s i n t h e l a s t i c e age; one which s p i l l e d down t h e s l o p e s s o u t h e a s t of Cedar Lake and a l a r g e r one t h a t flowed from hanging g l a c i e r s beneath t h e h i g h r i d g e s s o u t h of C l i f f and T e r r a c e l a k e s and converged w i t h t h e Cedar Lake g l a c i e r below Lower C l i f f Lake ( F i g s . 5 & 6 1. Cedar Basin l i e s w i t h i n an area of t h e Northern T r i n i t y Mtns. known a s "The Eddjzs."' The o r i g i n of t h e name i s u n c l e a r , b u t t h e s w i r l i n g p a t t e r n of r i d g e s and smoothed, g l a c i a l v a l l e y s s e e n i n an a e r i a l view does s u g g e s t t h e shapes of e d d i e s i n a r i v e r . The l a t e r a l moraine t h a t formed between t h e Gumboot Lake g l a c i e r t o t h e n o r t h and t h e Cedar Basin g l a c i e r forms t h e n o r t h w e s t e r n boundary of t h e RNA. The low l a t e r a l moraine s e p a r a t i n g t h e Cedar l a k e and C l i f f Lake g l a c i e r s i s c r o s s e d i n t h e open mountain c h a p a r r a l on t h e jeep road between t h e two l a k e s . FIGUREI 5: Upper Cliff, Cliff, and Lower Cliff lakes showing path of Cliff Lake glacier. Note t h e U-shaped valley of South Fork of Sacramento River beyond. From Lower Cliff Lake looking SW to glacially carved cirque holding Upper Cliff Lake (in clouds) FIGURE 6 : . The e f f e c t of g l a c i a t i o n c a n c l e a r l y be s e e n i n t h e p o l i s h e d gabbro on t h e bench n o r t h of Upper C l i f f Lake and 1 / 2 m i l e southwest of Cedar Lake. glaciation. A l l t h e l a k e s i n t h e b a s i n owe t h e i r o r i g i n s t o The s t e e p c l i f f f a c e behind C l i f f Lake r i s e s from t h e lake a t ca. 5800ft. t o 7149ft. i n l e s s than 1 / 2 m i l e . In contrast, t h e v a l l e y f l o o r from Cedar Lake t o t h e n o r t h w e s t boundary, a d i s t a n c e of n e a r l y a m i l e o n l y d r o p s from c a . 5750 t o 5 4 2 0 f t . Geology : A s was mentioned, Cedar Basin l i e s w i t h i n a huge a r e a of u l t r a - mafic r o c k s and i s e n t i r e l y u n d e r l a i n by them. Gabbro, a m s a l t and pepper" c o l o r e d rock with a consist.ency s i m i l a r t o g r a n i t e i s t h e I : dominant t y p e throughout t h e b a s i n . The b o u l d e r s a l o n g t h e o u t l e t t o Lower C l i f f Lake and t h e v e r t i c a l cliffs behind C l i f f and Terrace l a k e s a r e a l l made of gabbro. o u t c r o p s of o t h e r r o c k s . Within t h e g a b b r o i c mass a r e s m a l l Boulders of a p a r t l y s e r p e n t i n i z e d c r y s t a l - l i n e t a l c a r e o c c a s i o n a l a s a r e d i k e s of a n d e s i t e from on&-->to four f e e t thick. The o n l y o t h e r r o c k s i n t h e b a s i n occur n o r t h w e s t of Cedar Lake and i t s o u t l e t s t r e a m and a r e v a r i o u s l y a l t e r e d forms of p e r i d o t i t e (serpentinite). The n o r t h e a s t e r n boundary a r e a c o n s i s t s of v a r i o u s s i z e s of rounded s o c k s and b o u l d e r s d e p o s i t e d from t h e main r i d g e t o t h e w e s t i n a l a t e r a l moraine r i s i n g up from t h e northwestern s h o r e of Cedar L a k e . This u l t r a r n a f f c t e r s a n e i s composed of o l d r o c k s (Ordovician t o Carboniferous, I r w i n , 1 9 6 6 ) which e r u p t e d on o r below t h e sea floor. These r o c k s changed i n composition w i t h t h e a d d i t i o n of s e a water ( t o form s e r p e n t i n i t e ) o r o r i g i n a t e d from magma from t h e b a s a l t i c mantle of t h e e a r t h , w h i c h s o l i d i f i e d slowly w e l l under t h e s u r f a c e (to form gabbro) . Both o f t h e s e r o c k s s h a r e many chemical c h a r a c t e r i s t i c s , ' reflecting t h e i r s i m i l a r o r i g i n s beneath t h e ocean f l o o r o f f t h e s h o r e s of c o n t i n e n t s . S i m i l a r aged u l t r a - mafic r o c k s occur s o u t h e a s t i n t h e western S i e r r a , and f u r t h e r n o r t h e a s t i n t h e Blue and Wallowa mountains of N o r t h e a s t Oregon. of t h e s e mountains are thought t o be r e l a t e d All , and may have once formed a continuous mountain range on t h e w e s t e r n edge of North Ameri c a ( A l t and Hyndman 1 9 7 5 , 1 9 7 8 ) . Gabbroic r o c k s produce few economically i m p o r t a n t m i n e r a l s , and t h e small amount of s e r p e n t i n i t e i s u n l i k e l y t o c o n t a i n any more t h a n small t r a c e s of n i c k e l , chromium, a s b e s t o s , t a l c , c i n n a b a r , and o t h e r economic d e p o s i t s known from t h e Klamath u l t r a m a f i c t e r r a n e . The m i n e r a l c o n t e n t of t h e b a s i n ' s rocks p r o v i d e l i t t l e more of r e a l v a l u e t h a n a s e v e r e and i n t e r e s t i n g e d a p h i c environment i n f l u e n c i n g t h e s t r u c t u r e and composition of t h e l o c a l f l o r a . Climate : Cedar Basin i s under t h e sake c l i m a t i c regime a s most of Cali f o r n i a , b u t r e c e i v e s s l i g h t l y m o r e - e c e c - i p i t a t i o n i n t h e summer and w i n t e r t h a n many mountainous a r e a s t o t h e s o u t h ( S i e r r a , North Coast Range, and T ~ a n S v a r s eRanges). P r e c i p i t a t i o n a v e r a g e s between 70 and 80 i n c h e s a n n u a l l y (Kahrl, 1 9 7 9 ) , most of which f a l l s as snow i n the winter. Kahrl ( 1 9 7 9 ) shows t h e a v e r a g e A p r i l 1 snow depth f o r t h e Cedar Basin a r e a t o be over 100 i n c h e s , a s h i g h a s a n y a r e a i n t h e Klamath Province. Snow accumulates a t h i g h e l e v a t i o n f o r e s t e d as s i t e s t o a t l e a s t an average of 180 i n c h e s on t h e n o r t h e a s t s l o p e s F of Terrace Lake b a s i n , and commonly t o 1 4 4 i n c h e s a t o t h e r upper e l e v a t i o n f o r e s t e d s i t e s (evidence from h e i g h t o f l i c h e n growth on trees). I n c o n t r a s t , a t t h e lower e l e v a t i o n s on s o u t h e a s t f a c i n g s l o p e s snow r a r e l y a v e r a g e s deeper t h a n 36 i n c h e s . The c o l d , shaded upper slopes of the basin thus provide a distinctly different microclimate,conducive to the dominance of such trees as nlountain hernlock,from the lower elevation, sunny , quickly drying slopes covered with open mixed conifer forest. VEGETATION Shallow Lakes and Ponds: - Eight permanent or semipermanent bodies of water exist in the basin. They range from relatively deep and rockbound Cliff Lake to small, shallow ponds only 30 feet across. With the exception of CliEE Lakefall lakes and ponds in the basin have an obvious zone of aquatic vegetation inhabiting the shallower water areas. The best developed open water communities occur at Cedar and Lower Cliff lakes (Fig.7 ) because of their relatively constant water level, large size, and uniformly shallow depths. . The following species are either submerged, emergent, or anchored floating-leaved hydrophytes characteristic of the basin: Isoetes occidentalis Sparganium angusti~olium Isoetes bolanderi Scirpus validus Nuphar polysepalum Heleocharis montividensis Menyanthes trifoliata Potarnogeton natans var. parishii Dulichium axundinaceum Carex rostrata Boa and Meadow: At Cedar Basin there are no extensive wet meadowy areas. Apart from narrow, discontinuous fringing areas with characteristic meadow vegetation bordering Cedar and Lower Cliff lakes and several unnamed ponds, the majority of the permanently moist non-forest habitat in the basin is best classified as bog. The bog community around Cedar Lake, and to a lesser extent at tot. 874 Lower C l i f f Lake c o n s i s t s o f both r a i s e d , hummocky a r e a s topped l a r g e l y w i t h e r i c a c h a u s s h r u b s , and lower mucky a r e a s dominated by members of t h e Cyperaceae and such s p e c i e s a s D a r l i n g t o n i a .". c a l i f o r n i c a . (Fig . . 8 ) . Both t g p e s a r e u n d e r l a i n by t h i c k l a y e r s of p a k t i a l l y decomposed v e g e t a t i o n t h a t may f e e l shaky u n d e r f o o t . However, the r a i s e d a r e a s a r e b e t t e r d r a i n e d and s u p p o r t a n e a r l y c o n t i n u o u s shrub l a y e r of KaLmia, Ledum, Vaccinium o c c i d e n t a l e , and Spiraea douglasii. The hummocks form a n a t u r a l b a r r i e r between t h e open l a k e community and t h e low bog and meadow v e g e t a t i o n ( F i g . 9 ) . These hummocks may have o r i g i n a t e d from e i t h e r f l o a t i n g l o g s l o d g i n g near t h e s h o r e of t h e lake o r emergent b o u l d e r s and r o c k s . . On some younger f l o a t i n g l o g s and on t h e edges of hummocks grow t h i c k clumps of Brosera, small i n d i v i d u a l s of D a r l i n g t o n i a , T o l f i e l d i a , Narthecium, and sedges such a s Carex buxbaumii, and C. a q u a t a l i s . The uncommon, d i m u n i t i v e G a u l t h e r i a humifusa a l s o c r e e p s over emergent r o c k s and Logs i n t h i s zone, and small t r e e s of POC a r e f r e q u e n t l y i n t e r s p e r s e d among t h e shrubs. The low, mucky bog i d dominated by members of t h e sedge f a m i l y i n c l u d i n g ; S c i r p u s micPocarpus, Heleocharis montkvidensis, Carex ormantha, C. i n t e g r a , and C. - buxbaumii. A s t e r a l p i g e n u s s s p . a n d e r s o n i i grows i n t h i s zone a s does S c h o e n o l i r i o n album. I n s l i g h t l y more e l e v a t e d m o i s t a r e a s D a r l i n g t o n i a becomes dominant and o f t e n forms dense s t a n d s f u r t h e r away from t h e l a k e a s s o c i a t i n g w i t h POC and Ledum ( ~ i g , h J ) . Where more s o i l h a s developed such a s n e a r t h e i n l e t streams t o b o t h Cedar and Lower C l i f f l a k e s , herbaceous meadow v e g e t a t i o n p r e dominates i n c l u d i n g : Botrychium simplex Hypericurn a n a q a l l o i d e s S i d a l c e a oregana s s p . s p i c a t a Polyqonum b i s t o r t o i d e s Viola maclaskeyi Dodecatheon alpinurn s s p . rnaius F i g u r e 7:Lower C l i f f Lake w/ Nuphar ,Potamogeton, and o t h e r a q u a t i c s i n forground; f r i n g e o f POC and t r a n s i t i o n a l mixed conifer-red f i r f o r e s t i n background. F i g u r e 8 :Bog v e g e t a t i o n around Cedar Lake+&/ l o w mucky t y p e on l e f t , s l i g h t l y r a i s e d type (wy D a r l i n g t o n i a ) , and hummock w/ POC and K a l m i a i n backgkound. F i g u r e 9: Cedar Lake w/ hummock b o r d e r , % a k ~ v e g e t a t i o n ( i n d i s t - (: a n c e ) , mucky bog, and meadow fringe ( i n for- . ground) . Gentiana -- newberryi Sisyrinchium idahoense Gentians a m a r e l l a S. elrneri d Mimulus primuloides -- Juncus nevadensis Pedicularis attolens J. orthophyllus Parnassia p a l u s t r i s var. c a l i f o r n i c a Carex hoodi P o t e n t i l l a g r a c i l i s ssp. n u t t a l l i i C. - Lotus p i n n a t u s C. r a y n o l d s i i Epilobium l a c t i f l ~ r u m Deschamwsia c a e s ~ i t o s a P e r i d e r i d i a oregana A -g r o s t i s i d a h o e n s i s Belenium h o o p e s i i Gliceria s t r i a t a Aster occidentalis G -. T r i t e l i a hyacinthina Calarnagrostis c a n a d e n s i s laeviculmis elata The boy community a t Lower C l i f f Lake i s n o t a s w e l l developed a s a t Cedar Lake. F r i n g i n g v e g e t a t i o n a t Lower C l i f f included, Leucothoe, Ledum, S p i r a e a d o u q l a s i i , and Rosa p i s o c a r p a . However, ~ a l r n i a i s r a r e and Vaccinium o c c i d e n t a l e i s a p p a r e n t l y a b s e n t . Only a t t h e l a k e ' s s o u t h e r n end a r e t h e m f a i r l y l a r g e p a t c h e s of Drosera and D a ~ l i n g t o n i a . Other l a k e s and ponds i n t h e b a s i n do n o t s u p p o r t f r i n g i n g bog v e g e t a t i o n l a r g e l y because of a more widely f l u c t u a t i n g water t a b l e . P o r t Orford Cedar F o r e s t : I n Cedar Basin t h i s f o r e s t type (most s i m i l a r t o S A F t y p e 231) i s c l e a r l y a n e d a p h i c climax community r e l i a n t upon permanent moisture. P o r t Orford c e d a r groves f r i n g e Cedar, T e r r a c e , C l i f f , and Lower C l i f f l a k e s and f o l l o w t h e i r o u t l e t s t r e a m s a l l t h e way down t h e banks of t h e South Fork o f t h e Sackamento River. Groves a l s o o c c u r above t h e s e l a k e s and streams a t s e e p s , a l o n g i n t e r m i t t e n t r i v u l e t s , and s t r e a m s up t o n e a r l y 6400ft. The h i g h e s t e l e v a t i o n FIGURE 10: Edge of bottomland POC f o r e s t with u n d e r s t o r y of bog species; Darlingtonia c a l i f o r n i c a , Ledum glandulosum predominating. FIGURE 11: Bottomland POC f o r e s t South of Lower C l i f f Lake showing t h e many f a l l e n trees t y p i c a l of t h i s f.orest type. c e d a r i s a n i n d i v i d u a l c a . 30 f t . t a l l growing between t h e two t a r n s i n t h e upper T e r r a c e Lake b a s i n a t c a . 6350 f t . Because POC t e n d s t o t h r i v e i n t h e s e m o i s t s i t u a t i o n s throughout the b a s i n , the mountain a l d e r and willow-dominated h i g h mountain r i p a r i a n community t y p i c a l i n s i m i l a r s i t u a t i o n s throughout much POC i s q u i t e shade t o l e r a n t of C a l i f o r n i a does n o t o c c u r h e r e . (much more s o t h a n t h e l i g h t - l o v i n g a l d e r s and w i l l o w s ) and t e n d s t o exclude a l l o t h e r trees and l a r g e s h r u b s from t h e r i p a r i a n habitat available i n the basin. T h e r e a r e two r e d i l y d i s c e r n a b l e t y p e s of POC f o r e s t a t Cedar Basin. One o c c u r s a l o n g rocky s t r e a m s and l a k e s i d e s , where t h e surrounqing d ~ a i n a g ei s good and t h e m o i s t u r e i s r e s t r i c t e d t o w i t h i n a few f e e t of s u r 5 a c e w a t e r . The o t h e r i s i n r a t h e r f l a t bottomland a s around t h e western and s o u t h e r n shores of Cedar Lake, n e a r t h e i n l e t t o Lower C l i f f Lake, and n e a r t h e confluence of t h e Cedar and Lower C l i f f Lake o u t l e t s t r e a m s . Both t y p e s a r e s t r o n g l y dominated by POC (see T a b l e s 1,2,&3 and Appendix 3 ) , b u t t h e unders t o r y v e g e t a t i o n aD$ &he f o r e s t s t r u c t u r e d i f f e r s . The bottomland POC f o r e s t h a s t h e h i g h e s t d e n s i t y s t a n d s of any forest i n t h e b a s i n w i t h o f t e n between 4 0 i n height i n lOOm 2 & 5 0 t r e e s over 6 f t . (1600-210O/acre,see Table .I). The trees a r e o f t e n very s h a l l o w l y r o o t e d , and because of t h e s a t u r a t e d muddy, sandy, ax p e a t y s o i l o f t e n are uprooted. P a s s i n g through such groves means climbing over and under many f a l l e n t r u n k s ( ~ i g . 1 1 ) . Many r e c e n t l y f a l l e n trees ( a p p a r e n t l y r e s u l t i n g from heavy 1 9 8 1 s n o w f a l l ) have p o o l s o f water f i l l i n g t h e i r newly formed b a s a l c r a t e r s , a t t e s t i n g t o t h e high water t a b l e . Sample p l o t s 1,2, and 3 w e r e i n t h i s t y p e of f o r e s t around Cedar Lake. This type o f t e n l i e s immediately back from t h e bog and meadow s u r r o u n d i n g t h e l a k e s . TABLE 1 Tree d e n s i t y of Chamaecyparf s lawsoniana i n five lOxlOm p l o t s . Plot # M t . Al. POC L.P. D.F. I.C. W.F. WWP J.P. ...M.H. R.F. TABLE 2 S a p l i n g and s e e d l i n g d e n s i t y of C. - lawsoniana on lOxlOm p l o t s . Plot # M t . A l . POC L.P. D.F. I.C. W.F. WWP J.P. M.H. R.F. TABLE 3 Basal area ( s q . inches) of C. - lawsoniana t r e e s on lOxlOm p l o t s . Plot # Mt.Al. POC L.P. D.F. I.C. W.F. WWP J.P. M.H. R.F. Total Because d r a i n a g e i s slow and t h e s o i l s a t u r a t e d j u s t below the surface,many of t h e more shade t o l e r a n t bog and meadow s p e c i e s a r e s h a r e d , i n c l u d i n g Ledwn, G a u l t h e r i a , Leucothoe, and D a r l i n g t o n i a . However, t h e r e i s a c o r e of s p e c i e s which i s more o r l e s s r e s t r i c t e d t o t h e u n d e r s t o r y o f t h i s bottomland f o r e s t and t h e y i n c l u d e : Botrychium multifidum ssp. s i l a i f o l i u m Physocarpus c a p i t a t u s Athyrium f i l i x - f e m i n a Alnus t e n u i f o l i a Caltha howellii Cornus s t o l o n i f e r a Viola g l a b e l l a Linnaea b o r e a l i s s s p - l o n g i f o l i a V. - adunca L i l i u m kelleyanum P y r o l a secunda Allium validum P r u n e l l a v u l g a r i s ssp. l a n c e o l a t a Habenaria s ~ a r s i f l o r a Listera convallarioides Much of t h e bottomland f o r e s t around Cedar Lake a p p e a r s t o have s u f f e r e d f i r e damage perhaps 100 y e a r s ago. Many b a r e s n a g s , c h a r r e d a t t h e b a s e , s t i l l s t a n d ( F i g . 1 2 ) and no l i v i n g t r e e s l a r g e r t h a n ca. 2 2 i n c h e s dbh can be found. The l a r g e s t l i v i n g POC measured i n t h e bottomland t y p e f o r e s t was 4 8 " j u s t n o r t h o f Lower C l i f f Lake. The growth r a t e s of POC a p p e a r t o be t h e most r a p i d of any c o n i f e r i n t h e basin. This c e r t a i n l y h a s much t o do w i t h t h e c o n t i n u a l water supply a f f o r d e d t o most i n d i v i d u a l s . Table 4 compares t h e ages of b r e a s t h e i g h t s a p l i n g s of v a r i o u s s p e c i e s sampled i n t h e v i c i n i t y of Cedar Lake. The age of a 2 8 i n c h d i a m e t e r POC stump n e a r .bhe o u t l e t of Lower C l i f f Lake was 2 6 0 y e a r s . Ron Kelly r e p o r t s a stump measuring 4 2 i n c h e s i n d i a m e t e r and a n age of c a . 4 0 0 y e a r s n e a r t h e n o r t h e a s t boundary. The l a r g e s t i n d i v i d u a l POC measures ca. 5 2 i n c h e s dbh and i s probably 4 5 0 - 5 0 0 y e a r s o l d . grows a l o n g t h e o u t l e t s t r e a m of Lower Cliff Lake d r a i n e d s t r e a m and l a k e s i d e t y p e of POC f o r e s t . his i n d i v i d u a l i n the .well- V i e w S toward peak 7 1 4 9 from s e r p e n h i n i t e mixed conifer forest on moraine NW of Cedar Lake. Charred snags of POC along W s i d e o f Cedar Lake i n middle ground. FIGURE 12: TABLE 4 Ages of breast height saplings sampled near Cedar Lake. Species #1 - lawsoniana C. - jeffreyi P. A. concolor A - magnifica A. T. - mertensiana 56 P. - menziesii - 36 #2 #3 Sample p l o t s 4 and 5 a r e r e p r e s e n t a t i v e o f t h e s t r e a m s i d e rocky l a k e s h o r e type o f POC f o r e s t . The d e n s i t i e s of trees a r e n o t a b l y less, y e t b a s a l a r e a i s g r e a t e r t h a n o n t h e sampled bottomland p l o t s (Table 3 ) . This i s perhaps because fewer t r e e s c a n g e t e s t a b l i s h e d a l o n g t h e narrow, a f t e n s t e e p s i d e d , and v a r i a b l e watercourses. N e v e r t h e l e s s , t h e few t h a t can a r e a b l e t o s i n k t h e i r r o o t s down t o a permanent m o i s t u r e supply and perhaps grow with l e s s c o m p e t i t i o n for l i g h t and .space t h a n t h e bottomland i n d i v i d u a l s . The s t r e a m s i d e and rocky l a k e s i d e POC f o r e s t , a l t h o u g h r a r e l y extend(Fig. 13) ing more t h a n 5m from t h e w a t e r ' s e d g e , . h a s by f a r t h e h i g h e s t b a s a l a r e a of any f o r e s t t y p e sampled i n t h e b a s i n ( s e e Table I S ! , , up t o 2 3501in. 2/100m2 ( t h e e q u i v a a e n t of 984 f t / a c r e ) The u n d e r s t o r y . . s p e c i e s i n t h i s subtype a r e t y p i c a l of o t h e r m o i s t f o r e s t t y p e s found i n the b a s i n and i n c l u d e ; Leucothoe d a v i s a e , Goodyera o b l b n g i f o l i a , Chdmaphila uibellata , Pyrola p i c t a , Vacciniurn a r b u s c u l a , P t e r i d i u m aquilinum, and P e d i c u l a r i s semibarbata. - Mixed Conifer F o r e s t : Two t y p e s of mixed c o n i f e r f o r e s t (SAF t y p e 2 4 3 w i t h v a r i a t i o n s ) a l s o occur i n t h e b a s i n . One i s r e s t r i c t e d t o t h e s e r p e n t i n i t e mor- a i n e on t h e northwest s i d e of t h e a r e a , and t h e o t h e r o c c u r s on gabbro a t t h e lower e l e v a t i o n s i n t h e b a s i n . The s e r p e n t i n e t y p e i s an open f o r e s t w i t h dominance being t r a d e d between s e v e r a l s p e c i e s i n c l u d i n g ; Pseudotsuga m e n z i e s i i , - l a m b e r t i a n a . Calocedrus Abies c o n c o l o r , Pinus j e f f r e y i , and P. d e c u r r e n s and Pinus monticola a l s o commonly o c c u r i n t h i s f o r e s t P. c o n t o r t a murrayana o c c u r l o c a l l y . t y p e , and Pinus pondexosa and - The u n d e r s t o r y i s dominated by t y p i c a l mountain c h a p a r r a l s h r u b s - A -. p a t u l a i n c l u d i n g ; Quercus v a c c i n i f o l i a , A r c t o s t a p h y l o s nevadensis, , TABLE 5 Comparative s t a t i s t i c s f o r t h e t h r e e sampled f o r e s t types. Plat # av. p l o t e l e v . (feet) Basal A r e a ( ft 2 / a c r e ) height1 (ft.1 Rees/acre Snow Depth (feet) 1 e s t i m a t e d from r a n g e f i n d e r measurements of t h r e e t a l l e s t t r e e s 'estimated f r o m average distance f r o m ground t o edge of l i c h e n growth on three randomly chosen trees 2 Ceanothus p r o s t r a t u s , and Amalanchier p a l l i d a . Two o t h e r s h r u b s , Rhamnus c a l i f o r n i c u s s s p . o c c i d e n t a l i s and Ribes r o e z l i i appear t o be l o c a l l y r e s t r i c t e d t o t h i s f o r e s t t y p e . F i v e 5Ox20m ( 0 - l h a ) p l o t s w e r e sampled i n t h i s s u b t y p e . The r e s u l t s a r e p r e s e n t e d i n Tables 6 , 7 , and 8. The g e n e r a l exposure of t h e e n t i r e s e r p e n t i n i t e subtype i s s o u t h e a s t . the nutrient-poor, be very open. T h i s , coupled w i t h well drained substrate causes the f o r e s t t o o&ten Compared w i t h t h e o t h e r two sampled t y p e s o f f o r e s t i n t h e b a s i n , t h i s f o r e s t a v e r a g e s both t h e l o w e s t d e n s i t y of t r e e s and t h e s m a l l e s t b a s a l a r e a ( s e e Table 5 ) . Some i n d i c a t i o n of s h i f t - i n g dominance r e s u l t i n g from d i f f e r i n g m i c r o c l i m a t i c c o n d i t i o n s i s i n d i c a t e d by a s t r o n g e r tendency f o r P. j e f f r e y i dominate t h e d r i e s t , most s o u t h e r l y - f a c i n g s l o p e s and A. - c o n c o l o r , and i n c r e a s i n g l y - monticola t o dominate a t h i g h e r e l e v a t i o n s (above 6 0 0 0 f t . ) . P. P. - rnenziesii i s a l s o somewhat more common a t lower e l e v a t i o n s on t h i s subtype and may o c c a s i o n l y r e a c h s u r p r i s i n g l y l a r g e p r o p o r t i o n s (two t r e e s approachi.ng 60" dbh species seen i n t h e b a s i n ) . were measured and a r e t h e l a r g e s t o f any Conspiauous by i t s absence from t h i s t y p e i s Abies magnifica (even a t h i g h e r e l e v a t i o n s ) , a l t h o u g h t h i s s p e c i e s i s a common component of t h e gabbro mixed c o n i f e r subtype. The gabbro subtype d i f f e r s from t h e p r e v i o u s f o r e s t by i t s h i g h e r abundance of A. - magnifica and P. c o n t o r t a murrayana. A 1 1 o t h e r t r e e s p e c i e s noted on s e r p e n t i n i t e a l s o o c c u r , though P, - jeff r e y i and C -. d e c u r r e n s a r e of much lower importance. No v e g e t a t i o n sampling was done i n t h i s subtype, b u t both t h e s u b s t r a t e ( o f t e n w i t h a h i g h e r w a t e r t a b l e ) and t h e o v e r a l l n o r t h e r l y exposure of t h e lower e l e v a t i o n s o f t h e b a s i n appear t o i n f l u e n c e t h e d i f f e r e n t composition of t h i s s u b t y p e . The MCF f o r e s t on gabbro i s a l s o r a t h e r open, b u t n o t so much a s t h e s e r p e n t i n i t e t y p e . Understory s h r u b s a r e a l s o mostly t y p i c a l mountain c h a p a r r a l s p e c i e s , b u t Chrysolepkk TABLE 6 T r e e d e n s i t y on 5Ox20m Mixed C o n i f e r F o r e s t p l o t s . Plot # W.F. D.F. J.P. S.P. WWP L.P. I.C. TABLE 7 Sapling and seedling d e n s i t y o n 50x20111 Mixed Conifer F o r e s t p l o t s . plot # W.F. D.F. S.P. J.P. WWP L.P. I.C. TABLE 8 Basal area ( i n . 2 ) on Mixed C o n i f e r F o r e s t p l o t s . plot # W.F. 6 1032 D.F. 2356 S.P. J.P. W P L.P. I.C. 987 9 95 40 1158 total 5677 sempervirens, a s p e c i e s o f t e n c h a r a c t e r i s t i c o f open r e d f i r f o r e s t , appears l a r g e l y r e s t r i c t e d t o t h i s type. This f o r e s t shows a g r a d u a l c l i n a l s h i f t with i n c r e a s e d e l e v a t i o n toward t h e composition of r e d fir-mountain hemlock f o r e s t . I t s a f f i n i t y t o t h i s type can be recognized even a t lower e l e v a t i o n s by o c c a s i o n a l t r e e s of Tsuga mertensiana - and u n d e r s t o r y s h r u b s such a s Leucothoe d a v i s i a e and Vaccinium a r b u s c u l a i n more mesic a r e a s . Herbs and subshrubs a r e r a t h e r widely s c a t t e r e d i n both subt y p e s of MCF. A few such a s Lupinus c s o c e u s , Eriophyllum lanatum v a r . lanceolatum, C.onvolvulus malacophyllus, Lotus c r a s s i f o l i u s , Angelica c a l i f o r n i c a , Galium bibreale, Cirsium - a n d e r s o n i i , and Iris p u r d y i ,appear l o c a l l y r e s t r i c t e d t o t h e s e r p e n t i n i t e subtype. Others such a s : Xerophyllum t e n a x Phacelia sp. B e r b e r i s nervosa Cynoglossum o c c i d e n t a l e Linum perenne ssp. l e w i s i i P e d i c u l a r i s semibarbata Viola PurPurea P. - S i l e n e lemmonii Monardella o d o r a t i s s i m a s s p p a l l i d a racemosa Polvaonum d o u a l a s i i Solidago -c a l i f o r n i c a Frasera a l b i c a u l i s Senecio a r o n i c o i d e s Ipomopsis a g g r e q a t a Eupatoxium o c c i d e n t a l e Agoseris s p . Habenaria u n a l a s c e n s i s & Elymus g l a u c u s a r e c e n t e r e d i n t h e MCF zone i n t h e b a s i n , o c c u r r i n g more commonly on t h e gabbro subtype. L i s t s of a l l h e r b s and s h r u b s o c c u r r i n g on each of t h e 15 sample p l o t s c a n be found i n Appendix 3 . Re,d Fir-Mountain Hemlock F o r e s t : - This f o r e s t (most s i m i l a r t o SAF t y p e '207) i s t h e dominant t y p e on a l l n o r t h e r l y exposures a t h i g h e r e l e v a t i o n s i n t h e b a s i n . Five 5Ox20m ( 0 . l h a ) v e g e t a t i o n p l o t s were chosen i n t h i s f o r e s t t y p e and t h e r e s u l t s p r e s e n t e d i n Tables 9 , 10, and 11 and Appendix 3. There i s a l a r g e degree of v a r i a t i o n i n dominance and c o v e r i n t h i s t y p e . T h i s ranges from dense, c l o s e d s t a n d s dominated by r e d f i r ( e . g . p l o t 13 and Fig. 1 4 ) on n o r t h e a s t and northwest f a c i n g s l o p e s , t o c o l d , mesic v a l l e y bottom s t a n d s dominated ( o r co-dominated) by T. - mertensi a n a and o c c a s i o n a l l y P. - c o n t o r t a murrayana ( p l o t s 11 & 15, F i g . 151, t o open, r a t h e r s c e n e s c e n t r e d f i r f o r e s t ( p l o t 14), and very open - magnifica, P.montiaola, f o r e s t co-dominated by A. (plot 1 2 , and Fig.1) and P -. c o n t o r t a . The u n d e r s t o r y i s dominated i n open f o r e s t s with s h a l l o w s o i l by s h r u b s of t h e mountain c h a p a r r a l i n c l u d i n g Quercus v a c c i n i f o l i a , A r c t o s t a p h y l o s klarnathensis p rostratus. - ,A. nevadensis, A -. p a t u l a , and Ceanothus The endemic A. k l a m a t h e n s i s i s l o c a l l y a v e r y i m p o r t a n t member of t h i s t y p e , c o v e r i n g more t h a n 6 0 % of t h e g&ound i n some areas. I n t h e more c l o s e d canopy, shady s t a n d s t h e u n d e r s t o r y i s s p a r s e with pyrolaceous and s a p r o p h y t i c s p e c i e s such a s Pyrola p i c t a , Ch&maphila u m b e l l a t a r and C o r a l l o r h i z a maculata predominating. I n mesic s t a n d s Leucothoe, Vaccinium a r b u s c u l a , V. - scoparium, Anemone q ui nq u e f o l i a v a r . minor and P y r o l a secunda a r e o f t e n common. Even on t h e most mesic s i t e s w i t h i n t h i s f o r e s t tree s i z e and b a s a l a r e a coverage are n o t as high as i n many p a r t s of t h e S i e r r a Nevada r e d f i r f o r e s t . The g r e a t e s t b a s a l a r e a sampled i n Cedar Basin ( p l o t 11, 383 f t . 2/ a c r e ) i s l e s s t h a n % a s g r e a t a s r e d f i r f o r e s t sampled a t t h e M t . P l e a s a n t c a n d i d a t e RNA (Keeler-Wolf and Keeler-Wolf, 1981) o r t h e Onion Creek c a n d i d a t e RNA (Talby, 1 9 7 7 ) , FIGURE 13: Typical narrow r i p a r i a n s t a n d of POC a l o n g o u t l e t s t r e a m f r o m Terrace Lake. Red f i r dominates o n l y a few meters away t o the r i g h t . F I G U R E 14: Closed red firmountain hemlock f o r e s t on N E f a c i n g s l o p e above T e r r a c e Lake. Note sparse u n d e r s t o r y and l i c h e n growth i n d i c a t i n g average w i n t e r snow depth of ca. 180 ". TABLE 9 Tree d e n s i t y on 5 0 x 2 0 m r e d fir-mountain hemlock p l o t s . plot # W.F. R.F. M.H. L.P. WWP POC TABLE 10 S a p l i n g and s e e d l i n g density on 5 O x 2 0 m red f i r - m o u n t a i n h e m l o c k p l o t s . plot # W.F. R.F. M.H. L.P. WWP POC TABLE 11 Basal a r e a (sq. i n . ) on 5 O x 2 0 m r e d fir-mountain hemlock p l o t & . plot # W.F R.F. M.H. L.P. WWP POC total i n t y p i c a l S i e r r a red f i r f o r e s t . The d i f f e r e n c e i n s t a t u r e i s a l s o Many r e d f i r s a t M t . P l e a s a n t r e a c h 1 7 5 f t . i n h e i g h t substantial. and t h e average canopy h e i g h t i s ca. 1 3 8 f t . Average canopy h e i g h t i n t h e analagous f o r e s t a t Cedar Basin i s o n l y 8 3 f t . and t h e t a l l e s t f i r measured, o n l y 1 2 0 f t . DBH o f Cedar Basin r e d f i r . i s s i m i l a r l y r e - duced w i t h t h e l a r g e s t measured being 45.5" a s opposed t o many a t Mt. P l e a s a n t over 60"! These d i f f e r e n c e s i n d e n s i t y and s t a t u r e can probably b e s t be a t t r i b u t e d t o t h e s o . i l . t y p e . Gabbro u n d e r l i e s t h e l o c a l f o r e s t and i s a chemically d e f i c i e n t and h a r s h e r s u b s t r a t e t h a n t h e g r a n i t e u n d e r l y i n g t h e Mt. P l e a s a n t a r e a . Tolerance of u l t r a m a f i c s o i l s by A. rnagnifica has n o t been t e s t e d t o my knowledge. A However, i t i s probable t h a t l i k e Pseudotsuga (Zobel and Hawk, 1 9 8 0 ) , it e x h i b i t s reduced v i g o r on u l t r a m a f i c s o i l s . .The abundance of Tsuga mertensiana a t such r e l a t i v e l y low e l e v a t i o n s ( l o c a l l y dominant t o c a . 6050ft.), on t h e o t h e r hand, may r e s u l t from t h e p o t e n t i a l , f o r t h i s s p e c i e s t o t o l e r a t e t h e poor s o i l m a r a . e f f e c t i u e l y t h a n A. - magni f i c a . The r e l a t i v e l y open n a t u r e of t h i s and t h e mixed c o n i f e r f o r e s t s i n Cedar Basin i s c l e a r l y n o t t h e r e s u l t o f an overmature and s c e n e s c e n t tree crop. Regeneration of a l l t r e e s p e c i e s i s r e l a t i v e l y good a s i n d i c a t e d by a t y p i c a l l y uneven age c l a s s d i s t r i b u t i o n ( s e e Appendix 3) Mountain Chaparral : T h i s shrub dominated community c o v e r s l a r g e a r e a s of shallow, rocky s o i l i n t h e b a s i n . It is especially noticeable i n the eastern p o r t i o n where it e x t e n d s a c r o s s almost t h e e n t i r e west-facing s l o p e . The a r e a dominated by t h i s v e g e t a t i o n may a c t u a l l y be s u b s t a n t i a l l y l a r g e r t h a n t h e l i s t e d f i g u r e because of t h e many s m a l l unmappable pockets w i t h i n t h e openings of mixed c o n i f e r and r e d fir-mountain hemlock f o r e s t . Three s p e c i e s ; Q u e r c u s v a c c i n i f o l i a , A r c t o s taphy- l o u s n e v a d e n s i s , and A. - p a t u l a are t h e most w i d e s p r e a d and i m p o r t a n t p l a n t s of t h e b a s i n ' s mountain c h a p a r r a l . u f a r l y a b u n d a n t , d o m i n a t i n g i n most a r e a s . Q. v a c c i n i f o l i a i s p a r t i c O t h e r widespread, import- anlh s h r u b s are t h e low m a t - l i k e Ceanothus p r o s t r a t u s , Amalanchier p a l l i d a , and Holodiscus b o u r s i e r i , t h e l a t t e r p a r t i c u l a r l y abundant on s t a b l i z e d , . t a l u s a l o n g t h e r i d g e e a s t o f T e r r a c e and Lower C l i f f lakes. Prunus e m a r g i n a t a , Cdanothus v e l u t i n u s , and C h r y s o l e p i s s e m p e r v i r e n s a r e less common and more l o c a l s h r u b s o f t h i s community. The endemic A r c t o s t a p h y l o s k l a m a t h e n s i s i s a l o c a l dominant o f t h i s community w e s t and s o u t h w e s t o f T e r r a c e Lake. Many h e r b s i n t h i s community a r e a l s o s h a r e d w i t h t h e more open, x e r i c rock o u t c r o p community. Some o f t h e most common and c h a r a c t e r - i s t i c species include: Cheilanthes qracillima Penstemon d e u s t u s Onychium densum P. Streptanthus tortuosus P. - newberryi s s p . b e r r y i Arabis r e c t i s s i m a Castilleja arachnoidea A rabis platysperma Sedum o b t u s a t u m s s p . b o r e a l e Arenaria n u t t a l l i i ssp. qxegaria - Lornatium macrocarpurn A -. c o n q e s t a Kellogia g a l i o i d e s C a l y p t r i d i u m umbellatum Haplopappus g r e e n e i Erioqonum umbellatum v a r . umbellatum Chsysothamnus n a u s e o s u s a l b i c a u l i s Apbcynum pumilum Antennaria rosea Phlox d i f f u sa Stephanomeria l a c t u c i n a Nama l o b b i i Juncus p a r r y i Mimulus l a y n e a e Calamagrostis k o e l e r i o i d e s azuraus The l o c a l mountain c h a p a r r a l a p p e a r s t o be l a r g e l y an e d a p h i c climax. Unlike many a r e a s i n ~ a l i f o r n i a ,p a s t f i r e h i s t o r y d o e s n o t a f f e c t i h s presence o r d i s t r i b u t i o n a s much a s t h e shallow, rocky s o i l and s l o p e exposure. Large i s o l a t e d t r e e s w i t h o u t f i r e s c a r s on t h e west-facing s l o p e above Terrace Lake (Fig.16 ) and i n o t h e r a r e a s a t t e s t t o t h e permanence and r e l a t i v e l y f i r e - f r e e h i s t o r y of t h e surrounding mountain c h a p a r r a l . The widespread l o c a l presence o f t h i s community may r e f l e c t t h e t o l e r a n c e many of i t s c o n s t i t u e n t s have of extremely w e l l d r a i n e d , rocky u l t r a m a f i c s o i l . Rock O u t c r o ~ : On t h e s t e e p rock f a c e s s o u t h of C l i f f Lake and a t t h e head of t h e Terrace Lake c i r q u e , s e v e r a l species of rock-loving p l a n t s o c c u r i n a- unique community. The rock o u t c r o p environment v a r i e s g r e a t l y from x e r i c , exposed gabbro s l a b s t o shaded s e e p s n e s t l e d w i t h i n deep c r e v i c e s . (Fi'gs. 17 ., & 1.81. The f l o r a of t h e s e o u t c r o p s can t h u s c o n v e n i e n t l y be d i v i d e d i n t o x e r i c and rnesic components. Many of t h e dry-adapted s p e c i e s a r e s h a r e d w i t h t h e mountain c h a p a r r a l and a r e p r e v i o u s l y l i s t e d . : However, a few s p e c i e s appear r e s t r i c t e d t o a c t u a l e x t e n s i v e rock o u t c r o p s . Notable among t h e s e i s t h e l o c a l endemic penstemon, which i s a c t u a l l y one o f t h e most common p l a n t s i n i t s l i m i t e d h a b i t a t . Other c h a r a c t e r i s t i c s p e c i e s i n c l u d e Lewisia l e a n a , S i t a n i o n h y s t r i x , and S t i p a columbiana. T h e mesic phase o f t h e rock o u t c r o p community shares some s p e c i e s with t h e bogs and meadows, b u t t h e mkist-to-wet, y e t rocky (and u s u a l l y shady) environment pr0duce.s a unique s i t u a t i o n . Several h e r b s and s h r u b s a p p e a r l o c a l l y r e s t r i c t e d t o t h i s h a b i t a t : Cryptogramma a c r o s t i c h o i d e s -- Athyrium a l p e s t r e Adianturn pedatum v a r . a l e u t i c u m Delph,inium depauperatum Polystichum I o n c h i t i s Actaea r u b r a s s p . a r g u t a Polygonurn d a v i s a e Phyllodoce e m p e t r i f o r m i s South shore of Terrace Lake with dense forest dominated by mountain hemlock, lodgepole p i n e , and red f i r . Port Orford c e d a r and western w h i t e p i n e are 81.~0 :common. FIGURE 15: FIGURE 1 6 : Mountain chaparral east of T e r r a c e Lake showing l a r g e isolated Jeffrey p i n e and White fir. FIGURE IF: T y p i c a l x e r i c rock o u t c r o p a t head of Upper C l i f f Lake cirque. Red fir-mountain hemlock f o r e s t t o t h e l e f t . M t . Shasta i n d i s t a n c e . FIGURE 1 8 : Romanzoffia s i t c h e n s i s , a r a r e s p e c i e s of shaded rock c r e v i c e s , known i n C a l i f o r n i a o n l y from the Klamath Province. Cassiope m e r t e n s i a n a S i b b a l d i a procumbens Nemophila p a r v i f l o r a v a r . a u s t i n a e Sorbus s c o p u l i n a Epilobium a n g u s t i f o l i u m sitchensis Bovkinia maior Rhamnus p u r s h i a n a - Saxifraga ferruginea Acer glabrum v a r S. bryophora Arnica d i v e r s i f o l i a Heuchera p r i n g l e i Antennaria a l p i n a v a r . media Ribes nevadense Disporum h o o k e r i d . torreyi Spiraea densiflora Carex s p e c t a b i l i s Luetkea p e c t i n a t a . BOUNDARIES I n g e n e r a l , t h e o r i g i n a l boundaries d e l i m i t e d i n t h e Oat. 1 9 8 0 RNA p r o p o s a l a r e a c c e p t a b l e . The t o p o g r a p h i c b o r d e r s on t h e n o r t h - west, t h e s o u t h , and t h e e a s t a r e i n keeping w i t h t h e maintenance of t h e n a t u r a l i n t e g r i t y of t h e b a s i n . The owners of t h e c a . 1 2 0 a c r e s of p r i v a t e l a n d w i t h i n t h e b a s i n have a s t r o n g d e s i r e t o m a i n t a i n t h e i r land, i n p a r t i c u l a r t h e area a d j a c e n t t o t h e proposed RNA, i n a n a t u r a l s t a t e (Ron Kelly p e r s . com.). b u f f e r a r e a would seem n e c e s s a r y . Consequently, no The a r t i f i c i a l n o r t h e a s t e r n bound- a r y a c r o s s t h e basin f l o o r c o i n c i d e s w i t h t h e edge of t h e proposed D e v i l ' s Zoo timber s a l e , and r e p r e s e n t s t h e o n l y m o d i f i c a t i o n from t h e o r i g i n a l proposal. O r i g i n a l l y , a more a r c u a t e n o r t h e a s t boundary w a s drawn ( b e f o r e the e x t e n t of t h e timber s a l e was d e t e r m i n e d ) . The p r e s e n t s t r a i g h t - l i n e boundary was dr.awn w i t h t h e i n t e n t i o n of i n c l u d i n g a s mich a s p o s s i b l e of t h e bottomland POC f o r e s t and a d j a c e n t mixed c o n i f e r f o r e s t n e a r t h e edge of t h e timber s a l e . IMPACTS The major p o t e n t i a l and r e a l impacts on t h e Cedar Basin c a n d i d a t e RNA i n c l u d e camping, logging, and woodcutting p r e s s u r e , continued v e h i c l e u s e of road 39NO5Y, and t h e t h r e a t of c e d a r r o o t r o t i n f e s t i n g t h e l o c a l POC groves. The e x i s t i n g road i n t o t h e b a s i n has been i n e x i s t e n c e f o r many y e a r s (perhaps 5 0 o r more). A s evidenced by i t s poor c o n d i t i o n i t i s n o t much used e x c e p t by high-clearance four-wheel d r i v e - t y p e vehicles. The impact of v i s i t o r s a t Cedar Lake, t h e most a c c e s s i b l e a t t r a c t i o n i n t h e b a s i n , h a s been s u r p r i s i n g l y l i g h t o v e r t h e y e a r s . Although f o u r l o n g - e s t a b l i s h e d c a m p s i t e s e x i s t a t t h e l a k e , a l l a r e on t h e e a s t e r n s i d e away from t h e immediate s h o r e w i t h i t s s e n s i t i v e boggy hummocks and meadow f r i n g e . The western edges of t h e l a k e , which house t h e most e x t e n s i v e and d i v e r s e bog and meadow v e g e t a t i o n , are v e r y l i g h t l y e f f e c t e d , w i t h o u t even a noticreable f o o t p a t h . This l i g h t u s e i s probably a r e s u l t of t h e shallowness of t h e l a k e , making it u n l i k e l y t o s u p p o r t a l a r g e and a t t r a c t i v e ( t o fishermen) t r o u t population. S i m i l a r l y , t h e s i n g l e l a r g e c a m p s i t e a t Lower C l i f f Lake i s away f r o m t h e s h o r e on t h e n o r t h s i d e . L i g h t l y used p a t h s l e a d a l o n g t h e western and e a s t e r n edge of Lower C l i f f Lake, b u t t h e dense f r i n g i n g s t a n d s of cedar and t h e s a t u r a t e d boggy a r e a s on t h e e a s t e r n and s o u t h e r n s i d e s have n o t been n o t i u e a b l y d i s turbed. The l o n g - e s t a b l i s h e d campsite a l o n g t h e e a s t e r n s i d e o f C l i f f Lake i s a t t h e edge of t h e r a t h e r u n s e n s i t i v e mountain chaparr a l formation. The fring&..;of POC has n o t been s t r o n g l y e f f e c t e d , and because no bog o r meadow v e g e t a t i o n surrounds t h e lake,impacts on e x i s t i n g l a k e s h o r e v e g e t a t i o n a r e minimal. i n 1 9 8 2 were r e l a t i v e l y free of l i t t e r . . A l l campsites v i s i t e d No w e l l e s t a b l i s h e d campsites e x i s t a t any o t h e r p a r t s of t h e b a s i n . The p a s t e f f e c t s of l o g g i n g and woodcutting a r e most n o t i c e a b l e a l o n g t h e road t o Cedar Lake. S e l e c t i v e l o g g i n g o c c u r r e d many y e a r s ago i n t h e s e r p e n t i n i t e MCF n o r t h w e s t of Cedar Lake and on o t h e r p a r t s of t h e lower b a s i n . s c a t t e r e d (3-5/0. l h a ) . Stumps a t l e a s t 50 y e a r s o l d a r e widely Skid t r a i l s and o t h e r r e s i d u a l e f f e c t s a r e not generally noticeable. One l o g g i n g s p u r i s s t i l l a p p a r e n t a n the northwest s i d e o f Cedar Lake, though overgrown w i t h s a p l i n g s . Woodcutting u s e h a s i n c r e a s e d i n r e c e n t y e a r s i n t h e lower p a r t s of t h e b a s i n . Downed and dead s t a n d i n g t r e e s of POC and o t h e r s p e c i e s have been takeri, p a r t i c u l a r l y a l o n g t h e road t o Cedar Lake and around the lake, i t s e l f . Apparently, some l i v e t r e e s have been cut a s w e l l . A t p r e s e n t , t h e lower b a s i n h a s been posted a s a no woodcutting a r e a . However, chainsaws were heard on F o r e s t S e r v i c e l a n d w i t h i n t h e b a s i n i n mid-September. The timber v a l u e o f t h e a c c e s s i b l e f o r e s t e d p a r t s o f t h e b a s i n i s r e l a t i v e l y low, e x c e p t f o r t h e POC s t a n d s . The open, somewhat s t u n t e d n a t u r e of both t h e mixed c o n i f e r and r e d fir-mountain hemlock f o r e s t s suggests again t h a t t h e i r value i s higher f o r research than f o r marketable saw l o g s . Although t h e poor c o n d i t i o n of t h e road i n s u r e s , t o some d e g r e e i t s l i g h t u s e , t h e r e i s some evidence of more r e c e n t i n c r e a s e d impact. A few a r e a s show t h e e f f e c t s of d e t o u r i n g w i t h subsequent e r o s i o n and d e s t r u c t i o n of s o i l and surrounding v e g e t a t i o n . p a r t i c u l a r l y a p p a r e n t about + mile This i s n o r t h e a s t of Cedar Lake. Without q u e s t i o n t h e most t h r e a t e n i n g b a s i n i s t h e p o t e n t i a l s p r e a d of c e d a r r o o t r o t . impact on t h e Needless t o s a y , t h e value of t h e b a s i n a s an RNA would be s e r i o u s l y diminished i f t h e l o c a l c e d a r s were i n f e c t e d w i t h t h i s d i s e a s e . RECOMMENDATIONS I n my o p i n i o n , t h e most parsimonious way t o d e a l w i t h a l l of t h e above impacts i s t o g a t e t h e Cedar Basin road n e a r t o where it branches from road 40N37. Access by motor v e h i c l e s could t h u s be r e s t r i c t e d , b u t n o t precluded. T h i s i s n e c e s s a r y because t h e Kellys and p a r t i c u l a r l y t h e Dobsons r e q u i r e road a c c e s s t o t h e i r p r i v a t e c a b i n s a t C l i f f Lake. Also v e h i c l e a c c e s s could be g i v e n t o r e s e a r c h - e r s o r o t h e r p a r t i e s i n t e r e s t e d i n t h e n a t u r a l f e a t u r e s of t h e proposed RNA . Management problems w i t h g a t e s i n c l u d e vandalism and t h e l o g i s - t i c s of d i s t r i b u t i n g keys o r combinations t o t h e p r o p e r p e r s o n n e l . A g a t e might most p r u d e n t l y be p l a c e d a t a s p o t where c r o s s - c o u n t r y a c c e s s around t h e g a t e i s d i f f i c u l t . Perhaps t h e most l i k e l y s i t e i s a l o n g t h e Cedar asi in r o a d a b o u t 200 y a r d s from t h e main road where downhill s l o p e s on t h e moraine a r e s t e e p e r , and a cut-bank b o r d e r s t h e i n s i d e of t h e road. The i n i t i a l expense and maintenance of t h e g a t e r e q u i r e s c a r e f u l c o n s i d e r a t i o n . Being an owner of p r i v a t e l a n d i n t h e mountains, I am w e l l aware of t h e forms of vandalism and d e s t r u c t i o n t a k e n upon s i m i l a r locked g a t e s . Everything i s used; from ramming w i t h t r u c k s , t o b o l t c u t t e r s , p o r t a b l e c u t t i n g t o r c h e s , and dynamite. Thus, p r e c a u t i o n s such a s concealed, guarded padlocks, heavy gage s t e e l p o s t s and b a r s , and a s u b s t a n t i a l c o n c r e t e foundation f o r t h e posts should be t a k e n when p u t t i n g up t h e g a t e . Keys o r combinations could be d i s t r i b u t e d t o t h e p r i v a t e l a n d owners and could be h e l d a t t h e M t . Shasta Ranger S t a t i o n f o r o t h e r i n t e r e s t e d p a r t i e s arranging.' i n advance to w e . t h e area. Perhaps maintenance c o s t s could be d e f r a y e d by a s m a l l e n t r a n c e f e e ( s i m i l a r t o a campground f e e ) charged t o a l l u s e c s . LITERATURE CITED Alt, D. and D. Hyndman. 1975. Roadside Geology of Northern C a l i f o r n i a . Mountain P r e s s . Alt, D. and D . Hyndrnan. 1978. Roadside Geology of Oregon. Mountain Press. 1978. The C u l t i v a t e d C o n i f e r s i n North America. B a i l e y , L.H. Allen- h e l d Osmun/Universe Books. Davis, G.A. 1966. Metamorphic and g r a n i t i c h i s t o r y of t h e Klamath mountains. I n E . B a i l e y {ed. ) Geology o f Northern C a l i f o r n i - a , C a l i f . D i v i s i o n of Mines B u l l . 190. D i l l e r , J.S. U.S. 1 9 0 2 . Topographic development o f t h e Klamath Mountains. Geol. Survey B u l l . 196. Edwards, S . , Keeler-Wolf, T. W. Knight. 1983. A r c t o s t a p h y l o s klarnath- e n s i s , a new s p e c i e s of manzanita from S i s k i y o u County, C a l i f ornia. The Four Seasons 4 ( 6 ) : Franklin, J.F. and C.T. and Washington. Griffin, J.R. and W.B. Dyrness. 1973. Natural v e g e t a t i o n o f Oregon U.S. C r i t c h f i e l d . 1 9 7 2 . The d i s t r i b u t i o n of f o r e s t trees i n California. I n g l e s , L. 1965. F o r e s t S e r v i c e Gen. Tech. Rep. PNW-8. U.S. For. Serv. Res. Pap. PSW-82. Mammals of t h e P a c i f i c S t a t e s . S t a n f o r d . I r w i n , W.P. 1966. Geology of t h e Klamath Mountains Province. In E. B a i l e y ( e d . ) Geology of Northern C a l i f o r n i a . C a l i f . Div. Mines B u l l . 1 9 0 . Kahrl, W. (ed.) 1 9 7 9 . Keeler-Wolf, T. and V. t h e proposed M t . The C a l i f o r n i a Water A t l a s . S t a t e of C a l i f o r n i a . Keeler-Wolf . 1981. ~n e c o l o g i c a l survey of P l e a s a n t Research N a t u r a l Area, Plumas N a t i o n a l Forest, California. Klie junas, 5. and D. Adams Unpublished r e p o r t on f i l e a t PSW, Berkeley. . 1980. A n eva1uat:ion of Phytopthora r o o t r o t of P o r t Orford c e d a r i n C a l i f o r n i a . p e s t management r e p o r t 80-1. U.S. For. Serv. Munz, P. A. 1 9 5 9 . A C a l i f o r n i a F l o r a . U. Powell, R. 1974. of California. Raven, P. H. C. Press. Inventory of r a r e and endangered v a s c u l a r p l a n t s S p e c i a l P u b l i c a t i o n 1, C a l i f . Native P l a n t Soc. and D. A. A l e l r o d . 1978. O r i g i n and r e l a t i o n s h i p s of the California Flora. U. C. Pub. Bot. 72. T a l l e y , S. 1977. An e c o l o g i c a l survey of t h e Onion Creek c a n d i d a t e r e s e a r c h n a t u r a l area on t h e Tahoe N a t i o n a l F o r e s t . Unpublished r e p o r t on f i l e a t PSW Berkeley. Zobel, D . B,.. and G. M. Hawk. 1980. The environment of Chamaecyparis lawsoniana. Am. Midl. Nat.103:280-297. APPENDIX 1 VASCULAR PLANT LIST This list includes all species identified during my visits in August and September. Several additional species noted by other observers are also listed. Nomenc%ature for the most part is from Munz 1968. A California Flora and Supplement. The following smbols refer to habitat types: l=lake b&m=bog and meadow POC=Port Or ford cedar forest MCF=mixed conifer forest sf-mh=red fir-mountain hemlock forest mcmountain chaparral ro=sock outcrop (xeric) s&s=spring and seep (mesic rock outcrop) The symbol "KO" refers to plants known in California only from the Klamath Province. FERNS AND ALLIES: Botrychium multifidurn ssp. silaifolium POC B. simplex b&m Pteridium aqualinum var. pubescens POC, MCF Cheilanthes gracillima mc, ro Cryptogramma acrostichoides s&s, ro Onychium densum ro Adiantum pedatum var. aleuticum s&s Polystichum lonchitis s&s, ro P. lemmonii s & s , rot KO Cystopteris fragilis s&s AtQyrium filix-femina POC A. alpestre s&s CONIFERS Abies concolor MCF, rf-mh A. rnagnifica var. shastensis rf-mh, MCF A. nobilis MCF, rf-rnh (individuals with sometbut not all of the species characters) Abies l a s i o c a r p a ( r e p o r t e d by Keator) ? KO Pinus l a m b e r t i a n a MCF P. monticola MCF, rf-mh P. c o n t o r t a v a r . murrayana POC. MCF, rf-mh P. ponderosa MCF P. j e f f r e y i MCF Picea breweriana ( r e p o r t e d b$. Mary T a y l o r ) ? KO Tsuga m e r t e n s i a n a POC, MCF, r.f-mh Pseudotsuga menkigrsii,,MCF Calocedrus d e c u r r e n s MCF Chamaecyparis lawsoniana POC KO FLOWERING PLANTS Ranunculaceae C a l t h a h o w e l l i i POC, b&m Actaea r u b r a s s p . a r g u t a s L s Delphinium depauperatum s & s Anemone q u i n q u e f o l i a v a r . minor POC, MCF, rf-mh Aquilegia formosa s&s Berberidaceae B e r b e r i s nervosa MCF Nymphaceae Nuphar polysepalum 1 Malkaceae S i d a l c e a oregana s s p . s p i c a t a b&m Sasraceniaceae D a r l i n g t o n i a c a l i f o r n i c a b&m, POC Droseraceae Drosera r o t u n d i f o l i a b&m Linaceae Linum perenne s s p . l e w i s i i MCF Violaceae POC Viola g l a b e l l a V. purpurea MCF V. cuneata (Keator) MCF V. macloskeyi b&nI V. adunca b&m Bypericaceae Hypericum a n a g a l l o i d e s b & m Cruciferae S t r e p t a n t h u s t o r t u o s u s mc ~ h a l a s p iglaucum ( K e a t o r ) MCF Arabis r e c k i s s i m a m c A. platysperma mc, r o A. h o l b o e l l i i v a r . r e t r o f r a c t a MCF, m c Caryophyllaceae S t e l l a r i a sp. ? MCF Arenaria nuttallii ssp. gregaria mc, ro A. congesta mc, MCF, ro Silene lemmonii MCF Portulacaceae Lewisia leana ro, mc, KO Calyptridium umbellatum mc Polygonaceae Eriogonum umbellatum var. umbellatum Rumex angiocarpus b&m Polygonum bistortoides b&m P. douglasii MCF P. spergulariaeforma MCF P. davisiae s&s, ro ro, mc Primulaceae Dodecatheon alpinum ssp. majus b&m Ericaceae Ledum glandulosum var. californicum b&m, POC Rhododendron occidentale POC, MCF Kalmia polifolia var. microphylla b&m Phyllodoce empetriformis s&s, ro, KO Cassiope mertensiana s&s, ro Leucothoe davisiae POC, b&m, rf-mh Gaultheria humifusa b&m Arctostaphylos nevadensis MCF, rf-mh, mc,. A. patula mc, MCF, rf-mh A. mc, rf-mh Vaccinium occidentale b&m V, arbuscula POC, MCF, rf-mh V. scopariurn rf-mh KO Pyrolaceae Pyrola picta MCF, rf-mh Pyrola picta ssp. integra MCF, rf-mh P. secunda POC, rf-mh Chimaphila umbellata MCF, rf-mh Pterospora andromedea MCF, rf-mh Gentianaceae Gentiana newberryi b&m G. amarella b&m Menyanthes trifoliata 1 Frasera albicaulis MCF Apocynaceae Apocynum pumilum MCF, mc Convulvulaceae Convulvulus malacophyllus MCF Polemoniaceae Polemonium californicum rf-mh Phlox diffusa mc Microsteris gracilis MCF Ipomopsis aggregata MCF Linanthus ciliatus MCF Hydrophyllaceae Nemophylla parwiflora var. austinae s&s Phacelia sp. (Keator) ? Nama lobbii mc, ro Romanzoffia sitchensis s&s KO Boraginaceae Cynoglossum occidentale MCF Scrophulariaceae Mimulus primuloides ssp. linearifolius b&m M. layneae mc M. guttatus b&h, s&s Penstemon deustus mc, ro P. azureus mc, MCF P. parvulus mc P. laetus (Keator) ? P. newberryi ssp. berryi mc P. sp. nov.. a.0 KO Collinsia torreyi MCF, rf-mh Pedicularis semibarbata MCF, EOC P. attolens b&m P. racemosa MCF Castilleja arachnoidea mc KO C. miniata s&s KO Labiatae Scutellaria antirrhinoides MCF Prunella vulgaris ssp. lanceolata POC b&m Monardella odoratissima ssp. pallida MCF Cxassulaceae Sedum obtusatum ssp. boreale ro, mc, KO Saxifragaceae Parnassia palustris var. californica b&m Boykinia major s&s, POC Saxifraga bryophora s & s S. ferruginea s&s Mitella pentandra POC Heuchera merriami (Keator) ? Heuchera pringlei s&s, ro KO Ribes divaricatum var. klarnathense POC KO Ribes nevadense s&s Ribes roezlii MCF Rosaceae Physocarpus capitatus POC Spiraea douglasii b&m, POC S. densiflora s & s Luetkea pectinata s&s KO Holodiscus boursieri mc Ivesia sp. (Keator) ? Eotentilla glandulosa ssp. nevadensis MCF, b&m P. gracilis ssp. nuttallii b&m Fragafia platypetala POC Sibbaldia procumbens s&s Rubus leucodermis rf-mh R. parviflorus POC Rosa pisocarpa var. rivalis POC, MCF, b&m Prunus emarginata rnc Sorbus scopulina s & s Amelanchier pallida MCF, rf-mh, POC, mc Leguminosae Lupinus albicaulis MCF L. adsurgens ? MCF L. croceus MCF KO Trifolium longipes b & m , s&s T. sp. ? b&m Lotus crassifolius MCF L. pinnatus b&m L. oblangifolius b & m Betulaceae Alnus tenuifolia b&m, POC Fagaceae Chrysolepis sempervirens MCF, mc Quercus vaccinifolia MCE, rf-mh, mc Salicaceae Salix scouleriana POC, MCF Onagraceae Epilobium angustifolium s & s E. lactiflorum s&s, b&m Gayophytum nuttallii MCF G. humile MCF Rhamnaceae Rhamnus purshiana s&s R. californica ssp. occidentalis MCF Ceanothus velutinus mc Ceanothus prostratus mc, MCF, rf -mh Loranthaceae Arceuthobiwn campylopodum MCF Aceraceae Acer glabrum var. torreyi s&s Umbelliferae Sanicula nevadensis MCF Perideridia oregana b&m, s&s Ligusticum californicum s&s, POC Lomatium macrocarpum mc Angelica californica MCF Cornaceae Cornus stolonifera POC Rubiaceae Galium boreale MCF KO Kellogia galioides MCF, mc, rf-rnh Caprifoliaceae Linnaea borealis ssp. longiflora POC Symphoricarpos acutus POC, MCF ~oniceraconjugialis POC Compositae Madia minima MCF Helenium hoopesii bsm Eriophyllum lanatum var. lanceolatih MCF Haplopappus greenei mc, ro Solidago californica MCF Chrysothamnus nauseosus ssp. albicaulis MCF, mc Aster brickellioides var. glabratus MCF, ro, rf-mh, mc, A. occidentalis b&m A. alpigenus ssp. andersonii b&m Erigeron inornatus MCF, rf-mh Achillea lanulosa MCF, b&m Arnica discoidea var. elata MCF A. diversifolia s & s Senecio triangularis POC, b$m, s&s S. aronicoides MCF Antennaria alpina var. media s & s , ro A. rosea mc, ro Eupatorium occidentale MCF Cirsium andersonii MCF Microseris nutans b&m Agoseris sp. MCF Stephanomeria lactucina mc, ro Hieracium albiflorum MCF rf-mh H. cynoglossoides MCF, rf-mh H. bolanderi Potamogetonaceae Potamogeton natans 1 Liliaceae Xerophyllum tenax MCF Narthecium californicum b&m Tofieldia glutinosa ssp. occidentalis b&m Schoenolirion album b & m , POC Disporum hookeri s&s Zigadenus veneosus s&s, b&m Veratrum californicum POC, s & s , b&m Lilium washingtonianum var purpurescens MCF L. kelleyanum POC, s & s . Sparganiaceae Sparganium angustifolium 1 Amaryllidaceae Allium validum POC Tritelia hyacinthina b&m Iridaceae Iris purdyi MCF Sisyrinchiurn idahoense b&m KO KO Sisyrinchium elmeri b&m Orchidaaeae Habenaria unalascensis MCF H. dilatata var. leucostachys b&m H. sparsiflora s&s, POC Listera convallarioides POC Goodyera oblangifolia POC, MCF, rf-mh Corallorhiza maculata rf-mh Juncaceae Juncus parryi mc, ro, MCF 3. nevadensis b&m J. balticus b&m J. orthophyllus b&m Luzula divaricata rf-mh, mc Cyperaceae Scirpus microcarpus b&m S. validus 1 Heleocharis montividensis var. parishii 1, b&m Dulichium arundinaceum 1 Caxex rostrata 1, b&m C. hoodi b&m, s&s C. cjigas s & s C. spectabilis s&s C. integra b&m C. raynoldsii b&m C. buxbaumii b&m, s&s C. ormantha b&m C. laeviculmis b&m, s&s C. aquatalis b&rn C. leporinella b&m Grarninae Sitanion hystrix ro Deschampsia caespitosa b&m Poa canbyi ro Stipa columbiana ro Muhlenbergia filiforrnis ro, s&s Calamagrostis koelerioides ro, mc C. breweri b&m C. canadensis b&m, POC Danthonia intermedia MCF, ro Agrostis exarata b&m, MCF A. idahoensis b&m Glyceria striata b&m G. elata b&m, POC Elymus glaucus MCF APPENDIX 2 WRT'EBRATES KNOWN OR SUSPECTED FROM CEDAR BASIN FISH Brook Trout (Salvelinus fontina1is);in CliZE, Lower Cliff, Upper Cliff, and Terrace lakes Pacific Giant Salamander (Dicamptodon ensatus); known from headwaters of Sacramento River Rough-skinned newt (Taricha granulosa); seen in most lakes Ensatina (Ensatina eschscholtzi oregonensis); possibly occurs Black salamander (Aneides flavipunctatus) ; possibly occurs Western toad (Bufo - - boreas); seen around Cedar Lake Pacific treefrog (Hyla regilla) common around lakes & streams Red-legged frog (Rana aurora); seen in Cedar, Lower Cliff lakess.:: Sagebrush lizard (Sceloporus graciosus); seen in mtn. chaparral Western skink (Eumeces skiltonianus); possibly occurs No~thernalligator lizard (Gerrhonotus coeruleus shastensis); possibly occurs Rubber boa (Charina bottae) ; possibly occurs Ringneck snake (Diadolphus punctatus); possibly occurs Sharp4.tailed snake (Contia tenuis) ; possibly occurs Common garter snake (Thamnophis sirtalis fitchi); possibly occurs Mountain garter snake [Thamnophis e1egans);possibly occurs Oregon garter snake (Thamnophis couchi hydrophila); possible Racer (Coluber constrictor); possibly occurs Gopher snake (Pituophis melanoleucus); possibly occurs C a l i f o r n i a mountain k i n g s n a k e ( L a m p r o p e l t i s z o n a t a ) ; p o s s i b l e Western r a t t l e s n a k e ( C r o t a l u s v i r i d u s o r e g a n u s ) ; p o s s i b l e BIRDS T h i s list i n c l u d e s o n l y t h o s e s p e c i e s o b s e r v e d o r h e a r d i n t h e s t u d y area d u r i n g my v i s i t s i n mid-August and mid-September 1982. Great b l u e h e r o n ; s e e n and heard a t Cedar and C l i f f l a k e s Aug.&Sept. Pintail; a f l o c k o f n i n e w a s s e e n on S e p t . 16 a t Cedar Lake. Goshawk; one s e e n NW o f Cedar Lake S e p t 1 5 Sharp-shinned hawk; o n e s e e n b e i n g mobbed by S t e l l a r ' s j a y s a t Cedar Lake Aug. 1 5 Cooperms hawk; a n immature s e e n i n s e v e r a l p a r t s o f t h e b a s i n , S e p t . R e d - t a i l e d hawk; s e e n b o t h v i s i t s Mountain q u a i l ; s e e n i n mtn. c h a p a r r a l i n Aug. S p o t t e d s a n d p i p e r ; T e r r a c e Lake, Aug. h e a r d from mixed c o n i f e r f o r e s t NW o f Cedar Lake Spotted o w l ; on e v e n i n g of S e p t . 17 Rufous hummingbird i n Aug. ( ? ) p r o b a b l y t h i s s p e c i e s s e e n n e a r Cedar Lake B e l t e d K i n g f i s h e ~ ; commonly s e e n a t a l l , but Upper C l i f f l a k e , S e p t . Red-shafted (Common) f l i c k e r ; s e e n b o t h v i s i t s P i l e a t e d woodpecker; basin Red-breasted h e a r d o n S e p t . 1 6 , b 0 r i n g . s common i n lower sapsucker; Hairy woodpecker; s e e n W o f C l i f f Lake Aug 1 5 seen i n lower b a s i n i n Sept. White-headed woodpecker; s e e n Aug. and S e p t . Hammonds ( ? ) f l y c a t c h e r ; seen i n lower b a s i n i n S e p t . Dusky f l y c a t c h e r ; s e e n commonly i n August Western wood pewee; Olive-sided Violet-green s e e n i n August f l y c a t c h e r ; s e e n and h e a r d i n Aug. s w a l l o w ; s e e n f l y i n g o v e r r i d g e s i n Aug. S t e l l a r : ' : ~jay; common b o t h - v i s i t s Common raven; seen o v e r r i d g e s i n Aug. C l a r k ' s n u t c r a c k e r ; upper e l e v a t i o n s b o t h v i s i t s Mountain chickadee; common both v i s i t s Red-breasted n u t h a t c h ; common both v i s i t s Brown c r e e p e r ; s e e n both v i s i t s Dipper; Sacramento River n e a r NE boundary Aug. House wren; Rock wren; Mtn. c h a p a r r a l Aug. above Upper C l i f f Lake Aug. American r o b i n ; H e r m i t thrush; seen b o t h v i s i t s migrants seen i n Sept., Mountain b l u e b i r d ; probably b r e e d s h e r e , t o o summit of h i g h e s t peak Aug. towns end':^ s o l i t a r e ; f a i r l y common, upper e l e v a t i o n s both visitms Golden-crowned k i n g l e t ; lower b a s i n Aig. and Sept. Ruby-crowned k i n g l e t ; seen Sept S o l i t a r y v i r e o ; heard Aug . and . Sept . Warbling v i r e o ; heard Aug. Orange-crowned w a r b l e r ; s e e n Aug. and S e p t . N a s h v i l l e w a r b l e r ; s e e n n e a r Cedar Lake Aug. Yellow-rumped (Audubon ' s ) w a r b l e r ; Aug. and S e p t . H e r m i t w a r b l e r ; s e e n Aug. M a c G i l l i v r a y ' s w a r b l e r ; seen i n intn. c h a p a r r a l Aug. Wilson's w a r b l e r ; near Cedar Lake Aug. Brown-headed cowbird; s e e n n e a r Cedar Lake Aug. Western t a n a g e r ; s e e n i n A u g . Black-headed grosbeak; s e e n i n Aug. Evening grosbeak; f l o c k seen i n S e p t . n e a r T e r r a c e Lake Cassin's finch; seen both v i s i t s Pine s i s k i n ; seen b o t h v i s i t s Red c r o s s b i l l ; h e a r d i n S e p t . G r e e n - t a i l e d towhee; mtn. c h a p a r r a l b o t h v i s i t s Savannah sparrow; seen n e a r Lower C l i f f Lake Sept. Dark-eyed (Oregon) junco; common b o t h v i s i t s Chipping sparrow; s e e n i n ~ u g . White-crowned sparrow; m i g r a n t s s e e n i n S e p t . Fox sparrow; s e e n b o t h v i s i t s MAMMALS Dusky shrew (Sorex obscurus) ; p o s s i b l e Vagrant shrew (S. - vagrans); possible - Water Shrew (S. p a l u s t r i s ) ; p o s s i b l e Trowbridge shrew (S. t r o w b r i d g e i ) ; p o s s i b l e Broad-handed mole (Scapanus l a t i m a n u s ) ; p o s s i b l e L i t t l e brown myotis (Myotis Fringed rnyotis (M. lucifugus) ; possible thysanoides) ; possible C a l i f o r n i a myotis (M. - californicus); possible Long-e.ared myotis (M. e v o t i s ) ; p o s s i b l e Silvery b a t (Lasionycteris noctivagus) ; possible Hoary b a t ( L a s i u r u s c i n e r e u s ) ; p o s s i b l e Big brown b a t ( E p t e s c i u s f u s c u s ) ; p o s s i b l e B l a c k - t a i l e d h a r e {Lepus c a l i f o r n i c u s ) ; p o s s i b l e Snowshoe h a r e (L. - americanus); possible Mountain beaver (Aplodontia r u f a ) . ; p o s s i b l e Beechey ground s q u i r r e l (Otospermophilus b e e c h e y i ) p o s s i b l e Golden-mantled ground s q u i r r e l (Callospermophilus l a t e r a l i s ) ; common Yellow p i n e chipmunk (Eutamias amoenus) ; p o s s i b l e - townsendii) ; common Townsend's chipmunk (E. Douglas S q u i r r e l (Chickeree) (Tamiasciurus daucJlasii) ; very common Northern f l y i n g s q u i r r e l (Glaucomys s a b r i n u s ) ; p o s s i b l e Mazama pocket gopher (Thomomys mazama) ; common Deer mouse (Peromyscus m a n i c u l a t u s ) ; t h e most common mammal of t h e b a s i n . l i v e - t r a p p i n g on S e p t 17 &18 y i e l d e d 7 7 % c a p t u r e r a t e f o r t h i s species. Bushy-tailed wood r a t (Neotoma c i n e r e a ) ; possible Western red-backed mouse ( CSethrionomys o c c i d e n t a l i s ) ; p o s s i b l e Long-tailed meadow mouse (Microtus l o n g i c a u d u s ) ; p o s s i b l e Oregon meadow mouse (M. - oregoni ) ; possible Montane meadow mouse (M. montanus) ; p o s s i b l e Western jumping mouse (Zapus p r i n c e p s ) ; p o s s i b l e Porcupine ( E r e t h izon docsaturn) ; p o s s i b l e Red f o x (Vulpes f u l v a ) ; p o s s i b l e Coyote (Canis l a t r a n s ) ; chorusas h e a r d b o t h v i s i t s Black bear (Ursus americanus) ; f r e s h s c a t s and tracks s e e n both v i s i t : Raccoon (Procyon l o t o r ) ; possible Marten (Martes a m e r i c a n a ) ; t r a c k s i d e n t i f i e d i n 1980 F i s h e r (M. p e n n a n t i ) ; t r a c k s i d e n t i f i e d i n 1980 ~ o n g - t a i l e dweasel (Mustella £renatal); p o s s i b l e Wolverine (Gulo l u s c u s ) ; t r a c k s i d e n t i f i e d i n 1980 Mountain l i o n ( F e l i s c o n c o l o r ) ; t r a c k s s e e n i n Aug. near Cedar L. Bobcat (Lynx rufus),.; t r a c k s i d e n t i f i e d i n 1980 Mule d e e r (Odocoileus hemianus); f a i r l y common, s e e n both v i s i t s APPENDIX 3 GET AT ION PLOT DESCRIPTIONS* PLOT 1: Dense, c l o s e d POC g r o v e o n W. s i d e of Cedar L. Slopes g r a d u a l , 2-5O t o t h e NE. Typical bottomlarid, d a r k o r g a n i c s o i l , spmse u n d e r s t o r y . 20 1 totals 1 1 1 33 2 1 = 39 g r a n d t o t a l h e r b s and shrubs p r e s e n t : Athyrium f i l i x - f e m i n a , Anenome q u i n q u e f o l i a var. minor, Goodyera o b l o n g i f o l i a , P y r o l a s e c u n d a , ~ r n a l a n c h i e rp a l l i d a . *NOTE: p l o t s 1 t h r o u g h 5 are 10xlOm, p l o t s 6 t h r o u g h 15 a r e 5Ox20m. PLOT 2: Dense, closed POC forest W side of Cedar L. gore open on S. side of plot adjacent to stream. Slopes 2-5 , mesic herbae ceous understory in sunny opening. DBH (inches) POC .5 5 W.F. L.P. WWP D.F. 1 1.5 2 2.5 3 4 4.5 5 5.5 6 7 7.5 8.5 9 10 10.5 1 11 12 1 13.5 1 totals 43 4 3 1 1 1 = 52 herbs and shrubs present: Senecio triangularis, Carex sp.! Viola glabella, Pteridium aqutlinum, Anenlone quinquefolia var. rnlnor, Boykinia major, Lilium kelleyanum, Listera convallarioides, Pyrola secunda, Prunella vulgaris, Amalanchier pallida, Alnus tenuifolia, Rosa pisocarpa. 0 PLOT 3: More open POC bottomland forest. Slopes 0-5 , soil dark with afew gabbro boulders. Shrubby understory dominated by Ledum glandulosum. SW side of Cedar Lake. DBH (inches) POC TOTALS 15 W.F. R.F. L.P. 4 1 4 shrubs and herbs present: 3 1 = 28 Ledurn glandulosum, Leocothoe davisiae, yaccinium arbuscula, Linnaea borealis, Gaultheria humifusa, Amalanchier pallida, Liliurn kelleyanum, Rosa pisocarpa, Pteridium aquilinum, Goodyera oblongifolia. PLOT 4 : R i p a r i a n POC f o r e s t l O O m N of o u t l e t t o L o p r C l i f f Lake. General exposure NE, s t r e a m bank s l o p e s 15-30 ; s o i l i s rocky w i t h decomposed gabbrb.c:and some b o u l d e r s , free d r a i n i n g . Rhododendron o c c i d e n t a l e dominant u n d e r s t o r y shrub. DBH ( i n c h e s ) POC W.F. TOTALS 10:: 1 h e r b s and shrubs p r e s e n t : R.F. L.P. 1 1 WWP 1 Rhododendron o c c i d e n t a l e , Ledum gland- ulosum, Vaccinium a r b u s c u l a , Leocothoe d a v i s i a e , S c h o e n o l i r i o n album, Pedicularis - s e m i b a r b a t a , Pyrola picta. Riparian POC f o ~ e s talong seepy area at S end of Terrace Lake. Slopes 10-15 exposure NNW. Understory shaded, sparse,mostly ; duff overlying rocks and boulders of gabbro. PLOT 5 : . o : .. . DBH (inches) POC 2.5 1 4 1 4.5 1 5 1 5.5 2 M.H. WWP L.P. 2 1 1 18 24.5 TOTALS R.F. 1 21 3 4 2 1 = 31 Shrubs and herbs present: Leucothoe davisiae, Goodyera obloncjifolia, secunda. Chimaphila umbellata, Pyrola picta, P. \ PLOT 6 : Open s e r p e n t i n i t e mixed c o n i f e r f o r e s t c a . l o o m NW of edge of S. Fork Sacramento R. near confluence of o u t l e t streams. SE exposure 0-10° s l o p e s . S e r p e n t i n i t e b o u l d e r s a t upper end and Gabbro a t lower end of p l o t . Mtp. c h a p a r r a l s h r u b s c o v e r i n g c a . 30% of p l o t . DBH ( i n c h e s ) TOTALS W.F. D.F. 33 27 S.P. 17 I.C. L.P. J.P. 11 5 2 WWP 1 = 9.6 h e r b s and shrubs p r e s e n t : Quercus v a c c i n i f o l i a , Rhamnus c a l i f o r n i c a s s p . o c c i d e n t a l i s , Amalanchier p a l l i d a , Monardella o d o r a t i s s i m a s s p . p a l l i d a , Eriophyllum lanatum v a r . lanceolatum, Angelica c a l i f o r n i c a , Ceanothus p r o s t r a t u s , P t e r i d i u m aquilinum, ~ h i r n ~ u m b e l l a t u m , P y r o l a p i c t a , Arenaria c o n g e s t a , Convulvulus malacophyllus , A c h i l l e a pumilwn, P e d i c u l a r i s lanulosa,ola sp. ~ o t e ~ g l a n d u l o s Apocynum a , semibarbata, H'l'eracium cynoglossoides. P PLOT 7: Open s e r p e n t i n i t e mixed c o n i f e r f o r e s t n e a r N E c o r n e r of RNA c a 20m above road. Slopes 10-20° facing ESE; s o i l t h i n , rocky s e r p e n t i n i t e , x e r i c u n d e r s t o r y with Q. v a c c i n i f o l i ~ ,C-prastrat~ predominating some d u f f under l a r g e d o u g l a s f i r s . DBH ( i n c h e s ) W.F. 35 TOTALS I.C. S.P. 19 5 J.P. 6 D.F. 4 = 69 h e r b s and s h r u b s p r e s e n t : S a n i c u l a nevadensis, P y r o l a p i c t a , Kell o g i a q a i l o i d e s , P e d i c u l a r i s s e m i b a r b a t a , Cynoglossum o c c i d e n t a l e , Chimaphila u m b e l l a t a , Anemone q u i n q u e f o l i a v a r . minor, tlieracium c y n o g l o s s o i d e s , A r c t o s t a p h y l o s nevadensis, A. p a t u l a , Quercus v a c c i n i f o l i a Ceanothus ~ r o s t r a t u s Rhamnus-californica ssp. occident a l i s , ~ ~ m ~ h o r i o c a ar ~ z to us s , ~ i r s i u r na n d e r s o n i i . . . - PLOT 8 : Mixed c o n i f e r f o r e s t on S e r p e n t i n i G e , c a . 200m NW of SE f a c i n g s l o p e 25-30 g l a c i a l till, b o u l d e r s and l a r g e r o c k s , t h i n veneer o f s o i l . l i g h t l o g g i n g ( 3 stumps), very open f o r e s t w/ few young t r e e s Understory dominated by Q. v a c c i n i f o l i a , b u t m o i s t e r a r e a on S W w i t h Rhododendron and ~ o s a . NW s h o r e of Cedar L. . DBH ( i n c h e s ) D.F. W.F. S.P. 8.5 I.C. J.P. WWP 1 11 11.5 15 16 17 17.5 23 25 26 26.5 27 33 37 37.5 TOTALS 6 3 1 4 2 2 = 18 Herbs and s h r u b s : Juncus p a r r y i , Lotus c r a s s i c a u l i s , Vaccinium arbus c u l a , P t e r i d i u m aquilinum, Lupinus c r o c e u s , Lupinus s p . -~ o s <pisdcar.pa:l\.:~yrof h a : ~ . ~Quercus ic, v a c c i n i f o l i a , Arctostaphylos - , p a t u l a , Ribes r o e z l i i , Ceanothus p r o s t r a t u s , Rhamnus nevadensis, A ~ a l i f o r n i c a ,Rhododendron o c c i d e n t a l e , Chimaphila umbellata. S e r p e n t i n i t e mixed c o n i f - e r f o r e s t c a . 300m NW of Ce,dar Lake Slopes' '.25-30°, ENE exposure, open f o r e s t and u n d e r s t o r y , s u b s t s a t e g l a c i a l t i l l , Q. v a c c i n i f o l i a dominant shrub. PLOT 9 : DBH $ i n c h e s ) W.F. TOTALS 16 S.P. 3 D.F. WWP 4 2 I.C. 1 = 26 s h r u b s and h e r b s p r e s e n t : Q. v a c c i n i f o l i a , A r c t o s t a p h y l o s nevadensis, A. p a t u l a , ~ e a n o t h u s p r o s t r ~ t u sRhamnus , californica occidentalis, Fyrola p i c t a , e r o p h y l l u r n t e n a x , Goodyera oblong- PLOT $0: . : S e r p e n t i n i t e mixed c o n i f e r f o r e s t , open forest ca 50m from the t o p of r i d g e NW of Cedar Lake. Slope 20-30° s u b s t r a t e s e r p e n t i n i t e g l a c i a l till. Herbs and s h r u b s same as f o r p l o t #9. DBH (inches) W.F. .5 S.P. J.P. 2 1 3 1.5 2 2 5 2.5 3 3 1 3.5 1 4 2 4.5 1 5 1 1 WWP D . F. I.C. 1 1 2 1 1 1 6 7 1 9 1 9.5 1 10 10.5 12 13 15 15.5 2 17 1 17.5 1 18 1 22 1 22.5 1 28 TOTALS 29 1 13 2 6 2 3 = 52 PLOT 11: Lodgepole pine-mountain hemlock-red fir f o r e s t on v a l l e y bottom c a . 300m S of Terrace Lake. Slopes 15-20° t o NNE on lower part where mtn. hemlock dominant, l e v e l n e a r upper p a r t where l o d g e p o l e p i n e dominant. S o i l o r g a n i c A h o r i z o n w i t h s c a t t e r e d gabbro slabs and b o u l d e r s , POC a l o n g i n t e r m i t t e n t runoff r i l l . DBK ( i n c h e s ) M.H. R.F. POC. L.P. - WWP . W.F. - - TOTALS 118 48 26 29 10 8 = 239 h e r b s and shrubs: Leucothoe d a v i s i a e , A r c t o s t a p h y l o s nevadensis, Vaccinium a r b u s c u l a , ~ p ~ ~ m a l a n c h i e r d ~ i n i f o l i a , Chimaphila . - - . . - umbgllata, Anemone q u i n q u e f o l i a , Pyrola secunda, A s t e r brickellioides. . . PLOT 12: R e d f i r - l o d g e p o l e pine-western w h i t e p i n e f o r e s t , 400m S W of T e r r a c e Lake. open f o r e s t with s c a t t e r e d gabbro r o c k s and boulders.' Slope 5-20° f a c e s ENE. Understory dominated by Arctos t a p h y l o s k l a m a t h e n s i s ( t y p e l o c a l i t y of s p e c i e s ) DBH ( i n c h e s ) R.F. . L.P. TOTALS h e r b s and s h r u b s p r e s e n t : A . k l a m a t h e n s i s ( 6 3 % c o v e r ) , A. nevadensis ( 6 % c o v e r ) , Quercus v a c c i n i F o l i a ( 9 % cover), A. p a t u l a (3%c o v e r ) , Ceanothus prostratus(3% A s t e r b r i c k e i l i o i d e s , Arabis p l a t y sperma, Juncus p a r r y i , Luzula d i v a r i c a t a , Sedum obtusatum s s p b o r e a l e . -- PLOT 13: R e d f i r - m o u n t a i n hemlock f o r e s t ca0.75krn S W Terrace Lake at ca.06800ft.. Closed forest e x c e p t lower end, slopes s t e e p 30-45 , NE facing. Snow l o d g i n g e v i d e n t on many young trees ; u n d e r s t o r y m o s t l y d u f f e x c e p t lower p l o t w/ some open, rocky (gabbro) areas. DBH ( i n c h e s ) R.F. 124 TOTALS M.H. 49 WWP 6 = 179 h e r b s a n s s h r u b s present: A r c t o s t a p h y l o s klamathensis, Chimaphila u m b e l l a t a , Juncus p a r r y i , Sedum obtusatum ssp. boreale. T - PLOT 14: Red fir-western White pine forest, open . Senescent , several large snags, much young fir regeneration, ca. 0.75km S of Terrace lake. Slopes 15-20° on bench w/ some exposed gabbro outcrops at top of plot, facing ENE. open understory w/ shrubs at top. DBH (inahes) TOTALS R.F. M.H. 38 9 9 = 56 Herbs and shrubs: Juncus parryi, Chimaphila umbellata, Pyrola picta, Arctostaphylos klamathensis, Quercus v a c c i n i f o ~ n -e v a d e n s i s .