Hindawi Publishing Corporation Abstract and Applied Analysis Volume 2013, Article ID 624352, 5 pages http://dx.doi.org/10.1155/2013/624352 Research Article A Lotka-Volterra Competition Model with Cross-Diffusion Wenyan Chen and Ya Chen Department of Mathematics, Southeast University, Nanjing 210018, China Correspondence should be addressed to Wenyan Chen; wychen@seu.edu.cn Received 9 October 2012; Revised 24 January 2013; Accepted 29 January 2013 Academic Editor: Lan Xu Copyright © 2013 W. Chen and Y. Chen. This is an open access article distributed under the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited. A Lotka-Volterra competition model with cross-diffusions under homogeneous Dirichlet boundary condition is considered, where cross-diffusions are included in such a way that the two species run away from each other because of the competition between them. Using the method of upper and lower solutions, sufficient conditions for the existence of positive solutions are provided when the cross-diffusions are sufficiently small. Furthermore, the investigation of nonexistence of positive solutions is also presented. 1. Introduction In this paper, we deal with the following Lotka-Volterra competition model with cross-diffusions: −Δ (π’ + πΌV) = π’ (π − π’ − πV) , π₯ ∈ Ω, −Δ (π½π’ + V) = V (π − V − ππ’) , π₯ ∈ Ω, π’ = V = 0, (1) π₯ ∈ πΩ, where Ω is a bounded domain in Rπ with smooth boundary πΩ and all parameters π, π, π, π, πΌ, π½ are positive constants. π’ and V stand for the densities of the two competitors; π and π are the intrinsic growth rates of π’ and V, respectively; π and π are the competitive parameters between the two species; Here πΌ and π½ are referred to as cross-diffusions. Cross-diffusions express the two species run away from each other because of the competition between them. In this paper, the boundary condition is under homogeneous Dirichlet boundary condition which in biologically means that the boundary is not suitable for both species and they will all die on the boundary, and this is an ideal case. In order to describe the meaning of cross-diffusions in this model (1) from the biological point, we give the general model with intrinsic diffusion and cross-diffusion: ππ’ = div {π11 (π’, V) ∇π’ + π12 (π’, V) ∇V} + π (π’, V) , ππ‘ (2) πV = div {π21 (π’, V) ∇π’ + π22 (π’, V) ∇V} + π (π’, V) , ππ‘ where π’ and V stand for the densities of the two species, intrinsic diffusion parameters π11 (π’, V), π22 (π’, V) > 0, crossdiffusion parameters π12 (π’, V), π21 (π’, V), π½π’ = − {π11 (π’, V) ∇π’ + π12 (π’, V) ∇V} , π½V = − {π21 (π’, V) ∇π’ + π22 (π’, V) ∇V} (3) can be seen as the out-flux vector of π’ and V at π₯. The cross-diffusion parameters π12 (π’, V), π21 (π’, V) ≥ 0 imply that the two competitors π’ and V diffuse in the direction of lower contrary of their competitor to avoid each other. π(π’, V), π(π’, V) are response function and in this paper the classical Logistic Type is considered and πΌ, π½ ≥ 0. More biological meaning of the system can be seen in [1–3]. The method of upper and lower solutions is a useful tool to study the existence of solutions of elliptic systems. However, there are many difficulties in investigating the existences of positive solutions of strongly coupled elliptic systems. Recently, by changing general strongly coupled elliptic systems into weakly coupled ones, the author in paper [4] gives the method to judge the solutions existence of elliptic systems by using the Schauder theorem. Furthermore, the method can be used to solve the existence of solutions of strongly coupled elliptic systems. In [5] Ko and Ryu 2 Abstract and Applied Analysis investigate Lotka-Volterra prey-predator model with crossdiffusion: −Δπ’ = π’ (π1 − π’ − π12 V) , π₯ ∈ Ω, −π·Δπ’ − ΔV = V (π2 + π21 π’ − V) , π’ = V = 0, π₯ ∈ Ω, π’ = π΄∗ (π€, π§) , Here π· may be positive or negative. Using the developing method of upper and lower solutions in [4], the author gave a sufficient conditions for the existence of positive solutions to (4). Inspired by the paper [5], we investigate the existence and nonexistence of positive solutions to (1). The main goal of this paper is to provide sufficient conditions for the existence of positive solutions to (1) when the cross-diffusions πΌ and π½ are small. More precisely, we have the following theorem. Let π 1 > 0 be the principal eigenvalue of −Δ under homogeneous Dirichlet boundary condition. It is well known that the principal eigenfunction π corresponding to π 1 does not change sign in Ω and ||π||∞ = 1. Theorem 1. If min{π−ππ, π−ππ} > π 1 , then there exist positive constants πΌ = πΌ(π, π, π, π, Ω), π½ = π½(π, π, π, π, Ω), when πΌ < πΌ, π½ < π½, (1) has at least one positive solution. For πΌ = π½ = 0, (1) is the Lotka-Volterra competition model under homogeneous Dirichlet boundary condition. In [6, 7], the authors use different methods to prove the existence of positive solutions, a sufficient condition for the existence is min{π − ππ, π − ππ} > π 1 . The conclusion implies that weakly cross-diffusion does not affect the existence of positive solution. This paper is organized as follows. In Section 2, the existence theorem of solutions for a general class of strongly coupled elliptic systems is presented using the method of upper and lower solutions. In Section 3, sufficient conditions for the existence and nonexistence of positive solutions of (1) are investigated. Moreover, we give the corresponding results simply if the competitive system only has one cross-diffusion. 2. The Existence Theorem of Solutions for a Class of Strongly Coupled Elliptic Systems In this section, we presented the existence theorem of solutions for a general class of strongly coupled elliptic systems: π₯ ∈ Ω, −Δπ΅ (π’, V) = π2 (π’, V) , π₯ ∈ Ω, π’ = V = 0, (5) π₯ ∈ πΩ. Here let π΄, π΅, π1 , π2 satisfy the following hypotheses conditions. (H1) π, π are domain in R2 , (0, 0) ∈ π. (π΄, π΅) is a πΆ2 function about (π’, V) from π to π, π΄(0, 0) = π΅(0, 0) = 0, and have a continuous inverse (π΄∗ , π΅∗ ) ∈ πΆ2 (π, π). Then for all (π’, V) ∈ π, let π€ = π΄ (π’, V) , π§ = π΅ (π’, V) . V = π΅∗ (π€, π§) . (7) (4) π₯ ∈ πΩ. −Δπ΄ (π’, V) = π1 (π’, V) , There exists only one (π€, π§) ∈ π, satisfying (6) (H2) The function π΄∗ is increasing in π€ and decreasing in π§; π΅∗ is decreasing in π€ and increasing in π§. (H3) The functions π1 (π’, V), π2 (π’, V) are Lipschitz continuous in π, and there exist positive constants π1 , π2 such that for all (π’, V) ∈ π, the function π1 (π’, V) + π1 π΄(π’, V) is increasing in π’; the function π2 (π’, V) + π2 π΅(π’, V) is increasing in V. According to the hypothesis (H1), (5) can be rewritten as the following equal PDE equations: −Δπ€ + π1 π€ = π1 (π’, V) + π1 π΄ (π’, V) , π₯ ∈ Ω, −Δπ§ + π2 π§ = π2 (π’, V) + π2 π΅ (π’, V) , π₯ ∈ Ω, π’ = π΄∗ (π€, π§) , V = π΅∗ (π€, π§) , π€ = π§ = 0, π₯ ∈ Ω, (8) π₯ ∈ πΩ. Remark 2. According to the hypothesis (H1), (5) can also be equal to the following weakly coupled elliptic equations: −Δπ€ = π1 (π΄∗ (π€, π§) , π΅∗ (π€, π§)) := π1 (π€, π§) , π₯ ∈ Ω, −Δπ§ = π2 (π΄∗ (π€, π§) , π΅∗ (π€, π§)) := π2 (π€, π§) , π₯ ∈ Ω, (9) π€ = π§ = 0, π₯ ∈ πΩ. In its pure form, (9) is simpler than (8). However, due to the complicity of mixed functions π1 (π€(π₯), π§(π₯)) and π2 (π€(π₯), π§(π₯)), it is difficult to find the solutions of (9) directly. Therefore, we discuss (8). Assume functions π’, V, π’, V ∈ πΆ(Ω), π€, π§, π€, π§ ∈ πΆπΌ (Ω) β πΆ2 (Ω), the values of functions (π’, V) and (π’, V) are in π and the values of functions (π€, π§) and (π€, π§) are in π. To describe easily, let π = {π’ ∈ πΆ (Ω) : π’ (π₯) ≤ π’ (π₯) ≤ π’ (π₯)} , π = {π’ ∈ πΆ (Ω) : V (π₯) ≤ V (π₯) ≤ V (π₯)} . (10) According the definition of upper and lower solutions in [4] and conditions (H1)–(H3), we give the definition of upper and lower solutions of (5). Definition 3. A pair of functions ((π’, V, π€, π§), (π’, V, π€, π§)) are called upper and lower solutions of (9) provided that they Abstract and Applied Analysis 3 satisfy the relation (π’, V, π€, π§) ≥ (π’, V, π€, π§), and for all (π’, V) ∈ π × π, satisfy the following inequalities: −Δπ€ + π1 π€ ≥ π1 (π’, V) + π1 π΄ (π’, V) , π₯ ∈ Ω, −Δπ§ + π2 π§ ≥ π2 (π’, V) + π2 π΅ (π’, V) , π₯ ∈ Ω, −Δπ€ + π1 π€ ≤ π1 (π’, V) + π1 π΄ (π’, V) , π₯ ∈ Ω, −Δπ§ + π2 π§ ≤ π2 (π’, V) + π2 π΅ (π’, V) , π₯ ∈ Ω, π’ ≥ π΄∗ (π€, π§) , V ≥ π΅∗ (π€, π§) , π₯ ∈ Ω, π’ ≤ π΄∗ (π€, π§) , V ≤ π΅∗ (π€, π§) , π₯ ∈ Ω, π€ ≥ 0 ≥ π€, π§ ≥ 0 ≥ π§, Lemma 6. If the functions π’, V ∈ πΆ1 (Ω) satisfy π’|πΩ = V|πΩ = 0, π’|Ω > 0, (ππ’/ππ)|πΩ < 0, π is the outer unit normal vector of πΩ, then there exists positive constant π, such that π’(π₯) > πV(π₯), for all π₯ ∈ Ω. (11) π’ = 0, We can have the following conclusion from [4, Theorem 2.1]. Proposition 4. Assume that (8) has coupled upper and lower solutions ((π’, V, π€, π§), (π’, V, π€, π§)), then there exists at least one solution (π’, V, π€, π§), satisfying the relation (π’, V, π€, π§) ≤ (π’, V, π€, π§) ≤ (π’, V, π€, π§) . (12) Furthermore, (π’, V) is the solution of (5). Next, if π’, V, π’, V satisfy V = π΅∗ (π€, π§) , π’ = π΄∗ (π€, π§) , V = π΅∗ (π€, π§) , (13) π§ = π΅ (π’, V) , π€ = π΄ (π’, V) , π§ = π΅ (π’, V) , (14) (11) can be rewritten as π₯ ∈ Ω, (17) π₯ ∈ πΩ. Lemma 7. If π > π 1 , then (17) has a unique positive solution ππ satisfying ππ ≤ π. In addition, ππ is increasing with respect to π. 3. A Lotka-Volterra Competition Model with Two Cross-Diffusions In this section, the existence of positive solutions of (1) corresponding to πΌ ≥ 0, π½ ≥ 0, is investigated by applying Theorem 5 to prove Theorem 1. Proof. We seek some positive constants π , πΎ, πΏ, π , πΎ > π 1 sufficiently large and πΏ sufficiently small, Lemma 6 may guarantee the existence of ππ and ππΎ . It can be easily known from Hopf boundary lemma: ππ (π₯) < 0, ππ then π€ = π΄ (π’, V) , For the equation: −Δπ’ = π’ (π − π’) , π₯ ∈ πΩ. π’ = π΄∗ (π€, π§) , To make sure the upper and lower solutions reasonable, we give the following two lemmas; more details can be found in [8, 9]. πππ (π₯) < 0, ππ πππΎ (π₯) < 0, ππ ∀π₯ ∈ πΩ. Observe that min{π − ππ, π − ππ} > π 1 , using Lemma 7, we can have π , πΎ, πΏ, π < π < (π − π 1 )/π, π < πΎ < (π − π 1 )/π, satisfying the following three conditions: (i) πΏπ(π₯) < ππ (π₯), πΏπ(π₯) < ππΎ (π₯), for all π₯ ∈ Ω; −Δπ΄ (π’, V) + π1 π΄ (π’, V) ≥ π1 (π’, V) + π1 π΄ (π’, V) , π₯ ∈ Ω, (ii) (π(ππ − πΏπ)/ππ)(π₯) < 0, (π(ππΎ − πΏπ)/ππ)(π₯) < 0; −Δπ΅ (π’, V) + π2 π΅ (π’, V) ≥ π2 (π’, V) + π2 π΅ (π’, V) , π₯ ∈ Ω, (iii) πΏ < min{π − π 1 − ππΎ, π − π 1 − ππ }. −Δπ΄ (π’, V) + π1 π΄ (π’, V) ≤ π1 (π’, V) + π1 π΄ (π’, V) , π₯ ∈ Ω, −Δπ΅ (π’, V) + π2 π΅ (π’, V) ≤ π2 (π’, V) + π2 π΅ (π’, V) , π₯ ∈ Ω, π΄ (π’, V) ≥ 0 ≥ π΄ (π’, V) , π₯ ∈ πΩ. (15) π΅ (π’, V) ≥ 0 ≥ π΅ (π’, V) , Synthetically, we have the following result. Theorem 5. If there is a pair of functions ((π’, V), (π’, V)), satisfying (π’, V, π΄ (π’, V) , π΅ (π’, V)) ≥ (π’, V, π΄ (π’, V) , π΅ (π’, V)) , (16) and for all (π’, V) ∈ π × π, (15) is satisfied, then (5) has at least one solution (π’, V), satisfying the relation (π’, V) ≤ (π’, V) ≤ (π’, V). (18) Let π1 = 2π +ππΎ, π2 = 2πΎ+ππ . Using Lemma 7 again, there exist πΌ = πΌ(π, π, π, π, Ω) < 1, π½ = π½(π, π, π, π, Ω) < 1, for all (π, π) ∈ [0, πΌ) × [0, π½), for all π₯ ∈ Ω, satisfying (iv) ππ − πΏπ > π(ππΎ − πΏπ), ππΎ − πΏπ > π(ππ − πΏπ); (v) (π − π)ππ > π[π1 ππΎ − (π1 + π 1 )πΏπ], (πΎ − π)ππΎ > π[π2 ππ − (π2 + π 1 )πΏπ]; (vi) (π − π 1 − πΏ − ππΎ)πΏπ > π[(πΎ + π1 − ππΎ )ππΎ − π1 πΏπ]; (vii) (π − π 1 − πΏ − ππ )πΏπ > π[(π + π2 − ππ )ππ − π2 πΏπ]. We will verify πΌ, π½ satisfying Theorem 5. Suppose that (πΌ, π½) ∈ [0, πΌ) × [0, π½). Then we construct a pair of upper and lower solutions of the form (π’, V) = (ππ , ππΎ ) , (π’, V) = (πΏπ, πΏπ) , (19) 4 Abstract and Applied Analysis where πΏ satisfies conditions (i)–(iii). Let π΄ (π’, V) = π’ + πΌV, π΅ (π’, V) = π½π’ + V. Theorem 8. Any positive solutions (π’, V) of (1) have a priori bound; that is (20) π’ (π₯) ≤ Then π΄∗ (π€, π§) = π€ − πΌπ§ , 1 − πΌπ½ π΅∗ (π€, π§) = π§ − π½π€ . 1 − πΌπ½ π’= π2 (π’, V) = V (π − V − ππ’) . (22) And for all (π’, V) ∈ π, we have [π2 (π’, V) + π2 π΅ (π’, V)]V = π − 2V − ππ’ + π2 π€ − πΌπ§ , 1 − πΌπ½ (26) (23) V= π§ − π½π€ . 1 − πΌπ½ (27) Equation (1) can be rewritten as −Δπ€ = π§ − π½π€ π€ − πΌπ§ π€ − πΌπ§ (π − −π ), 1 − πΌπ½ 1 − πΌπ½ 1 − πΌπ½ π₯ ∈ Ω, −Δπ§ = π§ − π½π€ π§ − π½π€ π€ − πΌπ§ (π − −π ), 1 − πΌπ½ 1 − πΌπ½ 1 − πΌπ½ π₯ ∈ Ω, (π€, π§) = (0, 0) , [π1 (π’, V) + π1 π΄ (π’, V)]π’ = π − 2π’ − πV + π1 ≥ −2π − ππΎ + π1 = 0, π . π V (π₯) ≤ Proof. Let π€ = π’ + πΌV, π§ = π½π’ + V; then (21) By simply computing, (H1) and (H2) are satisfied, where π = [0, π ] × [0, πΎ], π = [0, π + πΌπΎ] × [0, πΎ + π½π ]. Note π1 (π’, V) = π’ (π − π’ − πV) , π , π (28) π₯ ∈ πΩ. Since (π’, V) > (0, 0), it easily follows that π€−πΌπ§ > 0, π§−π½π€ > 0. Assume that π§(π₯) attains its positive maximum at π₯0 ∈ Ω, then π (1 − πΌπ½) − π€ (π₯0 ) + πΌπ§ (π₯0 ) − ππ§ (π₯0 ) + ππ½π€ (π₯0 ) > 0 ≥ −2πΎ − ππ + π2 = 0. So (H3) is satisfied; observer that π’|πΩ = V|πΩ = π’|πΩ = V|πΩ = 0, (π’, V) ≥ (π’, V) and (iv) and (15) and the boundary conditions of (16) can be checked. Therefore, if we want to obtain the existence of solutions through [4, Theorem 2.1], we should only verify for all (π’, V) ∈ π × π, −Δπ΄ (π’, V) + π1 π΄ (π’, V) ≥ π1 (π’, V) + π1 π΄ (π’, V) , π₯ ∈ Ω, −Δπ΅ (π’, V) + π2 π΅ (π’, V) ≥ π2 (π’, V) + π2 π΅ (π’, V) , π₯ ∈ Ω, −Δπ΄ (π’, V) + π1 π΄ (π’, V) ≤ π1 (π’, V) + π1 π΄ (π’, V) , π₯ ∈ Ω, −Δπ΅ (π’, V) + π2 π΅ (π’, V) ≤ π2 (π’, V) + π2 π΅ (π’, V) , π₯ ∈ Ω. (24) Because π1 is decreasing in V, π2 is decreasing in π’, and π΄(π’, V) is increasing in V, π΅(π’, V) is increasing in π’, only to verify the following inequations: π (1 − πΌπ½) − ππ§ (π₯0 ) + ππ½πΌπ§ (π₯0 ) > 0, π§ (π₯) ≤ π§ (π₯0 ) ≤ π π (29) so that V = π§ − π½π’ ≤ π§ (π₯0 ) ≤ π . π Similarly, we can obtain the desired result π’≤ π . π (31) Theorem 9. If one of the following conditions: (i) π ≤ ππ, π 1 ≥ (π + ππ½(π/π))/(1 − πΌπ½); (ii) (1 − (πΌ + π½)/2)π 1 ≥ max{π, π}; −Δπ΄ (π’, V) + π1 π΄ (π’, V) ≥ π1 (π’, V) + π1 π΄ (π’, V) , π₯ ∈ Ω, −Δπ΅ (π’, V) + π2 π΅ (π’, V) ≥ π2 (π’, V) + π2 π΅ (π’, V) , π₯ ∈ Ω, −Δπ΄ (π’, V) + π1 π΄ (π’, V) ≤ π1 (π’, V) + π1 π΄ (π’, V) , π₯ ∈ Ω, −Δπ΅ (π’, V) + π2 π΅ (π’, V) ≤ π2 (π’, V) + π2 π΅ (π’, V) , π₯ ∈ Ω. (25) ∫ |∇π’|2 ππ₯ + πΌ ∫ ∇π’∇Vππ₯ = ∫ π’2 (π − π’ − πV) ππ₯, It is easy to check (25) by (v), (vi), and (vii). So from [4, Theorem 2.1], (1) has a solution (π’, V), in addition (π’, V) ≥ (π’, V) ≥ (π’, V) > (0, 0). πΌ ∫ |∇V|2 ππ₯ + ∫ ∇π’∇Vππ₯ = ∫ π’V (π − π’ − πV) ππ₯, In the end, before investigating the nonexistence of positive solutions of (1), we give its priori bound of positive solutions. (30) is satisfied, then (1) with πΌ < πΌ, π½ < π½ has no positive solution. Proof. Multiplying π’ and V to the first and second equations in (1), and integrating these equations on Ω, we have Ω Ω Ω Ω Ω Ω 2 π½ ∫ |∇π’| ππ₯ + ∫ ∇π’∇Vππ₯ = ∫ π’V (π − V − ππ’) ππ₯, Ω Ω Ω ∫ |∇V|2 ππ₯ + π½ ∫ ∇π’∇Vππ₯ = ∫ V2 (π − V − ππ’) ππ₯. Ω Ω Ω (32) Abstract and Applied Analysis 5 (i) Suppose, by contradiction that (1) has a positive solution (π’, V), then the second and fourth equations in (32) yield ≥ [(1 − πΌ+π½ ) π 1 − π] ∫ π’2 ππ₯ 2 Ω + [(1 − ∫ V3 ππ₯ + π½ ∫ π’V (π − π’) ππ₯ Ω πΌ+π½ ) π 1 − π] ∫ V2 ππ₯ ≥ 0. 2 Ω (36) Ω = − (1 − πΌπ½) ∫ |∇V|2 ππ₯ + π½ ∫ ππ’V2 ππ₯ Ω Ω (33) + ∫ V2 (π − ππ’) ππ₯. Ω Since π’ ≤ π/π by Theorem 8, the left-hand side of (33) must be positive. On the other hand, the Poincare inequality, β∇Vβ2πΏ2 ≥ π 1 βVβ2πΏ2 , for V ∈ π21 (Ω) and the given assumption shows the following contradiction: However, this results in a contradiction since the right-hand side of (35) is clearly strictly negative by the positivity of π’ and V. Remark 10. Before closing this section, more sufficient conditions of the nonexistence of positive solutions of (1) with πΌ + π½ > 0, πΌπ½ = 0 are investigated. Take πΌ = 0, π½ > 0 for example, then (1) may be reduced as −Δπ’ = π’ (π − π’ − πV) , π₯ ∈ Ω, −Δ (π½π’ + V) = V (π − V − ππ’) , 2 2 2 − (1 − πΌπ½) ∫ |∇V| ππ₯ + π½ ∫ ππ’V ππ₯ + ∫ V (π − ππ’) ππ₯ Ω Ω ≤ − [(1 − πΌπ½) π 1 − ππ½ (π’, V) = (0, 0) , Ω π − π] ∫ V2 ππ₯ ≤ 0. π Ω π₯ ∈ Ω, (37) π₯ ∈ πΩ. Using the same method, we can obtain that (37) has no positive solution, if one of the following conditions is satisfied: (34) (ii) A contraction argument is also used assuming that (1) has a positive solution (π’, V). Adding the first equation to the fourth equation, and then subtracting π ∫Ω π’2 ππ₯ + π ∫Ω V2 ππ₯ from the both sides, the following identity is obtained: (i) π 1 ≥ π + π½ππ; (ii) π 1 ≥ π; (iii) (1 − π½/2)π 1 ≥ max{π, π}; (iv) π < 1 < π/π and (1 − π)/π½ ≤ π 1 /(π + π½π) ≤ 1. 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