CHAPTER 38: ANGIOSPERM REPRODUCTION & BIOTECHNOLOGY BY: TREVOR GULLEDGE, ASHLEY LETO, AND JILL RICHARDS 38.1: FLOWERS, DOUBLE FERTILIZATION, AND FRUITS ARE UNIQUE FEATURES OF THE ANGIOSPERM LIFE CYCLE LIFE CYCLE OF PLANTS • ALTERNATES BETWEEN SPOROPHYTE AND GAMETOPHYTES GENERATIONS • SPOROPHYTE PRODUCE SPORES BY MEIOSIS • DOMINATE GENERATION • GAMETOPHYTES PRODUCE GAMETES FROM THE SPORES BY MITOSIS • FERTILIZAITON IS FUSION OF GAMETES MAKING A DIPLOID ZYGOTES • THE ZYGOTES DIVIDE AND FORM SPOROPHYTES • • FLOWER STRUCTURE STAMEN • FILAMENT HOLDS THE ANTHER • ANTHER PRODUCE POLLEN CARPEL • OVARY CONTAINS OVULES • STIGMA CATCHES POLLEN • STYLE HOLDS STIGMA • PISTIL ARE A GROUP OF CARPELS Flower Structure MALE GAMETOPHYTES • POLLEN • GENERATIVE CELL AND TUBE CELL • FORM FROM MICROSPORES • FORM FROM MICROSPOROCYTE • COME FROM MICROSPORANGIA LOCATED IN THE ANTHER FEMALE GAMETOPHYTES • EMBRYO SAC • FORM FROM MEGASPORES • FORM FROM MEGASPOROCYTE • COME FROM THE MEGASPORANGIUM POLLINATION • POLLINATION IS WHEN POLLEN HANGS ON A STIGMA OF A CARPEL • TYPES • WIND (ABIOTIC) • BEES • MOTHS AND BUTTERFLIES • BIRDS • FLIES • BATS DOUBLE FERTILIZATION • WHEN TWO SPERM ARE DISCHARGED INTO THE FEMALE GAMETOPHYTE • ONE FERTILIZES • ONE COMBINES WITH A POLAR NUCLEI FORMING A FOOD-STORING ENDOSPERM SEEDS • THE SEED HOLDS THE EMBRYO AND EITHER AN ENDOSPERM OR COTYLEDONS • SEED DORMANCY ALLOWS THE SEED TO GERMINATE IN THE RIGHT CONDITIONS FOR SURVIVING FRUIT • PROTECTS THE SEED • AIDS IN WIND DISPERSAL • ATTRACTS SEED-DISPERSING ANIMALS 38.2: REPRODUCTION OF FLOWERING PLANTS MECHANISMS OF ASEXUAL REPRODUCTION FRAGMENTATION: SEPARATION OF PARENT PLANT INTO INTO PARTS THAT DEVELOP INTO WHOLE PLANTS ASEXUAL REPRODUCTION PRODUCES CLONES OF THE PARENT APOMIXIS: ASEXUAL REPRODUCTION OF SEEDS IN WHICH A DIPLOID CELL IN THE OVULE CREATES AN EMBRYO, WHICH THEN MATURES INTO SEEDS DANDELIONS REPRODUCE ASEXUALLY ADVANTAGES/DISADVANTAGES OF ASEXUAL VERSUS SEXUAL REPRODUCTION ASEXUAL REPRODUCTION ADVANTAGES: NO NEED FOR POLLINATOR, PASS ON ALL GENETIC INFO, CLONES GENETICALLY WELL ADAPTED TO ENVIRONMENT, INCREASED DISPERSION DISADVANTAGES: MECHANISMS HAVE EVOLVED THAT MAKE IT DIFFICULT OR IMPOSSIBLE FOR A FLOWER TO SELF-FERTILIZE, IDENTICAL GENES INCREASE CHANCE OF EXTINCTION SEXUAL REPRODUCTION ADVANTAGES: GENERATES VARIATION, RESISTANCE TO EXTINCTION AND PATHOGENS DISADVANTAGES: NEED FOR POLLINATOR MECHANISMS THAT PREVENT SELF-FERTILIZATION PREVENTION INCREASES GENETIC VARIETY BY ENSURING THAT THE SPERM AND EGG CELLS COME FROM DIFFERENT PARENTS DIOECIOUS SPECIES: NO SELF-FERTILIZATION BECAUSE THE PLANTS EITHER HAVE CARPELS (CARPELLATES) OR STAMEN (STAMENATES) OTHER PLANTS HAVE BOTH STAMEN AND CARPELS, BUT THEY MATURE AT DIFFERENT RATES OR ARE IN AN ARRANGEMENT THAT MAKES IT DIFFICULT FOR A POLLINATOR TO POLLINATE SELF-INCOMPATIBILITY: A PLANT CAN REJECT ITS OWN POLLEN S-GENES DICTATE WHETHER POLLEN IS REJECTED OR ACCEPTED (POLLEN BLOCKING) PIN AND THRUM FLOWERS HAVE DIFFERENT LENGTH STAMENS AND STYLES WHICH FAVOR CERTAIN KINDS OF POLLINATORS (GENETIC VARIATION) AND MAKE IT DIFFICULT TO SELFPOLLENATE VEGETATIVE PROPAGATION AND AGRICULTURE CUTTINGS: PLANT FRAGMENTS THAT GROW AND DEVELOP INTO A NEW PLANT GRAFTING: WHEN A TWIG OR BUD OF ONE PLANT IS COMBINED WITH A CLOSELY RELATED SPECIES TO CREATE A SINGLE PLANT THE PLANT THAT PROVIDES THE ROOT SYSTEM=THE STOCK THE TWIG GRAFTED TO THE STOCK=THE SCION ALLOWS FOR COMBINATION OF BEST GENETIC QUALITIES TEST TUBE CLONING: FORM OF IN VITRO PLANT BREEDING USED IN THE PRODUCTION OF GM PLANTS (TRANSGENIC) AND THE INVENTION OF NEW PLANT VARIETIES (PROTOPLAST FUSION) FROM LEFT TO RIGHT: PROTOPLASM CULTURE, AFRICAN VIOLETS, AND WINE GRAPES 38.3: HUMANS MODIFY CROPS BY BREEDING AND GENETIC ENGINEERING PLANT BREEDING IN NATURE, PLANTS HYBRIDIZE ON THEIR OWN. HYBRIDIZATION HAS BEEN USED BY ANCIENT FARMERS AS WELL AS MODERN ONES ONCE PLANTS HAVE SUCCESSFULLY BRED, THE ONES WITH THE BEST TRAITS ARE CHOSEN MAIZE PLANT BIOTECHNOLOGY AND GENETIC ENGINEERING GENES FROM UNRELATED ORGANISMS ARE INTRODUCED INTO PLANTS GENETICALLY MODIFIED PLANTS CAN INCREASE THE AMOUNT AND THE QUALITY OF FOOD CAN BE USED AS ECOLOGICALLY SOUND FUEL. DEBATE OVER PLANT BIOTECHNOLOGY THERE ARE SOME UNKNOWN RISKS OF GMO’S HOWEVER, THE POTENTIAL FOR BENEFICIAL RESULTS MUST BE TAKEN INTO CONSIDERATION.