I. Populations and Variations • Population - a group of individuals belonging to the same species that occupy a given area. • In most natural populations, the manifestations of traits are not the same from one individual to the next. (variations) • Variations in traits are inherited. • Information about traits resides in hundreds of genes. • The genetic variation results in different phenotypes within a population. • more than 10600 combinations of genes are possible in human gametes. • Far more genetic variation is possible than can ever be expressed in the individuals of any population. II. Allele Frequencies and Gene Pools • Gene pool - all the genotypes that exist in a population • Remember that the different forms of a gene are called alleles. • Allele Frequencies - The relative abundance of each kind of allele in a population. • The number and type of alleles in a population is constantly changing. • Evolution occurs when there is a change in the allele frequency of a population • Changes in allele frequencies can occur in a number of different ways: 1) Mutation 2) Genetic Drift 3) Gene Flow 4) Natural Selection III. Mutation • Mutation - a heritable change in the genetic code (DNA) • Mutations are random events, they may be helpful or harmful Harmful Mutations: • Most mutations are harmful. Why? • Each individual inherits a combination of many genes that are already fine tuned by selection. • Therefore, the product of a mutant gene is likely to be less functional not more so. Helpful and Neutral Mutations • (mutations that survive in the population) • Every once in a great while a helpful mutation may arise and be beneficial to the organisms. • Some mutations have neutral effects or sometimes the effect of the mutant gene is masked by a dominant allele. • These genes are passed on by chance. IV. Genetic Drift • Allele frequencies can change randomly through generations because of chance. This process is called genetic drift GENETIC DRIFT Start with five different alleles Due to random events, only two different alleles are left after several generations. • Genetic drift is most rapid when population size is small. • During extreme cases of genetic drift, a population originates or is rebuilt from very few individuals • Founder Effect - a few individuals leave a population and establish a new one. By chance the allele frequencies for many traits may not be the same as in the original population. • Bottlenecks - disease, starvation, or some other disaster can nearly wipe out large populations. Even though the population recovers, the relative abundance of alleles has been altered at random V. Gene Flow • Allele frequencies change when individuals leave a population (migration) or enter a population. • This movement of individuals is called gene flow. VI. Natural Selection • Natural Selection - the differential survival and reproduction of individuals in a population based on the traits they posses • Natural selection may have a directional, stabilizing, or disruptive effects on the range of traits (phenotypes) in a population a)Directional selection - shifts allele frequencies in a constant direction in response to the environment b) Stabilizing selection - favors the most common phenotype in a population • Peccaries are eating those plants with low-spine-number causing their alleles to vanish from the gene pool • STABILIZING SELECTION • Preferring densely spined cacti, egglaying parasites more often destroy varieties of plants with larger numbers of spines. An infested cactus rarely survives. c)Disruptive Selection- favors characteristics at both ends of the range of traits in the population and operates against intermediate forms. • DISRUPTIVE SELECTION • Low-spine-number plants are not picked because they don't "look right", and highspine-number varieties are left alone because they are too hard to pick. VII. Speciation • Speciation - the process by which species originate • How does speciation occur? • Sometimes barriers arise between the parts of the population and create local breeding units. Then, two or more gene pools exist when there was only one. • If over time there is no gene flow between the two populations then selection, mutation, and genetic drift can operate differently in each population. • This can cause different traits to arise in each population. This is called divergence • When the divergence is great enough that the two populations will no longer interbreed, the two population have become different species. • Geographic Isolation -Populations can be separated by geographic barriers ex. mountains, rivers • Ex. The Blue-headed wrasse was separated from the Rainbow wrasse when the Isthmus of Panama was created. • Reproductive Isolation - any aspect of structure or behavior that prevents interbreeding • ex. the development of different breeding seasons or different mating behaviors • Over time, two species of frog have developed different mating calls through isolation. • Females respond only to the calls of their species.