README.

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Two files are available:
1) R code to run simulation:
2) R workspace containing individual level and population level wolf data from
Yellowstone, and results from simulation models:
The R code runs the IPM and returns data a baseline simulation and simulations
where intercept means are perturbed.
The R workspace contains simulation results and wolf data in the following
objects
A list of simulation results where means of intercept distributions are perturbed
(perturb.mean), a list of simulation results where variances of intercept
distributions are perturbed (perturb.var), a dataframe of individual-level wolf
data (wolf.lifehistory.data) and a dataframe of population level wolf data
(wolf.population.data).
The structure of perturb.mean and perturb.var lists:
perturb.mean[[1]] – results from baseline model
perturb.mean[[2]] – results from simulation where survival intercept
distribution mean increased by 5%
perturb.mean[[3]] – results from simulation where survival intercept
distribution mean increased by 10%
perturb.mean[[4]] – results from simulation where survival intercept
distribution mean increased by 20%
perturb.mean[[5]] – results from simulation where fertility intercept distribution
mean increased by 5%
perturb.mean[[6]] – results from simulation where fertility intercept distribution
mean increased by 10%
perturb.mean[[7]] – results from simulation where fertility intercept
distribution mean increased by 20%
perturb.mean[[8]] – results from simulation where offspring number intercept
distribution mean increased by 5%
perturb.mean[[9]] – results from simulation where offspring number intercept
distribution mean increased by 10%
perturb.mean[[10]] – results from simulation where offspring number intercept
distribution mean increased by 20%
perturb.mean[[11]] – results from simulation where growth < 41.7kg intercept
distribution mean increased by 5%
perturb.mean[[12]] – results from simulation where growth < 41.7kg intercept
distribution mean increased by 10%
perturb.mean[[13]] – results from simulation where growth < 41.7kg intercept
distribution mean increased by 20%
perturb.mean[[14]] – results from simulation where growth > 41.7kg intercept
distribution mean increased by 5%
perturb.mean[[15]] – results from simulation where growth > 41.7kg intercept
distribution mean increased by 10%
perturb.mean[[16]] – results from simulation where growth > 41.7kg intercept
distribution mean increased by 20%
perturb.mean[[17]] – results from simulation where inheritance intercept
distribution mean increased by 5%
perturb.mean[[18]] – results from simulation where inheritance intercept
distribution mean increased by 10%
perturb.mean[[19]] – results from simulation where inheritance intercept
distribution mean increased by 20%
perturb.var[[1]] – results from baseline model
perturb.var [[2]] – results from simulation where survival intercept
distribution mean increased by 5%
perturb.var[[3]] – results from simulation where survival intercept
distribution mean increased by 10%
perturb.var[[4]] – results from simulation where survival intercept
distribution mean increased by 20%
perturb.var[[5]] – results from simulation where fertility intercept distribution
mean increased by 5%
perturb.var[[6]] – results from simulation where fertility intercept distribution
variance increased by 10%
perturb.var[[7]] – results from simulation where fertility intercept
distribution variance increased by 20%
perturb.var[[8]] – results from simulation where offspring number intercept
distribution variance increased by 5%
perturb.var[[9]] – results from simulation where offspring number intercept
distribution variance increased by 10%
perturb.var[[10]] – results from simulation where offspring number intercept
distribution variance increased by 20%
perturb.var[[11]] – results from simulation where growth < 41.7kg intercept
distribution variance increased by 5%
perturb.var[[12]] – results from simulation where growth < 41.7kg intercept
distribution variance increased by 10%
perturb.var[[13]] – results from simulation where growth < 41.7kg intercept
distribution variance increased by 20%
perturb.var[[14]] – results from simulation where growth > 41.7kg intercept
distribution variance increased by 5%
perturb.var[[15]] – results from simulation where growth > 41.7kg intercept
distribution variance increased by 10%
perturb.var[[16]] – results from simulation where growth > 41.7kg intercept
distribution variance increased by 20%
perturb.var[[17]] – results from simulation where inheritance intercept
distribution variance increased by 5%
perturb.var[[18]] – results from simulation where inheritance intercept
distribution variance increased by 10%
perturb.var[[19]] – results from simulation where inheritance intercept
distribution variance increased by 20%
Simulated time series of cohort level population biology parameters
perturb.mean[[x]][[1]][1:4,1,] – time series of mean LRS for 1) all individuals, 2)
AA genotypes, 3) AB genotypes and 4) BB genotypes.
…[[x]][[1]][1:4,2,] – time series of variance in LRS for 1) all individuals, 2) AA
genotypes, 3) AB genotypes and 4) BB genotypes.
…[[x]][[1]][1:4,3,] – time series of generation length for 1) all individuals, 2) AA
genotypes, 3) AB genotypes and 4) BB genotypes.
…[[x]][[1]][1:4,4,] – time series of variance in generation length for 1) all
individuals, 2) AA genotypes, 3) AB genotypes and 4) BB genotypes.
…[[x]][[1]][1:4,5,] – time series of heritability of body weight for 1) all
individuals, 2) AA genotypes, 3) AB genotypes and 4) BB genotypes.
…[[x]][[1]][1:4,6,] – time series of additive genetic variance for body weight for
1) all individuals, 2) AA genotypes, 3) AB genotypes and 4) BB genotypes.
Simulated time series of per year population biology parameters
perturb.mean[[x]][[2]][1,1,] – time series of population size
…[[x]][[2]]2:4,1,] – time series of genotype frequencies for 2) AA genotypes, 3)
AB genotypes and 4) BB genotypes.
…[[x]][[2]][1:4,2,] – time series of mean survival rates for 1) all individuals, 2)
AA genotypes, 3) AB genotypes and 4) BB genotypes.
…[[x]][[2]][1:4,3,] – time series of mean reproductive success rates for 1) all
individuals, 2) AA genotypes, 3) AB genotypes and 4) BB genotypes.
…[[x]][[2]][1:4,4,] – time series of mean body weight for 1) all individuals, 2) AA
genotypes, 3) AB genotypes and 4) BB genotypes.
…[[x]][[2]][1:4,5,] – time series of variance in body weight for 1) all individuals,
2) AA genotypes, 3) AB genotypes and 4) BB genotypes.
Simulated time series of terms in the age-structured Price equation
…[[x]][[3]]1,1:4,] – time series of viability selection differentials for 1) all
individuals, 2) AA genotypes, 3) AB genotypes and 4) BB genotypes.
…[[x]][[3]]2,1:4,] – time series of fertility selection differentials for 1) all
individuals, 2) AA genotypes, 3) AB genotypes and 4) BB genotypes.
…[[x]][[3]]3,1:4,] – time series of growth contributions to delta z bar for 1) all
individuals, 2) AA genotypes, 3) AB genotypes and 4) BB genotypes.
…[[x]][[3]]4,1:4,] – time series of offspring-mother difference contributions
selection differentials for 1) all individuals, 2) AA genotypes, 3) AB genotypes
and 4) BB genotypes.
…[[x]][[3]]5:8,1:4,] – Price equation terms for delta sigma2 z (not reported in
paper).
Structure of dataframes containing wolf data
wolf.lifehistory.data - Individual level wolf data. Dataframe: each row is a wolf
identity, columns for id, year, age, sex, body weight, genotype.
wolf.population.data – population level wolf data. Dataframe: each row is a year,
columns are year, survival rate, reproductive success data, population size, mean
body weight, number of AA genotypes, number of AB genotypes, number of BB
genotypes.
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