Artifact 4.4

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Plecoptera:

Stoneflies

Overview:

Introduction

Taxonomy

Characteristics

Reproduction/Life cycle

Economic Importance

Plecoptera:

Pleco

”:folded “Ptera”: Wings

Hind wings fold under front wings when resting

Often seen resting on stones

Adults found around lakes and streams in cool, temperate parts of the world

Nymphs live in cool, well aerated lakes and streams

Small, ancient order of insects

Earliest fossils from Permian

Period

Triassic: first mammals, first dinosaurs

Permian Period:

Formation of Pangaea

286,000,000 yrs.ago

Taxonomy

• The Plecoptera order is divided into three suborders:

Archiperlaria

•2 Families

Filipalpia

•8 Families

Setipalpia

•4 families

Families and Species

In North America: 9

Families

465 Species

World wide: 15 families

Over 2,000 species

Three Major Families:

Perlidae: Common Stoneflies

Nemouridae: Spring Stoeflies

Taeniopterygidae: Winter stoneflies

Family: Perlidae

Common stoneflies

Largest family

Members most commonly collected

Perlidae

Found in mountains of eastern and western

North America

Emerge into adult form in spring & summer

Nemouridae

Spring emerging

Small & brown

Common in N. America

“x” pattern of veination near apex of wings

Taeniopterydigae

Medium sized, dark bodies Emerge in winter-spring

Taeniopteryx:

Common eastern and northern genus

Taenionema: common western group

Characteristics of Adults

Most have wings

Reduced or absent in some species

Membranous

Three Tarsi and Two Claws

Cerci: long, multi-segmented Long, segmented antenna

Mostly dull colored

Flat bodies

4-60 mm long

Hind legs not for jumping

Characteristics of Nymphs

Body shape: flat and elongated

Cerci: long and multisegmented

Tarsi: 3 segments

Thorax: Covered with large dorsal sclerites

Gills: branched and fillamented.

Sides of thorax and bases of legs

Nymphs

Thorax covered by dorsal sclerites

Legs widely separated

Long, segmented

Cerci

Reproduction &

Life Cycle

Males drum to attract females

Frequencies are species specific

Normal sequence:

Male drums, female drums back, male moves closer and drums again, female responds, cycle continues until they meet and mate.

Eggs deposited then hatch in 3-4 wks.

Eggs deposited either loose in the water or in a jelly in the water

Over 1,000 eggs deposited

Nymphs

Live in rivers and lakes

Most have gills

All require dissolved oxygen

Herbivorous and some carnivorous

Algae, diatoms, moss, immature mayfiles

Molt 12-36 times, 1-2 yrs. to mature

Full grown nymphs move to shoreline to molt one last time into adulthood.

Adults:

After emerging from water, fly or climb up nearby tree

Some adults feed, some do not

Non-feeding:

Have fully formed eggs

 mate quickly… die a few days later

Feeding species:

Females live for 4-5 weeks

Most males only live a few days

Drumming

Adult

Nymph

Boathouse

Mating

Eggs

Economic Importance

Indicate healthy streams and rivers

Important food source for game fish like trout and bass

The End

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