THEORY OF EVOLUTION Other Mechanisms of Change WHAT ARE WE LEARNING TODAY? Benchmarks Learning Objectives SC.912.L.15.13 – Describe the I will explain and describe conditions required for natural how mutation and genetic selection, including: overproduction recombination increase of offspring, inherited variation, and genetic variation. the struggle to survive, which result in I will explain and describe the differential reproductive success. scientific mechanisms, such SC.912.L.15.15 – Describe how as genetic drift, gene flow, mutation and genetic recombination and nonrandom mating, increase genetic variation. resulting in evolutionary change. SC.912.L.15.14 – Discuss mechanisms of evolutionary change other than natural selection, such as genetic drift and gene flow. WHAT ARE OTHER MECHANISMS OF EVOLUTION? Evolution is the change in a population’s genetic material over generations. Four other mechanisms of evolutionary change exist besides natural selection. Mutations Gene flow Genetic drift Nonrandom mating WHAT ARE MUTATIONS? A mutation is a change in the genetic material of a cell. Mutations affect evolution by producing totally new alleles. Germ mutations occur in the reproductive cells, or gametes, of an individual. These mutations can be passed on to the offspring. They can affect the survival of an individual organism or a genetic line of organisms. WHAT ARE MUTATIONS? Mutations can be: 1. Neutral- No affect • No pattern of increase or decrease in allele frequency 2. Harmful • Usually naturally selected against and rid of in short time 3. Beneficial • usually increase over time WHAT IS GENE FLOW? Gene flow is the process of genes moving from one population to another. Gene flow can occur through Immigration ( the movement of individuals into a population) Emigration (the movement of individuals out of a population) WHAT IS GENETIC DRIFT? Genetic drift is the process by which alleles frequencies in a population change as a result of random events, or chance. It can result in substantial changes within a population. Only significant in small and medium-sized populations. WHAT IS NONRANDOM MATING? Nonrandom mating occurs whenever individuals may choose partners. Influenced by geographic proximity and assortative mating Sexual selection occurs when certain traits increase an individual’s success at mating. REVIEW We have discussed how evolution by natural selection works • NATURAL SELECTION requires: 1. 2. 3. 4. Overproduction of offspring Inherited Variation Struggle to survive Differential Reproduction • We have discussed several ways natural selection can act on populations • • • • *Genetic drift Gene flow Mutations Non-random mating You should be able to explain/describe each of these concepts by now LETS GO OVER GENETIC DRIFT AGAIN • Genetic Drift is a RANDOM change in allele frequencies in a population due to a completely random event • 2 types of Genetic Drift Situations: 1. Bottleneck effect 1. Founder’s effect GENETIC DRIFT: BOTTLENECK EFFECT Sometimes a random disaster may kill many individuals in a population. Those alleles left may not be the same representation as the original population GENETIC DRIFT: FOUNDER’S EFFECT Migration of a small subgroup from a main population leads to a difference in allele frequencies from the original population’s The founding population will reproduce and pass on the alleles of the founders RATES OF EVOLUTION: PUNCTUATED EQUILIBRIUM VS GRADUALISM Gradualism • Small changes occurring little by little • Usually hard to notice over short period of time Punctuated Equilibrium • Change occurs in spurts • Period of little change, followed by period of huge change • Change is noticeable and abrupt • Change is slow, constant, and consistent Gradualism Punctuated Equilibrium 3 TYPES OF NATURAL SELECTION 1. Directional Selection • Better “fitness” toward 1 direction • Example: bigger the beak, the better fit 2. Stabilizing Selection • The intermediate trait is best fit • Example: the grey species is better fit than the white or black 3. Disruptive Selection • the extreme, or outer end traits are better fit than the intermediate trait • Example: the white and black species is better fit than the grey species 3 TYPES OF SELECTION MICROEVOLUTION VS MACROEVOLUTION • Microevolution is evolution based on the genetic level • Genetic drift • Mutation • Nonrandom mating • Gene flow • Macroevolution is evolution based on more visually observable changes in species • Extinction • Speciation • Adaptive radiation • Convergent Evolution • Co-evolution EXTINCTION AND SPECIATION • Extinction is when a species disappears forever from the earth as a result to its population being completely diminished • Speciation is the formation of new species from already existing species • But how do new species form? • Geographic isolation • Temporal isolation • Behavioral isolation Reproductive isolation New Species REPRODUCTIVE BARRIERS PRE- Reproductive Barriers • Habitat Isolation • Geographic isolation • Temporal isolation • Mechanical isolation • Behavioral isolation • Gametic (reproductive) islation POST- Reproductive Barriers • Hybrid Sterility • Reduced Hybrid Viability ADAPTIVE RADIATION A single species evolves over a relatively short period of time into several forms that live in different ways Galapagos finches evolved different beaks and behaviors that allow them to eat different kinds of food CONVERGENT EVOLUTION When distantly, unrelated species LOOK similar because they have evolved in similar environments CO-EVOLUTION Sometimes 2 different species live and interact so closely together, they evolve together The evolution of one species drives the evolution of the other EXAMPLE: • Flowers and pollinators