ORIGIN & EVOLUTION NOTES According to current evolutionary theory, life appeared on earth about 3.5 billion years ago. Evidence from microfossils and radiometric dating (Carbon-14 or Potassium-40) support this theory. Evolution—the theory that species change over time. Types of evolution include: Microevolution—change in allele frequencies within a population Macroevolution—evolutionary change at the species level or higher; that is, the formation of new species, new genera, and so forth. Important proponents of evolutionary theory in order of occurrence include: Charles Lyell—a Scottish geologist wrote Principles of Geology (1830). Lyell proposed that plant and animal species had arisen, developed variations, and then became extinct over time. He also believed that the Earth’s physical landscape changed over a long period of time. Page 1 of 6 Thomas Malthus—wrote an essay called The Principles of Population. In it, Malthus proposed that populations outgrew their food supplies, causing competition between organisms and a struggle for one species to survive against another. Jean Baptist de Lamarck—proposed the first mechanism for evolution called “the inheritance of acquired characteristics”. Charles Darwin—naturalist on ship HMS Beagle. Ship was chartered for a five year mapping and collecting expedition to South America and the South Pacific. Wrote a book called On the Origin of Species, which proposed the mechanisms for evolution to be a process called natural selection. Indirect evidence for evolution includes: The fossil record The geologic column or time scale—based upon the idea that the earths layers can be dated according to what fossils they contain, older organisms being found near the bottom of the column and younger organisms found nearer the top of the column. By matching rock layers with fossils, geologists can determine the age of the rocks, while paleontologists can determine the age of the fossils. This is called relative dating. Includes the Page 2 of 6 Precambrian, Paleozoic, Mesozoic and Cenozoic (most recent) Eras Comparative anatomy 1. Homologous structures—forelimb bones of some 2. Analogous structures—body parts of organisms 3. Vestigial structures—body structure that has no 4. Embryology—the study of how animal embryos mammals exhibit this. that do not have a common evolutionary origin, but are similar in function. Examples would include insect wings and bird wings. function in a present day organism but was probably useful to an ancestor. develop; during early stages of development many animal embryos look similar Biochemistry—comparisons of the proteins, DNA, or RNA of different species. Direct evidence for evolution includes: Pesticide resistance in insects and other organisms Antibiotic resistance in bacteria Processes involved in evolution include: Page 3 of 6 Natural selection—the process by which organisms with favorable adaptations survives, and reproduces at higher rates than organisms with less favorable adaptations. From all the information gathered by Darwin, two central concepts emerged to form the basis of his theory of evolution. First, Darwin observed that variations within a species were dependent on the environment. Adaptations are genetically coded traits that occur in organisms and enable them to be more successful in their environment. Darwin reasoned that the importance of these adaptations is to ensure the survival through reproduction of that species. The organisms that lack these adaptations will not reproduce as successfully. Secondly, the organisms on the Galapagos Islands had become geographically separated from one another. This resulted in reproductive isolation. There is no interbreeding between organisms of the same species that are located on different islands. For example, finches on one island could not breed with finches of the same species on another island. He theorized that within a population of a species, adaptations would arise due to reproductive isolation. The organisms would develop adaptations to their environment over time that would result in significant differences between the same species on different islands. Page 4 of 6 Types of natural selection include: 1. 2. 3. 4. 5. Stabilizing selection—individuals with the average form of a trait are selected. Directional selection—individuals with one extreme form of a trait are selected. Disruptive selection—individuals with either extreme form of a trait are selected. Sexual selection—preferential choice of a mate is based on the presence of a particular trait. Artificial selection—intentional breeding of organisms to produce certain traits Extinction—of the species that have existed on the planet, more than 99% are extinct. Mass extinctions are thought to be brought about by some sort of catastrophe or climate change that produces conditions too severe for most species to endure. Extinction is thought to be a major cause of evolution. Speciation is the evolution of a new species that occurs when interbreeding happens, or when the production of fertile offspring is prevented. In the physical world, natural barriers form and cause the breakup of populations to form smaller populations. Volcanoes, sea-level changes, and earthquakes are a few examples of natural occurrences that affect populations. This is known as geographic isolation. Geographic isolation prevents interbreeding, so gene exchange will cease. So over time, each smaller population will adapt to their new environment through the process of natural selection. Page 5 of 6 Eventually, this causes the gene pool of each group to become different so that it can be concluded that there is a new species formed. The rate of speciation can proceed in one of two timeframes: Gradual evolution (gradualism)—idea that evolution occurs a slow even rate over extremely long periods of time Punctuated equilibrium—trends in the fossil record cannot be attributed to gradual change within a lineage, but rather to quick bursts of evolutionary change. Patterns of evolution: Adaptive radiation—when diversity seems to have occurred in a newly evolved species in a relatively short time. They also believe it occurs when an organism colonizes a new area in which there is another organism that is lacking in survival skills. Researchers use the example of the finches Darwin observed on the Galapagos Islands. He counted over a dozen different kinds of finches that he believed evolved from a single founding species. Divergent evolution (adaptive radiation is one type of this)—pattern of evolution in which species that once were similar to an ancestors species diverge, or become increasingly distinct. Convergent evolution—occurs when unrelated species occupy similar environments in different parts of the world. Because they share similar environmental pressures, and therefore, similar pressures of natural selection, they tend to have many of the same characteristics. Page 6 of 6