Other Concepts of Species 1. Ecological Species Concept 2. Pluralistic Species Concept 3. Morphological Species Concept 4. Genealogical Species Concept 3.Morphological Species Concept: the oldest and still most practical, defines a species by a unique set of structural features. 4.Genealogical Species Concept: defines a species as a set of organisms with a unique genetic history—one tip of the branching tree of life. Speciation: Plants • Polyploids may cause new species because the change in chromosome number creates postzygotic barriers. • Polyploid Types: 1. Autopolyploid - when a species doubles its chromosome number from 2N to 4N. 2. Allopolyploid - formed as a polyploid hybrid between two species. – Ex: wheat 1.Ecological Species Concept: defines a species in terms of its ecological niche, the set of environmental resources that a species uses and its role in a biological community 2.Pluralistic Species Concept: concept may invoke reproductive isolation or adaptation to an ecological niche, or use both in maintaining distinctive, cohesive groups of individuals Speciation Event: Adaptive Radiation • Resources are temporarily infinite. • Most offspring survive. • New species form rapidly if isolation mechanisms work. • Result - little Natural Selection and the gene pool can become very diverse. Autopolyploid Allopolyploid Origin of Evolutionary Novelty • How do macroevolution changes originate? – Exaptation – Heterochrony – Homeosis Exaptation • When a structure that was adapted for one context is co-opted for another function. • Ex. – feathers and flying- adaptation for insulation and a exaptation for flight Eample: Trap-jaw ants, manibles were adaptation for manipulation of objects, adaptation for defense Heterochrony • Changes in the timing or rate of development. 1. Allometric Growth 2. Paedomorphosis What is the difference between this and ADAPTATION? Heterochrony: Allometric Growth 1. Allometric Growth – changes in the relative rates of growth of various parts of the body. • Ex. – skull growth in primates Heterochrony: Paedomorphosis 2. Paedomorphosis – when an adult retains features that are present in the juvenile form. Ex. – gills in adult salamanders Homeosis Changes in the basic body design or arrangement of body parts. Ex. – Hox gene clusters that gave rise to vertebrates from invertebrates. Refuting Misconceptions • Evolution is not goal oriented. It does not produce “perfect” species. Refuting Misconceptions • Evolution does not mean that life evolved by “chance.” •Evolution is not an explanation for the origin of life. Only an explanation of how life functions after its origin. •Natural Selection does not involve organisms “trying to adapt.” Refuting Misconceptions • Natural Selection does not give organisms what they “need” •Evolution as a “THEORY”